Forest dominated by myrtle beech Nothofagus cunninghamii, photographed by Simon Fraser.

Belongs within: Faganae.
Contains: Juglandoideae.

The Juglandanae was listed by Doweld (2001) as a taxon uniting the wind-pollinated families Myricaceae, Juglandaceae and Nothofagus; the molecular analysis of Rosidae by Wang et al. (2009) supports a clade of Myrica and Juglans within the Fagales. The Juglandaceae are a group of trees (rarely shrubs) found in Eurasia and the Americas, characterised by production of fruit as nuts surrounded by a thick husk formed from the involucre. Members of the Juglandaceae are grown for their edible nuts, particularly species of Juglans (walnuts) and Carya (hickory nuts). The Myricaceae are a family of small trees and shrubs, most of which are aromatic and resinous. Nothofagus, the southern beeches, is a genus of trees found in South America and Australasia, primarily in cooler habitats.

    |  i. s.: Normapolles CBH93
    |         Manningia [incl. Trudopollis] CBH93
    |         Caryanthus CBH93
    |         Plicapollis usitatus Tschudy 1975 CBH93, CP02
    |--Juglandaceae [Juglandales] WM09
    |    |  i. s.: Juglandiphyllum integrifolium CBH93
    |    |         Momipites Wodehouse 1933 CBH93, CP02
    |    |           |--M. actinus Nichols & Orr 1978 CP02
    |    |           |--M. fragilis CBH93
    |    |           |--M. leffingwellii Nichols 1973 CP02
    |    |           |--M. rotundus (Leffingwell 1971) Nichols 1973 CP02
    |    |           |--M. tenuipolus Anderson 1960 CP02
    |    |           |--M. triorbicularis (Leffingwell 1971) Nichols 1973 CP02
    |    |           `--M. ventoifluminis Nichols 1973 CP02
    |    |--Platycarya [Platycaryoideae] T00
    |    |--Juglandoideae T00
    |    `--Engelhardioideae T00
    |         |--Alfaroa T00
    |         |--Oreomunnea T00
    |         `--Engelhardia D07
    |              |--E. orsbergensis HL08
    |              `--E. spicata D07
    |--Myricaceae [Myricales, Myricineae] D01
    |    |--Canacomyrica T00
    |    |--Comptonia peregrina H93, C38
    |    |--‘Myricophyllum’ longepetiolatum von Ettingshausen 1893 CBH93
    |    |--Triatriopollenites CBH93
    |    `--Myrica WM09
    |         |--M. californica HL08
    |         |--M. cerifera WM09
    |         |--M. esculenta P88
    |         |--M. faya Z02
    |         |--M. gale RJ11
    |         |--M. hartwegii H93
    |         |--M. longifolia Unger 1850 [incl. M. ophir Unger 1850] HL08
    |         |--M. matheroniana [=Comptonia matheroniana] S89
    |         |--M. mexicana BT87
    |         |--M. picardae J87
    |         |--M. rubra MH98
    |         |--M. salicifolia PB27
    |         `--M. vernassiensis GT02
    `--Nothofagus Blume 1850 A61 [Nothofagaceae, Nothofagales D01]
         |  i. s.: N. × blairii (Kirk) Cockayne 1911 [=Fagus blairii Kirk 1885; N. cliffortioides × N. fusca] A61
         |--N. subg. Lophozonia LC95
         |    |--+--N. cunninghamii LC95
         |    |  `--+--N. menziesii (Hooker) Oerst. 1873 LC95, A61 [=Fagus menziesii Hooker 1844 A61]
         |    |     `--N. moorei LC95
         |    `--+--N. glauca LC95
         |       `--+--N. alpina LC95
         |          `--N. obliqua LC95
         `--+--N. subg. Fuscospora LC95
            |    |  i. s.: N. × apiculata (Colenso) Cockayne 1911 (see below for synonymy) A61
            |    |           |--N. a. var. apiculata A61
            |    |           `--N. a. var. dubia (Kirk) Cheeseman 1925 (see below for synonymy) A61
            |    |--N. alessandri LC95
            |    `--+--N. gunnii LC95
            |       `--+--N. solandri (Hooker) Oerst. 1873 LC95, A61 [=Fagus solandri Hooker 1844 A61, N. solanderi A61]
            |          |    |--N. s. var. solandri A61
            |          |    `--N. s. var. cliffortioides (Hooker) Poole 1958 (see below for synonymy) A61
            |          `--+--N. fusca (Hooker) Oerst. 1873 LC95, A61 (see below for synonymy)
            |             `--N. truncata (Col.) Cockayne 1926 LC95, A61 (see below for synonymy)
            `--+--N. subg. Nothofagus LC95
               |    |--+--N. antarctica LC95
               |    |  `--N. pumilio LC95
               |    `--+--N. nitida LC95
               |       `--+--N. betuloides LC95
               |          `--N. dombeyi LC95
               `--N. subg. Brassospora LC95
                    |--N. rubra LC95
                    |--+--N. discoidea LC95
                    |  `--+--N. aequilateralis LC95
                    |     `--+--N. balansae LC95
                    |        `--N. codonandra LC95
                    `--+--+--N. carrii LC95
                       |  `--+--N. flaviramea LC95
                       |     `--N. pullei LC95
                       `--+--+--N. crenata LC95
                          |  `--+--N. brassii LC95
                          |     `--N. starkenborghii LC95
                          `--+--N. perryi LC95
                             `--+--N. grandis LC95
                                `--N. resinosa LC95

