Microvalgus

Microvalgus vagans, copyright Christian Moeseneder.

Belongs within: Cetoniinae.

Microvalgus is a genus of small cetoniine beetles with a dorsal covering of scales found in warmer regions of the Old World.

<==Microvalgus Kraatz 1883 CW92
    |--*M. lapeyrousei (Gory & Percheron 1833) [=Valgus lapeyrousei] CW92
    |--M. apicalis Lea 1914 CW92
    |--M. bursariae Lea 1914 CW92
    |--M. castaneipennis (Macleay 1871) [=Valgus castaneipennis] CW92
    |--M. dubius Lea 1914 CW92
    |--M. fasciculatus Lea 1914 CW92
    |--M. flavipennis Arrow 1944 CW92
    |--M. glaber Lea 1914 CW92
    |--M. mucronatus Lea 1914 CW92
    |--M. nigriceps Lea 1914 CW92
    |--M. nigrinus (Macleay 1871) [=Valgus nigrinus] CW92
    |--M. quinquedentatus Lea 1914 CW92
    |--M. rufipennis Lea 1914 CW92
    |--M. scutellaris Blackburn 1894 CW92
    |--M. squamiventris Lea 1914 CW92
    |--M. vagans Lea 1914 CW92
    |    |--M. v. vagans CW92
    |    `--M. v. obscuripennis Lea 1914 CW92
    `--M. yilgarnensis Blackburn 1892 CW92

*Type species of generic name indicated

REFERENCES

[CW92] Cassis, G., & T. A. Weir. 1992. Cetoniinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 426–454. AGPS Press: Canberra.

Pseudoclithria

Pseudoclithria maura, from Friends of Queens Park Bushland.

Belongs within: Cetoniinae.

Pseudoclithria is an Australian genus of cetoniine beetles with costate elytra and non-metallic coloration.

<==Pseudoclithria van der Poll 1886 [Xiphoscelidini] CW92
    |--P. hirticeps (Macleay 1871) CW92 (see below for synonymy)
    |--P. adusta (Janson 1889) [=Diaphonia adusta] CW92
    |--P. anchoralis Lea 1914 CW92
    |--P. dejecta Lea 1914 CW92
    |--P. erythroptera Lea 1914 CW92
    |--P. fossor Lea 1914 CW92
    |--P. kershawi Lea 1914 CW92
    |--P. mastersii (Macleay 1871) [=Schizorhina mastersii] CW92
    |--P. maura (Janson 1874) CW92 [=Diaphonia maura CW92, Metallesthes maura M86]
    |--P. ruficornis (Westwood 1874) [=Diaphonia ruficornis] CW92
    `--P. rugosa (Schaum 1848) CW92 [=Schizorrhina (Diaphonia) rugosa CW92, Metallesthes rugosa M86]

Pseudoclithria hirticeps (Macleay 1871) CW92 [=Schizorhina hirticeps CW92, Chondropyga hirticeps M86; incl. Clithria bicostata Kraatz 1885 CW92, *Pseudoclithria bicostata CW92, Diaphonia hirtifrons Thomson 1878 CW92, Schizorhina nigrans Macleay 1871 CW92]

*Type species of generic name indicated

REFERENCES

[CW92] Cassis, G., & T. A. Weir. 1992. Cetoniinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 426–454. AGPS Press: Canberra.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part III. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 21–126.

Glycyphana

Brown flower beetle Glycyphana stolata, copyright Peter Woodard.

Belongs within: Cetoniinae.

Glycyphana is an Indo-Australian genus of relatively small cetoniine beetles with a dorsum covered by a matt bloom marked with patches of lighter-coloured microtrichia.

<==Glycyphana Burmeister 1842 CW92
    |  i. s.: G. albomaculata Mohnike 1871 G89
    |         G. bisignata Kraatz 1885 G89
    |         G. malayensis Guér. 1840 [=Euryomia malayensis] G89
    |         G. rufovittata Guér. 1840 [=Euryomia rufovittata; incl. G. rufovittata var. melanaria Kraatz 1885] G89
    |         G. sinuata Wall 1868 G89
    |         G. velutina Macleay 1886 [=Glyciphana velutina] Mac86
    |--*G. (Glycyphana) horsfieldii (Hope 1835) [=Cetonia horsfieldii] CW92
    |--G. (Caloglycyphana Mikšić 1968) CW92
    |    |--G. (*C.) binotata (Gory & Percheron 1833) [=Cetonia binotata] CW92
    |    |--G. (C.) papua (Wallace 1867) [=Eurynomia papua; incl. G. ochreonotata Lea 1914] CW92
    |    `--G. (C.) pulchra (Macleay 1871) [=Schizorhina pulchra; incl. G. subdepressa Blackburn 1890] CW92
    `--G. (Glycyphaniola Mikšić 1968) CW92
         |--G. (*G.) modesta (Fabricius 1792) [=Cetonia modesta] CW92
         |--G. (G.) australiana Mikšić 1971 CW92
         |--G. (G.) brooksi Bacchus 1974 CW92
         |--G. (G.) brunnipes (Kirby 1818) CW92 (see below for synonymy)
         |--G. (G.) darwinensis Bacchus 1974 CW92
         |--G. (G.) pusilla Bacchus 1974 CW92
         |--G. (G.) stolata (Fabricius 1781) (see below for synonymy) CW92
         `--G. (G.) varicorensis Burmeister 1842 CW92
              |--G. v. varicorensis CW92
              `--G. v. thursdayana Mikšić 1971 CW92

Glycyphana (Glycyphana) brunnipes (Kirby 1818) CW92 [=Cetonia brunnipes CW92; incl. C. conspersa Gory & Percheron 1833 CW92, Glycyphana obscura Mas86, C. viridiobscura B35]

Glycyphana (Glycyphaniola) stolata (Fabricius 1781) [=Cetonia stolata, C. fasciata Fabricius 1775 non Linnaeus 1758; incl. Glycyphana perversa Schaum 1844] CW92

*Type species of generic name indicated

REFERENCES

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d'Urville. Faune entomologique de l'océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[CW92] Cassis, G., & T. A. Weir. 1992. Cetoniinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 426–454. AGPS Press: Canberra.

[G89] Gestro, R. 1889. Sopra alcune cetonie dell’isola Nias e della costa occidentale di Sumatra raccolte dal Dott. Elio Modigliani. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 93–99.

[Mac86] Macleay, W. 1886. The insects of the Fly River, New Guinea, "Coleoptera". Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 136–157.

[Mas86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part III. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 21–126.

Lomapterina

Lomaptera sp., copyright Guido Bohne.

Belongs within: Cetoniinae.

The Lomapterina are an Indo-Australian group of flower beetles in which the basomedian lobe of the pronotum is expanded to entirely or almost entirely cover the scutellum. Many are a bright metallic green in coloration though members of Lomaptera subgenus Melanoptera are brown or blackish (Golding 2009).

<==Lomapterina G09
    |--Agestrata G09
    |--Mycterophalus duboulayi (Thomson 1878) G09
    |--Ischiopsopha Gestro 1874 G09
    |    |--*I. bifasciata (Quoy & Gaimand 1824) [=Cetonia bifasciata] CW92
    |    |--I. bourkei Blackburn 1895 CW92
    |    |--I. deyrollei Thomson 1878 CW92
    |    |--I. erratica Krikken 1983 CW92
    |    |--I. pulchripes (Thomson 1877) [=Lomaptera pulchripes] CW92
    |    |--I. scheini Schürhoff 1942 CW92
    |    `--I. wallacei (Thomson 1857) [=Lomaptera wallacei] CW92
    |         |--I. w. wallacei CW92
    |         `--I. w. wallasiana (Thomson 1860) CW92 (see below for synonymy)
    `--Lomaptera Gory & Percheron 1833 CW92
         |  i. s.: L. acanthopyga Lea 1914 CW92
         |         L. aurata Gestro 1879 CW92
         |         L. diaphonia Kraatz 1880 CW92
         |           |--L. d. diaphonia CW92
         |           `--L. d. nigripes Kraatz 1880 CW92
         |         L. inermis M86
         |         L. macrosticta Lea 1906 CW92
         |         L. pygmaea Kraatz 1880 CW92
         |--L. (Lomaptera) G09
         |    |--*L. (L.) bivittata Gory & Percheron 1833 CW92
         |    `--L. (L.) australis Wallace 1867 G09
         `--L. (Melanoptera) G09
              |--L. (M.) cinnamomea Thomson 1878 G09
              `--L. (M.) hackeri Lea 1906 G09

Ischiopsopha wallacei wallasiana (Thomson 1860) CW92 [=Lomaptera wallasiana CW92; incl. L. yorkiana Janson 1877 CW92, Ischiopsopha yorkiana C91]

*Type species of generic name indicated

REFERENCES

[CW92] Cassis, G., & T. A. Weir. 1992. Cetoniinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 426–454. AGPS Press: Canberra.

