Keroplatidae

Male fungus gnat, probably Macrocera, photographed by Tony DeSantis.


Belongs within: Sciaroidea.

The Keroplatidae is a group of fungus gnats whose larvae live in webs. Larvae of at least some species feed on small animals caught in the web.

Characters (from Vockeroth 2009): Body length 2.3–8.8 mm. Colour yellow to brown or black, often patterned. Usually three ocelli, middle ocellus small or rarely absent. Antenna with fourteen flagellomeres. Scutum uniformly setose or with bare longitudinal stripes; scutellum with posterior row of setae or with dorsal surface extensively setose. Scutellum sometimes with large membranous triangular basal area. Mediotergite bare. Pleural sclerites and prosternum variously haired or bare. Wing with Sc ending in C; R2+3 present, ending in R1 or C; base of M strong, weak or absent; Rs and M briefly fused at about one-third wing length. Foretibia with one apical spur, other tibiae with one or two spurs.

<==Keroplatidae
    |  i. s.: Orfelia nemoralis TC04 [=Platyura nemoralis M83]
    |         Asindulum GE05
    |         Arachnocampa Edwards 1924 GE05, TE27
    |           |--*A. luminosa (Skuse 1891) [=Bolitophila luminosa] TE27
    |           `--A. flava WT11
    |         Planivora GE05
    |         Neoditomyia L88
    |           |--N. spinosa L88
    |           `--+--N. andina L88
    |              `--N. troglophila L88
    |--Macrocerinae GE05
    |    |--Paramacrocera Tonnoir & Edwards 1927 TE27
    |    |    `--*P. brevicornis Edwards in Tonnoir & Edwards 1927 TE27
    |    `--Macrocera TE27
    |         |--M. annulata Tonnoir in Tonnoir & Edwards 1927 TE27
    |         |--M. antennalis Marshall 1894 TE27
    |         |--M. campbelli Edwards in Tonnoir & Edwards 1927 TE27
    |         |--M. centralis JP76
    |         |--M. fenestrata Edwards in Tonnoir & Edwards 1927 TE27
    |         |--M. fusca Tonnoir in Tonnoir & Edwards 1927 TE27
    |         |--M. glabrata Tonnoir in Tonnoir & Edwards 1927 TE27
    |         |--M. gourlayi Tonnoir in Tonnoir & Edwards 1927 TE27
    |         |--M. hudsoni Tonnoir in Tonnoir & Edwards 1927 TE27
    |         |--M. inconspicua Tonnoir in Tonnoir & Edwards 1927 TE27
    |         |--M. magna P76
    |         |--M. milligani Tonnoir in Tonnoir & Edwards 1927 TE27
    |         |--M. montana Marshall 1896 [incl. M. howletti Marshall 1896] TE27
    |         |--M. ngaireae Edwards in Tonnoir & Edwards 1927 TE27
    |         |--M. obsoleta Edwards in Tonnoir & Edwards 1927 TE27
    |         |--M. parva P76
    |         |--M. pulchra Tonnoir in Tonnoir & Edwards 1927 TE27
    |         |--M. ruficollis Edwards in Tonnoir & Edwards 1927 TE27
    |         |--M. scoparia Marshall 1896 TE27
    |         `--M. unipunctata Tonnoir in Tonnoir & Edwards 1927 TE27
    `--Keroplatinae [Ceroplatinae] TE27
         |--Pseudoplatyura truncata Tonnoir in Tonnoir & Edwards 1927 TE27
         |--Isoneuromyia TE27
         |--Cerotelion TE27
         |    |--C. bimaculatus Tonnoir in Tonnoir & Edwards 1927 TE27
         |    |--C. dendyi (Marshall 1896) [=Ceroplatus dendyi] TE27
         |    |--C. hudsoni (Marshall 1896) [=Ceroplatus hudsoni] TE27
         |    |--C. leucoceras (Marshall 1896) [=Ceroplatus leucoceras] TE27
         |    |--C. niger Tonnoir in Tonnoir & Edwards 1927 TE27
         |    |--C. tapleyi Edwards in Tonnoir & Edwards 1927 TE27
         |    `--C. vitripennis Tonnoir in Tonnoir & Edwards 1927 TE27
         |--Platyura TE27
         |    |--P. agricola Marshall 1896 TE27
         |    |--P. albovittata Tonnoir in Tonnoir & Edwards 1927 TE27
         |    |--P. brevis Tonnoir in Tonnoir & Edwards 1927 TE27
         |    |--P. brookesi Edwards in Tonnoir & Edwards 1927 TE27
         |    |--P. campbelli Tonnoir in Tonnoir & Edwards 1927 TE27
         |    |--P. carbonaria Tonnoir in Tonnoir & Edwards 1927 TE27
         |    |--P. chiltoni Tonnoir in Tonnoir & Edwards 1927 TE27
         |    |--P. curtisi Edwards in Tonnoir & Edwards 1927 TE27
         |    |--P. harrisi Tonnoir in Tonnoir & Edwards 1927 TE27
         |    |--P. lamellata Tonnoir in Tonnoir & Edwards 1927 TE27
         |    |--P. maculipennis Tonnoir in Tonnoir & Edwards 1927 TE27
         |    |--P. marshalli Tonnoir & Edwards 1927 [=P. flava Marshall 1896 (preoc.)] TE27
         |    |--P. novaezelandiae Tonnoir & Edwards 1927 [=P. magna Marshall 1896 (preoc.)] TE27
         |    |--P. ohakunensis Edwards in Tonnoir & Edwards 1927 TE27
         |    |--P. philpotti Tonnoir in Tonnoir & Edwards 1927 TE27
         |    |--P. proxima Tonnoir in Tonnoir & Edwards 1927 TE27
         |    |--P. punctifusa Edwards in Tonnoir & Edwards 1927 TE27
         |    |--P. ruficauda Tonnoir in Tonnoir & Edwards 1927 TE27
         |    |--P. rufipectus Tonnoir in Tonnoir & Edwards 1927 TE27
         |    |--P. rutila Edwards in Tonnoir & Edwards 1927 TE27
         |    `--P. subbrevis Tonnoir in Tonnoir & Edwards 1927 TE27
         `--Ceroplatus G60
              |--C. mangalorensis Edwards 1929 G60
              `--C. mastersi Skuse 1888 G60