Nothofagus × apiculata (Colenso) Cockayne 1911 [=Fagus apiculata Colenso 1884; N. solandri × N. truncata] A61

Nothofagus apiculata var. dubia (Kirk) Cheeseman 1925 [=Fagus fusca var. dubia Kirk 1889; incl. F. fusca var. obsoleta Kirk 1889] A61

Nothofagus fusca (Hooker) Oerst. 1873 LC95, A61 [=Fagus fusca Hooker 1844 A61; incl. F. punctata ms A61]

Nothofagus solandri var. cliffortioides (Hooker) Poole 1958 [=Fagus cliffortioides Hooker 1844, N. cliffortioides (Hooker) Oerst. 1873] A61

Nothofagus truncata (Col.) Cockayne 1926 LC95, A61 [=Fagus truncata Col. 1899 A61; incl. F. fusca var. colensoi Hooker 1854 A61, Nothofagus fusca var. colensoi (Hooker) Cheesem. 1925 A61, F. rotundifolia ms A61]

*Type species of generic name indicated


[A61] Allan, H. H. 1961. Flora of New Zealand vol. 1. Indigenous Tracheophyta: Psilopsida, Lycopsida, Filicopsida, Gymnospermae, Dicotyledones. R. E. Owen, Government Printer: Wellington (New Zealand).

[BT87] Baker, E. W., & D. M. Tuttle. 1987. The false spider mites of Mexico (Tenuipalpidae: Acari). United States Department of Agriculture, Technical Bulletin 1706: 1–237.

[CP02] Christopher, R. A., & D. C. Prowell. 2002. A palynological biozonation for the Maastrichtian stage (Upper Cretaceous) of South Carolina, USA. Cretaceous Research 23: 639–669.

[CBH93] Collinson, M. E., M. C. Boulter & P. L. Holmes. 1993. Magnoliophyta (‘Angiospermae’). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 809–841. Chapman & Hall: London.

[C38] Crawford, J. C. 1938. Some new or little known Thysanoptera. Proceedings of the Entomological Society of Washington 40 (2): 35–43.

[D07] Dash, S. S. 2007. Useful plants of Kabi Sacred Grove, Sikkim. Bulletin of the Botanical Survey of India 49: 79–88.

[D01] Doweld, A. B. 2001. The systematic relevance of fruit and seed structure in Bersama and Melianthus (Melianthaceae). Plant Systematics and Evolution 227: 75–103.

[GT02] Gomez, B., F. Thévenard, M. Fantin & L. Guisberti. 2002. Late Cretaceous plants from the Bonarelli Level of the Venetian Alps, northeastern Italy. Cretaceous Research 23: 671–685.

[HL08] Henniger, M., & R. Leder. 2008. Eozäne Makroflora der Querfurter Mulde. Mauritiana 20 (2): 229–251.

[H93] Hickman, J. C. (ed.) 1993. The Jepson Manual: Higher Plants of California. University of California Press: Berkeley (California).

[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum—Biological Sciences 32 (1): 1–136.

[LC95] Linder, H. P., & M. D. Crisp. 1995. Nothofagus and Pacific biogeography. Cladistics 11: 5–32.

[MH98] Morikawa, H., A. Higaki, M. Nohno, M. Takahashi, M. Kamada, M. Nakata, G. Toyohara, Y. Okamura, K. Matsui, S. Kitani, K. Fujita, K. Irifune & N. Goshima. 1998. More than a 600-fold variation in nitrogen dioxide assimilation among 217 plant taxa. Plant, Cell and Environment 21: 180–190.

[PB27] Pilsbry, H. A., & J. Bequaert. 1927. The aquatic mollusks of the Belgian Congo, with a geographical and ecological account of Congo malacology. Bulletin of the American Museum of Natural History 53 (2): 69–602, pls 10–77.

[P88] Polunin, I. 1988. Plants and Flowers of Malaysia. Times Editions: Singapore.

[RJ11] Rising, J. D., A. Jaramillo, J. L. Copete, P. G. Ryan & S. C. Madge. 2011. Family Emberizidae (buntings and New World sparrows). In: Hoyo, J. del, A. Elliott & D. A. Christie (eds) Handbook of the Birds of the World vol. 16. Tanagers to New World Blackbirds pp. 428–683. Lynx Edicions: Barcelona.

[S89] Squinabol, S. 1889. Res Ligusticae. VII.—Cenno preliminare sulla flora fossile di Santa Giustina. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 73–76.

[T00] Thorne, R. F. 2000. The classification and geography of the flowering plants: dicotyledons of the class Angiospermae (subclasses Magnoliidae, Ranunculidae, Caryophyllidae, Dilleniidae, Rosidae, Asteridae, and Lamiidae). The Botanical Review 66: 441–647.

[WM09] Wang, H., M. J. Moore, P. S. Soltis, C. D. Bell, S. F. Brockington, R. Alexandre, C. C. Davis, M. Latvis, S. R. Manchester & D. E. Soltis. 2009. Rosid radiation and the rapid rise of angiosperm-dominated forests. Proceedings of the National Academy of Sciences of the USA 106 (10): 3853–3858.

[Z02] Zherikhin, V. V. 2002. Ecological history of the terrestrial insects. In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 331–388. Kluwer Academic Publishers: Dordrecht.

Last updated: 5 September 2020.

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