[C91] CSIRO. 1991. The Insects of Australia: A textbook for students and research workers vol. 1. Melbourne University Press: Carlton (Victoria).

[G09] Golding, M. R. 2009. A Pictorial Field Guide to the Beetles of Australia. Part 5: Cetoniidae. Privately published.

[M86] Macleay, W. 1886. The insects of the Fly River, New Guinea, "Coleoptera". Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 136–157.

Last updated: 10 April 2022.

Diphucephala

Diphucephala sp., copyright Jean & Fred Hort.

Belongs within: Melolonthinae.

Diphucephala is an Australian genus of diurnal, metallic-coloured chafers often seen congregating on flowers.

Characters (from Macleay 1886): Mentum flat or convex, narrowed at the base, slightly rounded, truncate or slightly emarginate in front. Maxillae robust, the external lobe armed with five or six teeth. Palpi very short, the last joint of the labial obconic, that of the maxillary oval, more or less thick. Head moderately long, flat; clypeus large; nearly square, separated from the forehead by a fine suture, varying in the sexes, emarginate in front. Eyes moderate. Antennae short, of eight joints: first long, swelled at the apex, second nearly as long, obconic, third short, fourth elongate and compressed, fifth very short, the three last forming an oblong mass in males, oval in females. Thorax of varying length, more or less angular on sides, sinuate or truncate at base, often grooved or foveate on disk. Scutellum of curvilinear triangular form, very sharp at extremity. Elytra of an oblong square form, rounded with a perpendicular border at apex. Legs rather long; anterior tibiae bi- or unidentate at their extremities without terminal spur, the four posterior ciliated or velvety; tarsi of variable length, the three or four first joints of the anterior and two or three first of the intermediate dilated in the males and clothed with long hairs, claws robust, cleft at the summit. Pygidium perpendicular, of variable form and size.

<==Diphucephala Dejean 1821 HW92
    |--*D. sericea (Kirby 1818) HW92 [=Melolontha sericea HW92; incl. D. viridis Sturm 1826 Mas86]
    |--D. affinis Waterhouse 1837 HW92
    |--D. angusticeps Macleay 1886 HW92
    |--D. aurolimbata Blanchard 1850 HW92
    |--D. aurulenta (Kirby 1818) [=Melolontha aurulenta; incl. D. foveolata Boisduval 1835] HW92
    |--D. azureipennis Macleay 1886 HW92
    |--D. barnardi Macleay 1886 HW92
    |--D. beryllina Burmeister 1855 HW92
    |--D. carteri Blackburn 1906 HW92
    |--D. castanoptera Waterhouse 1837 HW92
    |--D. childrenii Waterhouse 1837 HW92
    |--D. coerulea Macleay 1883 HW92
    |--D. colaspidoides (Gyllenhall in Schönherr 1817) (see below for synonymy) HW92
    |--D. concinna Lea 1930 HW92
    |--D. crebra Blackburn 1906 HW92
    |--D. cribripennis Lea 1924 HW92
    |--D. cuprea Macleay 1886 HW92
    |--D. dentipes Oke 1951 HW92
    |--D. dicksoniae Lea 1930 HW92
    |--D. edwardsii Waterhouse 1837 HW92
    |--D. elegans Blackburn 1892 HW92
    |--D. fulgida Boisduval 1835 HW92
    |--D. furcata Guérin-Méneville 1830 [incl. D. acanthopus Boisduval 1835] HW92
    |--D. glabra Lea 1930 HW92
    |--D. hirtipennis Macleay 1883 HW92
    |--D. hirtipes Lea 1916 HW92
    |--D. hopei Waterhouse 1837 HW92
    |--D. humeralis Macleay 1886 HW92
    |--D. ignota Macleay 1886 HW92
    |--D. insularis Lea 1916 HW92
    |--D. lateralis Macleay 1886 HW92
    |--D. laticeps Macleay 1886 HW92
    |--D. laticollis Lea 1895 HW92
    |--D. latipennis Macleay 1883 HW92
    |--D. lineata Boisduval 1835 [incl. D. pilistriata Waterhouse 1837] HW92
    |--D. lineatocollis B35
    |--D. macrops Lea 1924 HW92
    |--D. mastersi Macleay 1886 HW92
    |--D. minima Macleay 1886 HW92
    |--D. montana Lea 1930 HW92
    |--D. nigritarsis Lea 1917 HW92
    |--D. nitens Macleay 1886 HW92
    |--D. nitidicollis Macleay 1886 HW92
    |--D. obscura Macleay 1886 HW92
    |--D. obsoleta Macleay 1886 HW92
    |--D. parviceps Lea 1924 HW92
    |--D. parvula Waterhouse 1837 HW92
    |--D. prasina Macleay 1886 [incl. D. kershawi Blackburn 1897] HW92
    |--D. puberula Blackburn 1906 HW92
    |--D. pubescens Macleay 1886 HW92
    |--D. pubiventris Burmeister 1855 HW92
    |--D. pulchella Waterhouse 1837 (see below for synonymy) HW92
    |--D. pulcherrima Blackburn 1906 HW92
    |--D. purpureitarsis Macleay 1886 HW92
    |--D. pygidialis Lea 1916 HW92
    |--D. pygmaea Waterhouse 1837 HW92
    |--D. quadratigera Blanchard 1850 HW92
    |--D. rectipennis Blackburn 1906 HW92
    |--D. regalis Lea 1917 HW92
    |--D. richmondia Macleay 1886 HW92
    |--D. rufipes Waterhouse 1837 HW92
    |--D. rugosa Boisduval 1835 HW92 [incl. D. aenea Sturm 1826 Mas86, D. spencii Waterhouse 1837 HW92]
    |--D. rugosula Dalla Torre 1912 [=D. rugosa Macleay 1886 non Boisduval 1835] HW92
    |--D. smaragdula Boisduval 1835 HW92
    |--D. sordida Blackburn 1906 HW92
    |--D. spreta Blackburn 1892 HW92
    |--D. tantilla Lea 1917 HW92
    |--D. tarsalis Lea 1916 HW92
    `--D. waterhousei Burmeister 1855 HW92

Nomen nudum: Diphucephala effulgens Macleay 1886 Mac86

Diphucephala colaspidoides (Gyllenhall in Schönherr 1817) [=Melolontha colaspidoides; incl. D. lineatocolis Boisduval 1835, D. splendens Macleay 1826] HW92

Diphucephala pulchella Waterhouse 1837 [incl. D. pusilla Waterhouse 1837, D. smaragdula Burmeister 1855 non Boisduval 1835] HW92

*Type species of generic name indicated

REFERENCES

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d'Urville. Faune entomologique de l'océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[HW92] Houston, W. W. K., & T. A. Weir. 1992. Melolonthinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 174–358. AGPS Press: Canberra.

[Mac86] Macleay, W. 1886. Miscellanea entomologica. No. I.—The genus Diphucephala. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 381–402.

[Mas86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part III. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 21–126.

Aplopsis

Aplopsis sp., copyright Liz MacRaild.

Belongs within: Melolonthinae.

Aplopsis is an Australian genus of chafers with the body clothed by pale setae arising on the elytra from elongate, annulate punctures.

Characters (from Britton 1957, as Haplopsis): Anterior tibiae with basal tooth on outer edge situated on proximal half of tibia, the tooth small with its point directed towards apex of tibia; antennae with six to nine segments including a club of three lamellae. Body clothed with pale setae which are usually slender and pointed. Setae very regularly distributed except on elytra where they are arranged in longitudinal bands, with relatively narrow bare spaces between. Setae arising from characteristic elongate annulate punctures. Anterior edge of pronotal hypomera usually marked by a ridge or produced as a free edge enclosing a pocket for reception of antenna. Abdomen usually with a longitudinal ridge on sternites on each side, corresponding with outer edges of elytra; tergite and sternite of propygidium fused. Legs slender, posterior tibiae with or without a slight transverse setose ridge on upper side.

