Assemblage of Pleurocystites squamosus, from here.

Belongs within: Dichoporita.
Contains: Callocystitidae, Echinoencrinitidae.

The Glyptocystitida are a Lower Ordovician to Upper Devonian group of cystoids in which the pore groups have well-exposed openings.

Characters (from Kesling 1967): Theca in most genera composed of four BB, five ILL, five LL, five RR, and five OO, though plate O1 is tripartite in many genera and R1 is bipartite in some; a few anomalous plate divisions known to occur in some specimens. Rhombs developed as distinct pectinirhombs, most as full rhomgs and a few as demirhombs, with well-exposed pores or slits. Column present, in most forms with expanded, flexible proximal part.

<==Glyptocystitida [Glyptocystidea, Regularia, Rhombiporitidae]
    |--+--+--Callocystitidae K67
    |  |  |--Echinoencrinitidae K67
    |  |  `--Rhombifera Barrande 1867 [Rhombiferidae] K67
    |  |       `--*R. bohemica Barrande 1867 K67
    |  `--Cystoblastidae K67
    |       |--Hesperocystis Sinclair 1945 K67
    |       |    `--*H. deckeri Sinclair 1945 K67
    |       `--Cystoblastus Volborth 1867 K67
    |            |--*C. leuchtenbergi Volborth 1867 K67
    |            `--C. kokeni K67
    `--+--Pleurocystitidae [Pleurocystidae] K67
       |    |--Amecystis Ulrich & Kirk 1921 K67
       |    |    `--*A. laevis (Raymond 1921) [=Pleurocystites laevis] K67
       |    |--Regulaecystis Dehm 1932 K67
       |    |    `--*R. pleurocystoides Dehm 1932 K67
       |    `--Pleurocystites Billings 1854 [=Pleurocystis Carpenter 1891; incl. Dipleurocystis Jaekel 1918] K67
       |         |--*P. squamosus Billings 1854 [=*Pleurocystis squamosus] K67
       |         |--P. anglicus (Jaekel 1899) [=*Dipleurocystis anglicus] K67
       |         |--P. anticostiensis K67
       |         |--P. bassleri K67
       |         |--P. beckeri K67
       |         |--P. clermontensis K67
       |         |--P. elegans K67
       |         |--P. exornatus K67
       |         |--P. filitextus K67
       |         |--P. foriolus K67
       |         |--P. gibbus K67
       |         |--P. matutinus K67
       |         |--P. mercerensis K67
       |         |--P. multistriatus K67
       |         |--P. procerus K67
       |         |--P. quadratus K67
       |         |--P. robustus K67
       |         |--P. rugeri K67
       |         |--P. slocomi K67
       |         `--P. watkinsi K67
       `--+--Glyptocystites Billings 1854 (see below for synonymy) K67
          |    |--*G. multiporus Billings 1854 K67
          |    |--G. batheri K67
          |    |--G. ehlersi K67
          |    |--G. grandis K67
          |    `--G. regnelli K67
          `--Cheirocrinidae [Chirocrinidae] K67
               |--Leptocystis Jaeckel 1899 K67
               |    |--*L. tertia (Barrande 1887) [=Homocystites tertius] K67
               |    |--L. atava K67
               |    |--L. constricta K67
               |    `--L. gigantea K67
               |--Homocystites Barrande 1887 [=Homocystis Bather 1889] K67
               |    |--*H. alter Barrande 1887 K67
               |    |--H. anatiformis K67
               |    |--H. angulatus K67
               |    |--H. degener K67
               |    |--H. forbesi K67
               |    |--H. insignis [=Cheirocrinus insignis] K67
               |    |--H. sculptus K67
               |    |--H. striatus K67
               |    `--H. walcotti K67
               `--Cheirocrinus Eichwald 1856 [=Chirocrinus Haeckel 1896 non Angelin 1878] K67
                    |--*C. penniger (Eichwald 1842) [=Cyathocrinus penniger] K67
                    |--C. atavus Jaekel 1899 B64
                    |--C. constrictus Bather 1913 B64
                    |--C. dilatus K67
                    |--C. giganteus (Leuchtenberg 1843) B64
                    |--C. granulatus K67
                    |--C. holmi K67
                    |--C. hyperboreus K67
                    |--C. interruptus K67
                    |--C. leuchtenbergi K67
                    |--C. merrileesi Brown 1964 B64
                    |--C. nodosus K67
                    |--C. ornatus K67
                    |--C. radiatus K67
                    `--C. volborthi K67

Glyptocystites Billings 1854 [=Glyptocystis Bather 1900; Glyptocystidae, Glyptocystinae, Glyptocystitidae] K67

*Type species of generic name indicated


[B64] Brown, I. A. 1964. An Ordovician cystoid (Pematozoa, Echinodermata) from Western Australia. Journal of the Royal Society of Western Australia 47 (1): 3-7.

[K67] Kesling, R. V. 1967. Cystoids. In Treatise on Invertebrate Paleontology pt. S. Echinodermata 1. General characters. Homalozoa-Crinozoa (except Crinoidea) (R. C. Moore, ed.) vol. 1 pp. S85-S267. The Geological Society of America, Inc., and The University of Kansas: Lawrence (Kansas).

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