*Type species of generic name indicated

REFERENCES

[G60] Gatenby, J. B. 1960. The Australasian mycetophilid glowworms. Transactions of the Royal Society of New Zealand 88 (3): 577-593.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[JP76] Joost, W., & E. Plassmann. 1976. Mycetophiliden aus dem Zentralkaukasus (Diptera: Mycetophilidae). Senckenbergiana Biologica 57 (1-3): 67-68.

[L88] Liebherr, J. K. 1988. General patterns in West Indian insects, and graphical biogeographic analysis of some circum-Caribbean Platynus beetles (Carabidae). Systematic Zoology 37 (4): 385-409.

[M83] Martin, N. A. 1983. Miscellaneous observations on a pasture fauna: An annotated species list. DSIR Entomology Division Report 3: 1-98.

[P76] Plassmann, E. 1976. Vierter Nachtrag zu der Mycetophiliden-Sammlung des Senckenberg-Museums Frankfurt am Main (Diptera; Mycetophilidae). Senckenbergiana Biologica 57 (1-3): 77-79.

[TC04] Toft, R. J., & P. J. Chandler. 2004. Three introduced species of Mycetophilidae (Diptera: Sciaroidea) established in New Zealand. New Zealand Entomologist 27: 43-49.

[TE27] Tonnoir, A. L., & F. W. Edwards. 1927. New Zealand fungus gnats (Diptera, Mycetophilidae). Transactions and Proceedings of the New Zealand Institute 57: 747-878.

Vockeroth, J. R. 2009. Keroplatidae (predatory fungus gnats). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1, pp. 259-263. NRC Research Press: Ottawa.

[WT11] Wiegmann, B. M., M. D. Trautwein, I. S. Winkler, N. B. Barr, J.-W. Kim, C. Lambkin, M. A. Bertone, B. K. Cassel, K. M. Bayless, A. M. Heimberg, B. M. Wheeler, K. J. Peterson, T. Pape, B. J. Sinclair, J. H. Skevington, V. Blagoderov, J. Caravas, S. N. Kutty, U. Schmidt-Ott, G. E. Kampmeier, F. C. Thompson, D. A. Grimaldi, A. T. Beckenbach, G. W. Courtney, M. Friedrich, R. Meier & D. K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the USA 108 (14): 5690-5695.

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