<==Aplopsis Blanchard 1850 (see below for synonymy) HW92
    |--*A. lineoligera Blanchard 1850 HW92 (see below for synonymy)
    |--A. ablata Britton 1957 [=Haplopsis ablata] HW92
    |--A. borealis Britton 1957 [=Haplopsis borealis] HW92
    |--A. caviceps (Lea 1917) [=Caulobius caviceps] HW92
    |--A. darlingtoni Britton 1957 [=Haplopsis darlingtoni] HW92
    |--A. debilis (Blackburn 1898) [=Haplopsis debilis] HW92
    |--A. decipiens (Burmeister 1855) [=*Ocnodus decipiens; incl. Haplopsis polita Britton 1957] HW92
    |--A. discedens (Blackburn 1898) [=Caulobius discedens] HW92
    |--A. evexa (Britton 1957) [=Haplopsis evexa] HW92
    |--A. ferruginea (Blackburn 1892) [=Ocnodus ferrugineus] HW92
    |--A. grisea (Burmeister 1855) HW92 [=Colpocrania grisea HW92, Haplopsis grisea L19]
    |--A. hamilla Britton 1957 [=Haplopsis hamilla] CW92
    |--A. henryi Britton 1957 [=Haplopsis henryi] CW92
    |--A. longipes (Blackburn 1907) [=*Atermonocheila longipes, Haplopsis longipes] HW92
    |--A. mayae Britton 1957 [=Haplopsis mayae] CW92
    |--A. olliffi Blackburn 1898 [=Haplopsis olliffi] CW92
    |--A. punctulata (Blackburn 1898) [=Caulobius punctulatus] CW92
    |--A. rutila Britton 1957 [=Haplopsis rutila] CW92
    |--A. serricollis Lea 1919 [=Haplopsis serricollis] CW92
    |--A. setosa Britton 1957 [=Haplopsis setosa] CW92
    |--A. spadix Britton 1957 [=Haplopsis spadix] CW92
    |--A. viridis Blackburn 1892 [=Haplopsis viridis] CW92
    `--A. wilsoni Britton 1957 [=Haplopsis wilsoni] CW92

Aplopsis Blanchard 1850 [=Haplopsis Gemminger & Harold 1869; incl. Atermonocheila Blackburn 1907, Colpocrania Burmeister 1855, Ocnodus Burmeister 1855] HW92

*Aplopsis lineoligera Blanchard 1850 HW92 [=Haplopsis lineoligera M86; incl. Colpocrania pilosa Burmeister 1855 HW92, H. pilosa M86]

*Type species of generic name indicated

REFERENCES

Britton, E. B. 1957. A Revision of the Australian Chafers (Coleoptera: Scarabaeidae: Melolonthinae) vol. 1. British Museum (Natural History): London.

[HW92] Houston, W. W. K., & T. A. Weir. 1992. Melolonthinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 174–358. AGPS Press: Canberra.

[L19] Lea, A. M. 1919. Notes on some miscellaneous Coleoptera with descriptions of new species.—Part V. Transactions of the Royal Society of South Australia 43: 166–261, pls 25–27.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part III. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 21–126.

Automolius

Automolius sp., copyright Jackie Miles.

Belongs within: Melolonthinae.

Automolius is an Australian genus of chafers in which the elytra are shortened, leaving at least part of the last two abdominal segments exposed.

Characters (from Britton 1957, as Automolus): Antennae seven-, eight- or nine-segmented, including a three-segmented club; elytra more or less shortened, not reaching pygidium, part or whole of propygidium exposed; abdomen without a longitudinal ridge on each side; tergite and ventrite of propygidium fused at sides; whole body clothed with rather regularly arranged semierect setae; female usually with a bare shining area on each elytron near apex on outer side; posterior tibia rather slender, moderately expanded to apex with a slight setose ridge on upper side.

<==Automolius Britton 1978 [=Automolus Burmeister 1855 non Reichenbach 1853] HW92
    |--*A. angustulus (Burmeister 1855) (see below for synonymy) HW92
    |--A. brooksi (Britton 1957) [=Automolus brooksi] HW92
    |--A. brunneus (Lea 1917) [=Automolus brunneus] HW92
    |--A. compactus (Blackburn 1898) [=Caulobius compactus] HW92
    |--A. cookii (Macleay 1886) [=Liparetrus cookii] HW92
    |--A. depressus (Blanchard 1850) (see below for synonymy) HW92
    |--A. evanescens (Blackburn 1898) [=Caulobius evanescens] HW92
    |--A. fulgineus (Britton 1957) [=Automolus fulgineus] HW92
    |--A. funereus (Blackburn 1906) [=Automolus funereus] HW92
    |--A. granulatus (Blackburn 1888) [=Liparetrus granulatus] HW92
    |--A. hispidus (Macleay 1886) [=Liparetrus hispidus; incl. L. aureus Blackburn 1888] HW92
    |--A. humilis (Blanchard 1850) (see below for synonymy) HW92
    |--A. immitis (Blackburn 1906) [=Caulobius immitis] HW92
    |--A. irrasus (Blackburn 1906) [=Automolus irrasus] HW92
    |--A. maechidioides (Macleay 1886) [=Liparetrus maechidioides] HW92
    |--A. major (Blackburn 1906) [=Automolus major] HW92
    |--A. nanus (Britton 1957) [=Automolus nanus] HW92
    |--A. opaculus (Blackburn 1906) [=Automolus opaculus] HW92
    |--A. ordinatus (Macleay 1886) [=Liparetrus ordinatus] HW92
    |--A. phoxus (Britton 1957) [=Automolus phoxus] HW92
    |--A. pictus (Blackburn 1906) [=Automolus pictus] HW92
    |--A. poverus (Blanchard 1850) [=Liparetrus poverus; incl. L. striatipennis Macleay 1886] HW92
    |--A. pygmaeus (Burmeister 1855) [=Liparetrus pygmaeus] HW92
    |--A. rotundus (Blackburn 1906) [=Caulobius rotundus] HW92
    |--A. semitifer (Blackburn 1906) [=Automolus semitifer] HW92
    |--A. sordidus (Britton 1957) [=Automolus sordidus] HW92
    |--A. sulcatus (Britton 1957) [=Automolus sulcatus] HW92
    |--A. tridentifrons (Lea 1917) [=Automolus tridentifrons] HW92
    |--A. valgoides (Blanchard 1850) [=Liparetrus valgoides] HW92
    `--A. vulgaris (Britton 1957) [=Automolus vulgaris] HW92

*Automolius angustulus (Burmeister 1855) [=*Automolus angustulus; incl. Liparetrus alpicola Blackburn 1892, L. burmeisteri Macleay 1886] HW92

Automolius depressus (Blanchard 1850) [=Liparetrus depressus; incl. L. bicolor Blackburn 1888, Automolus melancholicus Lea 1917] HW92

Automolius humilis (Blanchard 1850) [=Liparetrus humilis; incl. L. concolor Macleay 1864 non Erichson 1842, Automolus propygidialis Lea 1917, L. unicolor Masters 1871] HW92

*Type species of generic name indicated

REFERENCES

Britton, E. B. 1957. A Revision of the Australian Chafers (Coleoptera: Scarabaeidae: Melolonthinae) vol. 1. British Museum (Natural History): London.

[HW92] Houston, W. W. K., & T. A. Weir. 1992. Melolonthinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 174–358. AGPS Press: Canberra.

Phyllotocus

Phyllotocus macleayi caught in a spider web, copyright Mike Gordon.

Belongs within: Melolonthinae.

Phyllotocus is an Australian genus of often relatively brightly coloured chafer beetles.

Characters (from Britton 1957): Labrum transverse, clearly separated from the clypeus by a suture; clypeus reflexed at sides, without a transverse ridge, the general surface of clypeus and frons slightly convex; antennae eight- or nine-segmented, including a three-segmented club; tarsal claws slightly curved, slender, simple, inner claw of anterior tarsus usually enlarged in male; posterior angle of pronotum obtuse or rounded, not closely applied to shoulders of the elytra; maxillae with lacinia rounded and setose; prosternum with a long slender process in the middle just behind anterior coxae, process with strong spines at apex.

