Ditomyiidae

Symmerus annulatus, photographed by J. K. Lindsey.

Belongs within: Sciaroidea.

The Ditomyiidae are a group of fungus gnats characterised by wings with the media and cubitus connected, and vein R₄ relatively long (Tonnoir & Edwards 1927). An incomplete eye bridge may be present above the antennae in some genera; the eyes are almost touching in Symmerus (Vockeroth 1981).

Characters (from Tonnoir & Edwards 1927, as Ditomyiinae): R₄ present and rather long, generally half or more than half as long as Rs; Sc short and ending free; posterior divisions of pronotum with one or more longish bristles.

<==Ditomyiidae [Ditomyiinae]
    |--Ditomyia potomaca V81
    |--Rhipidita [incl. Calliceratomyia] V09
    |--Celebesomyia inocellata O98
    |--Australosymmerus [incl. Austrosymmerus] V09
    |    `--A. acutus V09
    |--Symmerus WT11
    |    |--S. annulatus WT11
    |    |--S. coqulus V81
    |    |--S. (Symmerus) lautus V81
    |    `--S. vockerothi V81
    |--Centrocnemis TE27
    |    |--C. basalis Tonnoir in Tonnoir & Edwards 1927 TE27
    |    |--C. fumipennis Tonnoir in Tonnoir & Edwards 1927 TE27
    |    |--C. nitida Tonnoir in Tonnoir & Edwards 1927 TE27
    |    |--C. tillyardi Tonnoir in Tonnoir & Edwards 1927 TE27
    |    `--C. trivittata Edwards in Tonnoir & Edwards 1927 TE27
    `--Nervijuncta Marshall 1896 (see below for synonymy) TE27
         |--*N. nigrescens Marshall 1896 TE27
         |--N. bicolor Edwards in Tonnoir & Edwards 1927 TE27
         |--N. flavoscutellata Tonnoir in Tonnoir & Edwards 1927 TE27
         |--N. harrisi Edwards in Tonnoir & Edwards 1927 TE27
         |--N. hexachaeta Edwards in Tonnoir & Edwards 1927 TE27
         |--N. hudsoni (Marshall 1896) [=*Cyrtoneura hudsoni, *Arctoneura hudsoni] TE27
         |--N. longicauda Edwards in Tonnoir & Edwards 1927 TE27
         |--N. marshalli Edwards in Tonnoir & Edwards 1927 TE27
         |--N. nigricornis Tonnoir in Tonnoir & Edwards 1927 TE27
         |--N. nigricoxa Edwards in Tonnoir & Edwards 1927 TE27
         |--N. ostensackeni Tonnoir in Tonnoir & Edwards 1927 TE27
         |--N. parvicauda Edwards in Tonnoir & Edwards 1927 (see below for synonymy) TE27
         |--N. pilicornis Edwards in Tonnoir & Edwards 1927 TE27
         |--N. pulchella Edwards in Tonnoir & Edwards 1927 TE27
         |--N. punctata Tonnoir in Tonnoir & Edwards 1927 (see below for synonymy) TE27
         |--N. ruficeps Edwards in Tonnoir & Edwards 1927 [incl. N. ruficeps var. continua Tonnoir & Edwards 1927] TE27
         |--N. tridens (Hutton 1881) [=Platyura tridens, *Casa tridens, *Huttonia tridens] TE27
         `--N. wakefieldi (Edwards 1921) (see below for synonymy) TE27

Nervijuncta Marshall 1896 [incl. Cyrtoneura Marshall 1896 (preoc.), Arctoneura Hutton 1901, Huttonia Marshall 1896 nec Pickard-Cambridge 1879 nec Kirk 1882, Casa Hutton 1901] TE27

Nervijuncta parvicauda Edwards in Tonnoir & Edwards 1927 [incl. N. parvicauda var. suffusa Tonnoir & Edwards 1927] TE27

Nervijuncta punctata Tonnoir in Tonnoir & Edwards 1927 [incl. N. punctata var. robusta Tonnoir in Tonnoir & Edwards 1927] TE27

Nervijuncta wakefieldi (Edwards 1921) [=Arctoneura wakefieldi; incl. N. wakefieldi var. abbreviata Tonnoir & Edwards 1927] TE27

*Type species of generic name indicated

REFERENCES

[O98] Oosterbroek, P. 1998. The Families of Diptera of the Malay Archipelago. Brill: Leiden.

[TE27] Tonnoir, A. L., & F. W. Edwards. 1927. New Zealand fungus gnats (Diptera, Mycetophilidae). Transactions and Proceedings of the New Zealand Institute 57: 747–878.

[V81] Vockeroth, J. R. 1981. Mycetophilidae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 223–246. Research Branch, Agriculture Canada.

[V09] Vockeroth, J. R. 2009. Ditomyiidae (ditomyiid fungus gnats). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 253–255. NRC Research Press: Ottawa.

[WT11] Wiegmann, B. M., M. D. Trautwein, I. S. Winkler, N. B. Barr, J.-W. Kim, C. Lambkin, M. A. Bertone, B. K. Cassel, K. M. Bayless, A. M. Heimberg, B. M. Wheeler, K. J. Peterson, T. Pape, B. J. Sinclair, J. H. Skevington, V. Blagoderov, J. Caravas, S. N. Kutty, U. Schmidt-Ott, G. E. Kampmeier, F. C. Thompson, D. A. Grimaldi, A. T. Beckenbach, G. W. Courtney, M. Friedrich, R. Meier & D. K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the USA 108 (14): 5690–5695.

Last updated: 18 June 2018.