<==Phyllotocus Fischer 1823 [incl. Macrothops Boisduval 1835, Macrotops (l. c.)] HW92
    |--*P. macleayi Fischer von Waldheim 1823 HW92, A06
    |    |--P. m. macleayi HW92 (see below for synonymy)
    |    `--P. m. pallidus Lea 1919 HW92
    |--P. apicalis Macleay 1864 [incl. P. navicularis erythroderes Lea 1919] HW92
    |--P. armstrongi Britton 1957 HW92
    |--P. australis (Boisduval 1835) (see below for synonymy) HW92
    |--P. basalis Lea 1919 [=P. bimaculatus basalis; incl. P. bimaculatus insularis Lea 1919] HW92
    |--P. bimaculatus Erichson 1842 HW92
    |--P. chapmani Britton 1957 HW92
    |--P. darlingtoni Britton 1957 HW92
    |--P. decipiens Lea 1919 HW92
    |--P. erythropterus Blanchard 1850 HW92
    |--P. gilvus Britton 1957 HW92
    |--P. giveni Britton 1957 HW92
    |--P. kingii Macleay 1864 HW92
    |--P. laterofuscus Lea 1919 HW92
    |--P. lottinii (Boisduval 1835) M86, HW92 [=Macrothops lottinii HW92]
    |--P. luridus Britton 1957 non Cheiragra lurida Macleay 1864 (not preoc. if in dif. gen.) HW92
    |--P. marginalis Bohem. 1858 M86
    |--P. marginipennis Macleay 1864 [incl. P. marginatus Macleay 1864] HW92
    |--P. melanurus Britton 1957 HW92
    |--P. meyricki Blackburn 1888 HW92
    |--P. moestus (Boisduval 1835) (see below for synonymy) HW92
    |--P. navicularis Blanchard 1850 (see below for synonymy) HW92
    |--P. nigripennis Lea 1919 [=P. bimaculatus nigripennis] HW92
    |--P. occidentalis Blackburn 1888 [incl. P. occidentalis apicifuscus Lea 1919] HW92
    |--P. pelus Britton 1957 HW92
    |--P. ruficollis Macleay 1864 [incl. P. sericeus Macleay 1871] HW92
    |--P. rufipennis (Boisduval 1835) [=Macrothops rufipennis] HW92
    |--P. scutellaris Macleay 1864 HW92
    |--P. ustulatus Blanchard 1850 HW92
    `--P. variicollis Macleay 1871 HW92

Phyllotocus australis (Boisduval 1835) [=Macrothops australis; incl. P. discoidalis Burmeister 1855, P. iridescens Macleay 1864, P. palliatus Macleay 1864] HW92

*Phyllotocus macleayi macleayi Fischer 1823 HW92 [incl. P. assimilis Macleay 1864 HW92, Macrothops praeusta Boisduval 1835 HW92, P. praeustus M86]

Phyllotocus moestus (Boisduval 1835) [=Macrothops moesta; incl. P. marginicollis Boheman 1858, P. oblongus Boheman 1858, P. velutinus Boheman 1858] HW92

Phyllotocus navicularis Blanchard 1850 [incl. P. navicularis apicalis Lea 1919 non Macleay 1864, P. navicularis rufibasis Lea 1919] HW92

*Type species of generic name indicated

REFERENCES

[A06] Ahrens, D. 2006. The phylogeny of Sericini and their position within the Scarabaeidae based on morphological characters (Coleoptera: Scarabaeidae). Systematic Entomology 31: 113–144.

Britton, E. B. 1957. A Revision of the Australian Chafers (Coleoptera: Scarabaeidae: Melolonthinae) vol. 1. British Museum (Natural History): London.

[HW92] Houston, W. W. K., & T. A. Weir. 1992. Melolonthinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 174–358. AGPS Press: Canberra.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part III. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 21–126.

Maechidius

Maechidius sp., from Morwell National Park Online.

Belongs within: Melolonthinae.

Maechidius is an Australasian genus of chafers, at least some of which live in association with termite nests.

Characters (from Britton 1957): Two teeth on outer edge of anterior tibia, the basal tooth usually on distal half of tibia, obtuse, not acute and closely applied to the edge; anterior edge of pronotal hypomera produced forwards to form a blade-like free edge enclosing a pocket into which antenna can be withdrawn; edge obtusely angulate when viewed from side; claws with or without pulvilli; antennae seven-, eight- or nine-segmented including a three-segmented club; disc of pronotum and elytra often with characteristic annular setiferous punctures; body form depressed; abdomen with a strong longitudinal ridge on each side; tergite and sternite of propygidium at least partly fused; colour reddish brown to black.

<==Maechidius Macleay 1819 [incl. Geobatus Boisduval 1835] HW92
    |--*M. spurius (Kirby 1818) HW92 (see below for synonymy)
    |--M. algonus Britton 1957 HW92
    |--M. amanus Britton 1957 HW92
    |--M. antennalis Blackburn 1898 HW92
    |--M. ater Waterhouse 1875 HW92
    |--M. atratus Burmeister 1855 HW92
    |--M. bidentulus Fairm. 1877 M86
    |--M. brevis Waterhouse 1875 HW92
    |--M. caesius Britton 1957 HW92
    |--M. calabyi Britton 1957 HW92
    |--M. capitalis Blackburn 1907 HW92
    |--M. caviceps Blackburn 1888 HW92
    |--M. cavus Britton 1957 HW92
    |--M. chadwicki Britton 1957 HW92
    |--M. charaxus Britton 1957 HW92
    |--M. clypealis Blackburn 1894 HW92
    |--M. coelus Britton 1957 HW92
    |--M. conspicuus Lea 1917 HW92
    |--M. corrosus Waterhouse 1875 [incl. M. sculptipennis Lea 1917, M. sexdentatus Waterhouse 1875] HW92
    |--M. crassifrons Britton 1957 HW92
    |--M. crenaticollis Blackkburn 1888 [incl. M. collaris Blackburn 1898] HW92
    |--M. davidsoni Britton 1957 HW92
    |--M. emarginatus Waterhouse 1875 HW92
    |--M. eutermiphilus Lea 1924 HW92
    |--M. excisicollis Blackburn 1898 HW92
    |--M. excisus Waterhouse 1875 HW92
    |--M. fissiceps Macleay 1888 HW92
    |--M. froggatti Macleay 1888 HW92
    |--M. geminus Britton 1957 HW92
    |--M. gibbicollis Blackburn 1898 HW92
    |--M. hackeri Lea 1919 HW92
    |--M. hopianus Westwood 1842 [=M. hopeanus (l. c.); incl. M. obscurus Macleay 1871] HW92
    |--M. hoplocephalus Lea 1917 HW92
    |--M. insularis Lea 1917 HW92
    |--M. kanallus Britton 1957 HW92
    |--M. kiwanus Britton 1957 HW92
    |--M. konandus Britton 1957 HW92
    |--M. kurantus Britton 1957 HW92
    |--M. lateripennis Lea 1917 HW92
    |--M. latus Waterhouse 1875 HW92
    |--M. longitarsus Waterhouse 1875 HW92
    |--M. macleaynus Westwood 1842 (see below for synonymy) HW92
    |--M. major Blackburn 1888 HW92
    |--M. mellianus Westwood 1842 [incl. M. sinuaticeps Blackburn 1889] HW92
    |--M. metellus Britton 1957 HW92
    |--M. milkappus Britton 1957 HW92
    |--M. modicus Blackburn 1898 HW92
    |--M. occidentalis Britton 1957 HW92
    |--M. olor Britton 1957 HW92
    |--M. ordensis Blackburn 1898 HW92
    |--M. parandus Britton 1957 HW92
    |--M. parvulus Macleay 1871 [incl. M. acutangulus Waterhouse 1875] HW92
    |--M. pellus Britton 1957 HW92
    |--M. pilosus Blackburn 1891 HW92
    |--M. proximus Britton 1957 HW92
    |--M. puncticollis Lea 1924 HW92
    |--M. pygidialis Britton 1957 HW92
    |--M. raucus Blackburn 1907 HW92
    |--M. relictus Blackburn 1907 HW92
    |--M. rufus Westwood 1842 HW92
    |--M. rugosicollis Macleay 1871 HW92
    |--M. rugosipes Blackburn 1888 [incl. M. imitator Blackburn 1898] HW92
    |--M. savagei Lea 1917 HW92
    |--M. serratus Britton 1957 HW92
    |--M. sordidus (Boisduval 1835) (see below for synonymy) HW92
    |--M. squamipennis Lea 1917 HW92
    |--M. stradbrokensis Lea 1919 HW92
    |--M. tibialis Blackburn 1892 HW92
    |--M. tridentatus Britton 1963 HW92
    |--M. tumidifrons Britton 1957 HW92
    |--M. variolosus Macleay 1871 HW92
    `--M. wilsoni Britton 1957 HW92

Maechidius macleaynus Westwood 1842 [=M. macleajanus (l. c.), M. macleayanus (l. c.); incl. M. multistriatus Blackburn 1898] HW92

Maechidius sordidus (Boisduval 1835) [=*Geobatus sordidus; incl. M. raddonianus Westwood 1842, M. raddonanus (l. c.)] HW92

*Maechidius spurius (Kirby 1818) HW92 [=Trox spurius HW92; incl. T. australis B35, Maechidius kirbianus Westwood 1845 HW92, M. kirbyanus (l. c.) HW92]

*Type species of generic name indicated

REFERENCES

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d'Urville. Faune entomologique de l'océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

Britton, E. B. 1957. A Revision of the Australian Chafers (Coleoptera: Scarabaeidae: Melolonthinae) vol. 1. British Museum (Natural History): London.

[HW92] Houston, W. W. K., & T. A. Weir. 1992. Melolonthinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 174–358. AGPS Press: Canberra.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part III. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 21–126.

Gnaphalopoda

Gnaphalopoda sp., copyright Dominique Poirier-Ducrocq.

Belongs within: Melolonthinae.

Gnaphalopoda is an Australasian genus of chafers characterised by the presence of dense tufts of setae on the undersides of the tarsi.

<==Gnaphalopoda Reiche 1860 [incl. Xylostygnus Broun 1886] HW92
    |--*G. leptopoda (Montrouzier 1860) [=Rhisotrogus leptopoda] HW92
    |--G. ascia Britton 1987 HW92
    |--G. austrina Britton 1987 HW92
    |--G. bidentata (Lea 1917) [=Ocnodus bidentatus] HW92
    |--G. biloba (Lea 1917) [=Ocnodus bilobus] HW92
    |--G. carnei Britton 1987 HW92
    |--G. crassa Britton 1987 HW92
    |--G. eremia Britton 1987 HW92
    |--G. fallax (Blackburn 1892) [=Ocnodus fallax] HW92
    |--G. frons Britton 1987 HW92
    |--G. lepida (Blackburn 1900) [=Ocnodus lepidus] HW92
    |--G. lesouefi Britton 1987 HW92
    |--G. lirella Britton 1987 HW92
    |--G. lugubris (Blackburn 1892) [=Ocnodus lugubris; incl. O. simplex Lea 1917] HW92
    |--‘*Xylostygnus’ piceus Broun 1886 HW92
    |--G. porosa (Blackburn 1907) [=Ocnodus porosus] HW92
    |--G. proxima Britton 1987 HW92
    |--G. scissiceps (Blackburn 1907) [=Ocnodus scissiceps] HW92
    |--G. solida Britton 1987 HW92
    |--G. spinicollis (Blackburn 1892) [=Ocnodus spinicollis] HW92
    |--G. suspiciosa (Blackburn 1907) [=Ocnodus suspiciosus] HW92
    |--G. tridentata (Lea 1917) [=Ocnodus tridentatus] HW92
    `--G. unidentata (Lea 1917) [=Ocnodus unidentatus] HW92

*Type species of generic name indicated

REFERENCES

[HW92] Houston, W. W. K., & T. A. Weir. 1992. Melolonthinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 174–358. AGPS Press: Canberra.

Heteronyx

Heteronyx sp., from PaDIL.

Belongs within: Melolonthinae.

Heteronyx is a remarkably diverse genus of chafers found in Australia, species of which have toothed tarsal claws.

Characters (from Britton 2000): Small to moderate size (length 3.5–17 mm); tarsal claws with a tooth beneath, sometimes bifid; clypeus not extended forwards and upwards as a reflexed lamina above the labrum; labrum often reflexed upwards and visible before the clypeus as viewed from above; without a vertical median carina in an anterior emargination; maxillae with 5 or 6 inwardly directed, sharp, black teeth on the lacinia; disc of pronotum and elytra usually with a uniform clothing of short, semierect or recumbent setae; antennae 8 or 9-segmented, including an asymmetric club of three very short lamellae; anterior tibiae almost always with 2 teeth on the outer edge in addition to the apical process and never with a minute, acute tooth very close to the base; tarsi without a pulvillus or setose pad between the claws; abdomen almost always with a lateral, longitudinal ridge on each side; elytra usually without longitudinal striae, rarely with more than slightly impressed sutural striae; body colour testaceous, castaneous, piceous or black.

<==Heteronyx Guérin-Méneville 1831 (see below for synonymy) HW92
    |--*H. australis Guérin-Méneville 1831 (see below for synonymy) B00
    |--H. acutifrons Blackburn 1889 HW92
    |--H. additus Blackburn 1909 [incl. H. amoenus Blackburn 1910] B00
    |--H. advena Blackburn 1890 HW92
    |--H. aequaliceps Blackburn 1890 HW92
    |--H. aequalis Blackburn 1889 [incl. H. debilicollis Blackburn 1909, H. taeniensis Blackburn 1909] B00
    |--H. affinis Blackburn 1909 HW92
    |--H. agrestis Burmeister 1855 HW92
    |--H. agricola Blackburn 1910 HW92
    |--H. alienus Blackburn 1892 HW92
    |--H. alpicola Blackburn 1892 HW92
    |--H. ambiguus Blackburn 1910 HW92
    |--H. amboinensis Moser 1920 B00
    |--H. apertus Blackburn 1910 HW92
    |--H. aphodioides Blanchard 1850 [incl. H. incultus Blackburn 1889, H. potens Blackburn 1889] B00
    |--H. apicispinosa Heller 1914 B00
    |--H. arcanus Blackburn 1892 [incl. H. umbrinus Blackburn 1909] B00
    |--H. aridus Blackburn 1889 HW92
    |--H. aruanus Moser 1920 B00
    |--H. asperatus Moser 1920 B00
    |--H. aspericollis Blackburn 1889 HW92
    |--H. asperifrons Blackburn 1909 HW92
    |--H. augustae Blackburn 1889 [incl. H. anceps Blackburn 1889, H. decorus Blackburn 1892] B00
    |--H. badius Macleay 1888 [incl. H. darwini Blackburn 1889, H. macleayi Blackburn 1910] B00
    |--H. beltanae Blackburn 1889 HW92
    |--H. bidentatus Blackburn 1889 HW92
    |--H. blandus Blackburn 1909 HW92
    |--H. bolivianus Moser 1919 B00
    |--H. borealis Blackburn 1889 HW92
    |--H. bovilli Blackburn 1890 HW92
    |--H. brevior Fairmaire 1883 B00
    |--H. calidus Blackburn 1910 HW92
    |--H. callabonnae Blackburn 1909 HW92
    |--H. capillatus Macleay 1888 HW92
    |--H. castanescens Moser 1924 B00
    |--H. castaneus Blanchard 1846 [incl. H. obscurus Blanchard 1846 non Le Guillou 1844] B00
    |--H. cervina (Boisduval 1835) (see below for synonymy) B00
    |--H. chlorotica (Gyllenhal 1817) (see below for synonymy) B00
    |--H. cliens Blackburn 1910 HW92
    |--H. collaris Blackburn 1890 HW92
    |--H. comans Blackburn 1909 HW92
    |--H. concolor Macleay 1871 (see below for synonymy) B00
    |--H. confertus Blackburn 1910 HW92
    |--H. constans Blackburn 1889 [incl. H. yilgarnensis Blackburn 1890] B00
    |--H. corpulentus Macleay 1888 HW92
    |--H. corumbanus Moser 1921 B00
    |--H. costulatus Blackburn 1910 HW92
    |--H. crassus Blackburn 1889 HW92
    |--H. cribriceps Blackburn 1892 HW92
    |--H. crinitus Blackburn 1910 HW92
    |--H. cunnamullae Blackburn 1910 HW92
    |--H. cuyabanus Moser 1919 B00
    |--H. cygneus Blackburn 1889 HW92
    |--H. darlingensis Blackburn 1889 HW92
    |--H. declaratus Blackburn 1909 HW92
    |--H. dentipes Blackburn 1889 HW92
    |--H. dimidiata (Erichson 1842) B00 (see below for synonymy)
    |--H. disjectus Blackburn 1910 [incl. H. nudus Blackburn 1910] B00
    |--H. diversiceps Blackburn 1890 HW92
    |--H. doctus Blackburn 1890 HW92
    |--H. doddi Blackburn 1908 HW92
    |--H. electus Blackburn 1889 HW92
    |--H. elongatus Blanchard 1850 [incl. H. monticola Blackburn 1909] B00
    |--H. elytrurus Blackburn 1909 HW92
    |--H. excisus Blackburn 1890 B00
    |--H. ‘excisa’ Heller 1912 non H. excisus Blackburn 1890 B00
    |--H. exectus Blackburn 1909 HW92
    |--H. fervidus Blackburn 1892 HW92
    |--H. fimbriatus Moser 1924 [=H. ciliatus Moser 1920 (preoc.)] B00
    |--H. flavus Blackburn 1890 [incl. H. angustus Blackburn 1890] B00
    |--H. fortis Blackburn 1889 HW92
    |--H. fraserensis Blackburn 1892 HW92
    |--H. frenchi Blackburn 1909 HW92
    |--H. froggatti Macleay 1888 HW92
    |--H. frontalis Blackburn 1889 [incl. H. nitidus Blackburn 1890] B00
    |--H. fulvohirtus Blackburn 1889 (see below for synonymy) B00
    |--H. fumata (Erichson 1842) (see below for synonymy) B00
    |--H. furvus Blackburn 1892 HW92
    |--H. glabrata (Erichson 1842) [=Silopa glabrata, H. glabratus] B00
    |--H. grandis Blackburn 1900 HW92
    |--H. granulatus Blackburn 1910 HW92
    |--H. granulifer Blackburn 1889 HW92
    |--H. granum Burmeister 1855 HW92
    |--H. helmsi Blackburn 1892 [incl. H. waterhousei Blackburn 1910] B00
    |--H. heynei Moser 1919 B00
    |--H. hirsutosetosus Moser 1920 B00
    |--H. hirsutus Blackburn 1910 HW92
    |--H. hirtuosus Blackburn 1890 HW92
    |--H. hispidulus Blackburn 1908 HW92
    |--H. horridus Blackburn 1889 HW92
    |--H. humilis Blackburn 1910 HW92
    |--H. imitator Blackburn 1909 HW92
    |--H. impar Blackburn 1910 [incl. H. convexicollis Blackburn 1910] B00
    |--H. incognitus Blackburn 1892 [incl. H. punctipes Blackburn 1909] B00
    |--H. incola Blackburn 1889 HW92
    |--H. infirmus Blackburn 1910 HW92
    |--H. infuscatus Macleay 1871 (see below for synonymy) B00
    |--H. inornatus (Blackburn 1900) [=*Anacheirotus inornatus] HW92
    |--H. insignis Blackburn 1889 HW92
    |--H. insularis Fairmaire 1883 B00
    |--H. interioris Blackburn 1909 HW92
    |--H. intrusus Blackburn 1910 HW92
    |--H. iridiventris Blackburn 1890 HW92
    |--H. irrasus Lea 1926 HW92
    |--H. jejunus Blackburn 1889 [incl. H. longulus Blackburn 1890, H. lubricus Blackburn 1890] B00
    |--H. johannis Blackburn 1912 HW92
    |--H. labratus Moser 1926 B00
    |--H. laeviceps Blackburn 1889 HW92
    |--H. laminatus Blackburn 1890 HW92
    |--H. lateritius Blackburn 1889 [incl. H. validus Blackburn 1910] B00
    |--H. laticeps Burmeister 1855 [incl. H. coatesi Blackburn 1909, H. holosericeus Macleay 1871] B00
    |--H. lilliputanus Blackburn 1890 HW92
    |--H. lindi Blackburn 1889 [incl. H. submetallicus Blackburn 1889] HW92
    |--H. lividus Blackburn 1889 HW92
    |--H. lobatus Blackburn 1889 [incl. H. viator Blackburn 1890] B00
    |--H. luteolus Blackburn 1909 HW92
    |--H. maculatus Blackburn 1889 HW92
    |--H. magnus Moser 1924 B00
    |--H. major Blackburn 1910 [incl. H. moestus Blackburn 1910] B00
    |--H. maluensis Brenske 1889 B00
    |--H. marcidus Blackburn 1892 HW92
    |--H. marginatus Blackburn 1890 HW92
    |--H. merus Blackburn 1892 [incl. H. oodnadattae Blackburn 1909] B00
    |--H. metropolitanus Blackburn 1909 HW92
    |--H. mimus Blackburn 1890 HW92
    |--H. minimus Lea 1926 HW92
    |--H. minutus Blackburn 1910 HW92
    |--H. moluccanus Moser 1920 B00
    |--H. montanus Blackburn 1890 [incl. H. miser Blackburn 1910, H. ordinarius Blackburn 1910] B00
    |--H. mulwalensis Blackburn 1889 HW92
    |--H. mundus Blackburn 1909 HW92
    |--H. nasutus Blackburn 1889 [incl. H. tridens Blackburn 1892] B00
    |--H. neglectus Blackburn 1910 HW92
    |--H. nigella (Erichson 1842) B00 (see below for synonymy)
    |--H. nigrescens Blackburn 1909 [incl. H. nubilus Blackburn 1909] B00
    |--H. nigricans Burmeister 1855 [incl. H. comes Blackburn 1910] B00
    |--H. nigritulus Moser 1926 B00
    |--H. nigrita Blanchard 1850 B00 (see below for synonymy)
    |--H. normalis Blackburn 1889 HW92
    |--H. novitius Blackburn 1910 HW92
    |--H. obesa Burmeister 1855 [=H. obesus; incl. H. occidentalis Blackburn 1889] B00
    |--H. oblongus Blanchard 1850 (see below for synonymy) B00
    |--H. ohausiana Saylor 1938 B00
    |--H. orbus Blackburn 1909 HW92
    |--H. ovatus Blanchard 1850 [incl. H. inconspicuus Blackburn 1910] B00
    |--H. pallidulus Macleay 1871 HW92
    |--H. papuanus Moser 1920 B00
    |--H. parvulus Macleay 1888 HW92
    |--H. pauxillus Blackburn 1910 HW92
    |--H. pellucida Burmeister 1855 [incl. H. distortus Lea 1926, H. scutatus Macleay 1888] B00
    |--H. peregrinus Blackburn 1890 [incl. H. macilentus Blackburn 1910] B00
    |--H. perkinsi Blackburn 1909 HW92
    |--H. piceoniger Macleay 1888 HW92
    |--H. piceus Blanchard 1850 [incl. H. ponderosus Blackburn 1909, H. spissus Blackburn 1909] B00
    |--H. pilosellus Blanchard 1850 [incl. H. copiosus Blackburn 1909] B00
    |--H. pilosus Blanchard 1846 HW92
    |--H. placidus Blackburn 1910 HW92
    |--H. planata Burmeister 1855 (n. d.) B00
    |--H. planiceps Blackburn 1910 HW92
    |--H. posticalis Blackburn 1890 [incl. H. approximans Blackburn 1910] B00
    |--H. praecox (Erichson 1842) B00 (see below for synonymy)
    |--H. proditor Blackburn 1892 HW92
    |--H. proprius Blackburn 1910 [incl. H. austrinus Blackburn 1910] B00
    |--H. prosper Blackburn 1909 HW92
    |--H. protervus Blackburn 1892 [incl. H. tropicus Blackburn 1909] B00
    |--H. proxima Burmeister 1855 B00 [=H. proximus HW92]
    |--H. pubescens (Erichson 1842) (see below for synonymy) B00
    |--H. pumilus Sharp 1877 B00 [incl. H. brevicornis Blackburn 1889 B00, HW92]
    |--H. punctatissimus Montrouzier 1857 B00
    |--H. puncticollis Blackburn 1890 HW92
    |--H. punctipennis Blackburn 1889 [incl. H. cribripennis Blackburn 1912] B00
    |--H. pustulosus Blackburn 1890 HW92
    |--H. pygidialis Blackburn 1889 HW92
    |--H. pygmaeus Blackburn 1910 HW92
    |--H. quadraticollis Blackburn 1890 HW92
    |--H. queenslandicus Blackburn 1909 HW92
    |--H. randalli Blackburn 1890 [incl. H. femoralis Blackburn 1909, H. severus Blackburn 1892] B00
    |--H. rectangulus Blackburn 1910 HW92
    |--H. relictus Blackburn 1909 [incl. H. mildurensis Blackburn 1910] B00
    |--H. rhinastus Blackburn 1890 HW92
    |--H. rhinoceros Blackburn 1892 HW92
    |--H. rothei Blackburn 1890 HW92
    |--H. rotundiceps Blanchard 1850 [incl. H. debilis Blackburn 1889] B00
    |--H. rubescens Blanchard 1850 HW92
    |--H. ruficollis Macleay 1871 HW92
    |--H. rofumarginatus Blanchard 1850 HW92
    |--H. rufopiceus Macleay 1888 [incl. H. breviceps Blackburn 1889] HW92
    |--H. rugosipennis Macleay 1871 [incl. H. dux Blackburn 1909, H. incomptus Blackburn 1909] B00
    |--H. rusticus Blackburn 1890 HW92
    |--H. salebrosus Blackburn 1909 HW92
    |--H. satelles Blackburn 1889 HW92
    |--H. scalptus Blackburn 1890 (see below for synonymy) B00
    |--H. schenklingi Moser 1919 B00
    |--H. sequens Blackburn 1894 [incl. H. ignobilis Blackburn 1910, H. sollicitus Blackburn 1910] B00
    |--H. setifer Blackburn 1890 HW92
    |--H. sexualis Blackburn 1909 HW92
    |--H. siccus Blackburn 1892 HW92
    |--H. simius Blackburn 1890 HW92
    |--H. simplicicollis Blackburn 1909 HW92
    |--H. simulator Blackburn 1889 [incl. H. intermedius Blackburn 1909, H. thoracicus Blackburn 1909] B00
    |--H. sloanei Blackburn 1889 HW92
    |--H. solidus Blackburn 1889 HW92
    |--H. spadicea Burmeister 1855 [=H. spadiceus] B00
    |--H. spretus Blackburn 1889 HW92
    |--H. squalidus Blackburn 1908 HW92
    |--H. striatus Blackburn 1909 [incl. H. seriatus Blackburn 1909] B00
    |--H. suavis Blackburn 1909 HW92
    |--H. subferruginea Burmeister 1855 B00 [=H. subferrugineus HW92]
    |--H. subfuscus Macleay 1888 (see below for synonymy) B00
    |--H. subglaber Macleay 1888 HW92
    |--H. substriatus Macleay 1871 HW92
    |--H. sulcifrons Blackburn 1909 HW92
    |--H. sydneyanus Blackburn 1890 HW92
    |--H. tarsalis Blackburn 1909 [incl. H. griffithi Blackburn 1909] B00
    |--H. tasmanicus Blackburn 1909 [incl. H. quaesitus Blackburn 1909] B00
    |--H. tenebrosus Blackburn 1909 HW92
    |--H. tepperi Blackburn 1889 [incl. H. subfortis Blackburn 1908] B00
    |--H. terrena Blackburn 1892 (see below for synonymy) B00
    |--H. tindalei Britton in Houston & Weir 1992 [=H. insularis Lea 1926 non Fairmaire 1883] HW92
    |--H. torvus Blackburn 1889 HW92
    |--H. transversicollis Macleay 1888 HW92
    |--H. tridentatus Lea 1924 HW92
    |--H. tristis Blackburn 1889 HW92
    |--H. umbilicatus Fauvel 1914 B00
    |--H. unicolor Blanchard 1850 [incl. H. auricomus Blackburn 1889, H. campestris Blackburn 1910] B00
    |--H. unicus Blackburn 1900 HW92
    |--H. variegatus Blackburn 1889 HW92
    |--H. vicinus Blackburn 1910 HW92
    |--H. victoris Blackburn 1889 [incl. H. socius Blackburn 1909] B00
    |--H. viduus Blackburn 1910 [incl. H. glaber Lea 1926, H. modestus Blackburn 1910] B00
    `--H. zalotus Blackburn 1910 HW92

Heteronyx Guérin-Méneville 1831 [incl. Anacheirotus Blackburn 1900, Cotidia Boisduval 1835, Hostilina Blanchard 1850, Silopa Erichson 1842] HW92

*Heteronyx australis Guérin-Méneville 1831 [=H. castaneus Macleay 1871 non Blanchard 1846, H. conjunctus Blackburn 1910, H. erichsoni Blackburn 1910, H. farinensis Blackburn 1910, H. hackeri Blackburn 1910, H. olliffi Blackburn 1910, H. pseudocastaneus Houston & Weir 1992, H. pubescens Macleay 1871 non Silopa pubescens Erichson 1842, H. vagans Blackburn 1890] B00

Heteronyx cervina (Boisduval 1835) [=Sericesthis cervina, H. cervinus; incl. Silopa tempestiva Erichson 1842, H. tempestivus] B00

Heteronyx chlorotica (Gyllenhal 1817) [=Melolontha chlorotica, Scitala chlorotica; incl. H. fissiceps Blackburn 1890] B00

Heteronyx concolor Macleay 1871 [incl. H. cowelli Blackburn 1894, H. oscillator Blackburn 1890, H. pauper Blackburn 1910, H. puer Blackburn 1910, H. sparsus Blackburn 1889, H. suturalis Blackburn 1895] B00

Heteronyx dimidiata (Erichson 1842) B00 [=Silopa dimidiata HW92; incl. *Cotidia australis Boisduval 1835 non Heteronyx australis Guérin-Méneville 1831 HW92, B00, HW92, Heteronyx fallax Blackburn 1890 B00, H. fraternus Blackburn 1890 B00, H. jubatus Blackburn 1890 B00, H. striatipennis Blanchard 1850 B00]

Heteronyx fulvohirtus Blackburn 1889 [incl. H. dispar Blackburn 1908, H. litigiosus Blackburn 1908, H. rudis Blackburn 1908] B00

Heteronyx fumata (Erichson 1842) [=Silopa fumata; incl. S. hepatica Erichson 1842, Heteronyx laticollis Blanchard 1850, H. rapax Blackburn 1890, H. rubriceps Blanchard 1850] B00

Heteronyx infuscatus Macleay 1871 [incl. H. fictus Blackburn 1910, H. lucidus Blackburn 1910, H. rotundifrons Blackburn 1889] B00

Heteronyx nigella (Erichson 1842) B00 [=Silopa nigella B00; incl. H. holomelaena Blanchard 1850 B00, H. holomelaenus HW92, H. maurulus Blackburn 1910 B00, H. obscurus Le Guillou 1844 B00, H. pinguis Blackburn 1890 B00]

Heteronyx nigrita Blanchard 1850 B00 [=H. nigritus HW92; incl. H. nigrinus Blackburn 1890 B00, H. vacuus Blackburn 1890 B00]

Heteronyx oblongus Blanchard 1850 [incl. H. consanguineus Blackburn 1892, H. dubius Blackburn 1899, H. piger Blackburn 1889, H. raucinasus Blackburn 1889] B00

Heteronyx praecox (Erichson 1842) B00 [=Silopa praecox B00, *Hostilina praecox HW92; incl. H. carpentariae M86, H. testaceus Blackburn 1889 B00]

Heteronyx pubescens (Erichson 1842) [=Silopa pubescens; incl. H. carteri Blackburn 1909, H. deceptor Blackburn 1890, H. gracilipes Blackburn 1889] B00

Heteronyx scalptus Blackburn 1890 [=H. sculptus (l. c.); incl. H. difficilis Blackburn 1912, H. xanthotrichus Blackburn 1910] B00

Heteronyx subfuscus Macleay 1888 [incl. H. sordidus Blackburn 1910, H. subcylindricus Blackburn 1910, H. subvittatus Macleay 1888] B00

Heteronyx terrena Blackburn 1892 [=H. terrenus (l. c.); incl. H. elongatulus Dalla Torre 1912, H. elongatus Blackburn 1909 non Blanchard 1850, H. eremita Blackburn 1909, H. hothamensis Blackburn 1909, H. ingratus Blackburn 1909] B00

*Type species of generic name indicated

REFERENCES

[B00] Britton, E. B. 2000. A review of Heteronyx Guérin-Méneville (Coleoptera: Scarabaeidae: Melolonthinae). Invertebrate Taxonomy 14: 465–589.

[HW92] Houston, W. W. K., & T. A. Weir. 1992. Melolonthinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 174–358. AGPS Press: Canberra.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part III. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 21–126.

Scitala

Shiny pasture scarab Scitala sericans, from Insects of Tasmania.

Belongs within: Melolonthinae.

Scitala is an Australian genus of black or brown chafers lacking dorsal setae and with striae on the elytra paired so alternate intervals differ in width.

<==Scitala Erichson 1842 HW92
    |--*S. sericans Erichson 1842 HW92 (see below for synonymy)
    |--S. aureorufa (Blanchard 1850) (see below for synonymy) HW92
    |--S. calescens Blackburn 1907 HW92
    |--S. celata Britton 1987 HW92
    |--S. citra Britton 1987 HW92
    |--S. coracina Britton 1987 HW92
    |--S. eremia Britton 1987 HW92
    |--S. fustula Britton 1987 HW92
    |--S. glabra (Blanchard 1850) [=Sericesthis glabra] HW92
    |--S. hirticeps (Blackburn 1907) [=Anodontonyx hirticeps; incl. A. opalescens Lea 1919] HW92
    |--S. howdeni Britton 1987 HW92
    |--S. iridipennis Britton 1987 HW92
    |--S. juvenis Blackburn 1907 HW92
    |--S. minuta Britton 1987 HW92
    |--S. nana Britton 1987 HW92
    |--S. nemesis Blackburn 1907 HW92
    |--S. nigrolineata Boisd. 1835 M86
    |--S. pruinosa Dalm. 1823 [incl. S. australis Boisd. 1835, S. geminata Boisd. 1835] M86
    |--S. pullata Boisd. 1835 M86
    |--S. rorida Burmeister 1855 [incl. S. hospes Blackburn 1907] HW92
    |--S. satelles Britton 1987 HW92
    |--S. simulans Britton 1987 HW92
    |--S. sororia Britton 1987 HW92
    |--S. uptoni Britton 1987 HW92
    `--S. volux Britton 1987 HW92

Scitala aureorufa (Blanchard 1850) [=Sericesthis aureorufa; incl. Sc. ambigua Blackburn 1907, Sc. coxalis Blackburn 1907, Sc. subsericans Blackburn 1907] HW92

*Scitala sericans Erichson 1842 HW92 [incl. Sc. aenescens Burmeister 1855 HW92, Sericesthis rugosiceps Blanchard 1850 HW92, Sc. rugosiceps M86]

*Type species of generic name indicated

REFERENCES

[HW92] Houston, W. W. K., & T. A. Weir. 1992. Melolonthinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 174–358. AGPS Press: Canberra.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part III. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 21–126.

Sericesthis

Pruinose scarab Sericesthis geminata, from here.

Belongs within: Melolonthinae.

Sericesthis is an Australian genus of chafers characterised by elytra with striae grouped in pairs so odd-numbered intervals are narrower than adjacent even-numbered intervals.

<==Sericesthis Boisduval 1835 [incl. Anodontonyx Sharp 1890] HW92
    |--*S. geminata Boisduval 1835 HW92
    |--S. accola Britton 1987 HW92
    |--S. anepsia Britton 1987 HW92
    |--S. antennalis (Blackburn 1907) [=Anodontonyx antennalis] HW92
    |--S. armaticeps (Macleay 1871) [=Scitala armaticeps; incl. Sc. impressa Brenske 1900] HW92
    |--S. bisetosa Britton 1987 HW92
    |--S. brooksi Britton 1987 HW92
    |--S. calignea Britton 1987 HW92
    |--S. carnei Britton 1987 HW92
    |--S. cinnamea Britton 1987 HW92
    |--S. consanguinea (Blackburn 1907) [=Anodontonyx consanguineus; incl. A. noxius Arrow 1992] HW92
    |--S. coolumensis Britton 1987 HW92
    |--S. dispar Blackburn 1890 HW92
    |--S. egens Britton 1987 HW92
    |--S. elderi Britton 1987 HW92
    |--S. erosa Blackburn 1890 [incl. S. puncticollis Blackburn 1890] HW92
    |--S. fovea Britton 1987 HW92
    |--S. harti (Sharp 1890) [=Anodontonyx harti; incl. S. planiceps Blackburn 1890] HW92
    |--S. ignota Britton 1987 HW92
    |--S. illawarrae Britton 1987 HW92
    |--S. incisa Britton 1987 HW92
    |--S. incola Britton 1987 HW92
    |--S. ino (Blackburn 1907) [=Scitala ino] HW92
    |--S. insularis (Lea 1919) [=Anodontonyx insularis] HW92
    |--S. iris Britton 1987 HW92
    |--S. janetae Britton 1987 HW92
    |--S. latens Britton 1987 HW92
    |--S. lutea Britton 1987 HW92
    |--S. magna Britton 1987 HW92
    |--S. major Britton 1987 HW92
    |--S. metincisa Britton 1987 HW92
    |--S. micans Blackburn 1890 HW92 (see below for synonymy)
    |--S. minima Britton 1987 HW92
    |--S. miskoi Britton 1987 HW92
    |--S. nemoralis (Blackburn 1907) [=Scitala nemoralis; incl. Sc. convexicollis Blackburn 1907] HW92
    |--S. nigra (Lea 1919) [=Anodontonyx niger] HW92
    |--S. nigrolineata Boisduval 1835 HW92 (see below for synonymy)
    |--S. palumae Britton 1987 HW92
    |--S. parallela Blackburn 1890 HW92
    |--S. parvipes Blackburn 1890 HW92
    |--S. propria Britton 1987 HW92
    |--S. proxima Britton 1987 HW92
    |--S. pullata B35
    |--S. rectangula (Blackburn 1907) [=Andontonyx rectangulus] HW92
    |--S. rufescens Britton 1987 HW92
    |--S. serena Britton 1987 HW92
    |--S. stipata Britton 1987 HW92
    |--S. storeyi Britton 1987 HW92
    |--S. suberosa Britton 1987 HW92
    |--S. suturalis (Macleay 1871) [=Scitala suturalis; incl. Sc. pruinosella Brenske 1900] HW92
    |--S. tetrica (Blackburn 1907) [=Anodontonyx tetricus] HW92
    |--S. vera Britton 1987 HW92
    `--S. vigilans (Sharp 1890) [=*Anodontonyx vigilans; incl. A. chalceus Blackburn 1907] HW92

Sericesthis micans Blackburn 1890 HW92 [=Anodontonyx micans L19, Omaloplia micans B35; incl. A. creber Blackburn 1907 HW92, A. gravicollis Blackburn 1907 HW92, A. indignus Blackburn 1907 HW92]

Sericesthis nigrolineata Boisduval 1835 HW92 [=Anodontonyx nigrolineata L19; incl. Scitala languida Erichson 1842 HW92, Melolontha pruinosa Dalman 1823 non Wiedeman 1819 HW92, Sericesthis pruinosa Blanchard 1850 HW92, Scitala rugosula Harold 1869 HW92]

*Type species of generic name indicated

REFERENCES

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d'Urville. Faune entomologique de l'océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[HW92] Houston, W. W. K., & T. A. Weir. 1992. Melolonthinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 174–358. AGPS Press: Canberra.

[L19] Lea, A. M. 1919. Notes on some miscellaneous Coleoptera with descriptions of new species.—Part V. Transactions of the Royal Society of South Australia 43: 166–261, pls 25–27.

Rhopaea

Brown cockchafer Rhopaea magnicornis, from Brisbane Insects.

Belongs within: Melolonthini.

Rhopaea is an Australasian genus of chafers characterised by a dense dorsal covering of semi-erect setae and strongly toothed tarsal claws.

Characters (from Britton 1978): Anterior face of clypeus as deep as labrum, bearing scattered setiferous punctures all over its surface, not limited to a single transverse row; anterior edge of clypeus as seen from above straight or convex in the middle, never broadly bilobed; antenna 10-segmented with a 5- to 8-lamellate club, the lamellae much longer in the male than in the female; head, pronotum, scutellum and elytra clothed with short, fine, semierect setae, sometimes with sparse, long, erect setae in addition; never bearing scales; pronotum without defined or raised anterior and posterior margins; tarsal claws each with a strong tooth in the middle beneath; apices of parameres flanged.

Rhopaea Erichson 1847 HW92
    |--*R. verreauxii Blanchard 1851 HW92
    |--R. assimilis Blackburn 1911 HW92
    |--R. australis (Blackburn 1888) [=Holophylla australis] HW92
    |--R. decipiens Lea 1919 HW92
    |--R. heterodactyla (Germar 1848) [=Melolontha heterodactyla; incl. R. soror Blackburn 1892] HW92
    |--R. hirtuosa Blackburn 1898 HW92
    |--R. laticollis Blackburn 1911 HW92
    |--R. magnicornis Blackburn 1888 HW92
    |--R. nigricollis Lea 1919 HW92
    `--R. pilosa Blackburn 1911 HW92

*Type species of generic name indicated

REFERENCES

Britton, E. B. 1978. A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae) vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series 60: 1–150.

[HW92] Houston, W. W. K., & T. A. Weir. 1992. Melolonthinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 174–358. AGPS Press: Canberra.