Demarziella

Demarziella sp., copyright Rebecca Harris.

Belongs within: Scarabaeinae.

Demarziella is an Australian genus of small dung beetles with oblique mid coxae and unarmed head and pronotum in both sexes.

Characters (from Matthews 1976, as Notopedaria): Oblong, feebly convex, fuscous, often bronzed and occasionally metallic, antenna1 clubs flavous, dorsally with stout and decumbent bristles combined with some very small setae. Total length 2.9-5.0 mm. Head subcircular, genal angles absent or very obtuse. Clypeal margin bidentate, the teeth widely separated by an arcuate emargination. Frons unarmed. Dorsal portion of eyes moderately to very narrow, the ventral portion large and globose, end of canthus remote from occipital edge, which is curved out to meet it. Underside of clypeal edge with a strong angular ridge, without a pit or groove. Labium trapezoidal, with a very shallow anterior emargination not fringed by setae. Labial palpi with first and second segments subtriangular, very wide, subequal in size and much larger than third segment, which is minute. Demarcation between gula and submentum more or less transverse. Antennae of nine segments. Hind angles of head prominent. Pronotum subquadrate, not strongly convex, unarmed but with various shallow depressions and elevations, and sparse, stout bristles often arranged in a transverse row. Deflexed areas of pronotum deeply excavated anteriorly to receive fore femora, the excavation bounded posteriorly by a complete carina running to lateral edges, a second carina paralleling lateral edges posteriorly. Lateral edges not well defined, with a regular row of erect bristles. Scutellum hidden. Mesosternum about four times as wide as long at widest point, mesometasternal suture feebly bent or curved forward. Median lobe of metasternum narrow, about 2.5 times as long as wide at narrowest point, middle coxal cavities oblique and very broadly margined anteriorly, this margin oftcn incomplete. Underside generally very densely and coarsely punctate, with short bristles. Elytra with ten striae, including the one along epipleural edge, eighth represented only by a short anterior section and ninth present only for posterior half, very close to tenth. Tenth greatly enlarged and deepened anteriorly. Striae usually unevenly spaced, the odd-numbered intervals wider and often higher than even-numbered ones. Striae variable, often widened or geminate. Intervals variable, but fifth always subcarinate, the elytral surface deflexed along it. Some stout bristles on elytra. Epipleura of moderate width, their upper edges carinate. Hind wings always fully developed. Legs with fore tarsi present, short, held in transverse grooves in tibial truncation. Fore spur small to minute. Outer edge of fore tibia with three teeth, the distal edge transversely truncate. Middle and hind tibiae strongly expanded distally, without any transverse carinae. The distal edges more or less straight and transverse, with a fringe of very short, stout bristles as well as some longer setae. Two short unequal spurs on middle tibiae, one short spur on hind. Middle and hind tarsi short, the segments generally subparallel, the first two subequal, the rest gradually diminishing in size, claws small, simple. All leg segments with short bristles. Abdomen with all sternites fused, the last considerably longer along midline in both sexes; with large shallow punctures but not setae. Pygidium fully margined, with a groove along dorsal edge, setose. Parameres of aedeagus asymmetrical.

<==Demarziella Balthasar 1961 [incl. Notopedaria Matthews 1976] CW92
    |--*D. mirifica Balthasar 1961 [incl. Notopedaria occidentalis Matthews 1976] CW92
    |--D. alternata (Lea 1923) [=Pedaria alternata] CW92
    |--D. eungella (Matthews 1976) [=Notopedaria eungella] CW92
    |--D. geminata (Macleay 1871) CW92 (see below for synonymy)
    |--D. imitatrix Balthasar 1967 [incl. D. costata Petrovitz 1975, Notopedaria tuberculata Matthews 1976] CW92
    |--D. interrupta (Carter 1936) (see below for synonymy) CW92
    |--D. metallica (Carter 1936) [=Pedaria metallica] CW92
    |--D. planitarsis Petrovitz 1971 CW92
    |--D. pratensis (Matthews 1976) [=Notopedaria pratensis] CW92
    |--D. scarpensis (Matthews 1976) [=Notopedaria scarpensis] CW92
    |--D. storeyi (Matthews 1976) [=Notopedaria storeyi] CW92
    |--D. sylvestris (Matthews 1976) [=Notopedaria sylvestris] CW92
    |--D. tropicalis (Matthews 1976) [=Notopedaria tropicalis] CW92
    `--D. yorkensis Matthews & Stebnicka 1986 CW92

Demarziella geminata (Macleay 1871) CW92 [=Aphodius geminatus CW92, Pedaria geminatus M86; incl. D. proxima Petrovitz 1963 CW92, D. umbilicata Petrovitz 1975 CW92]

Demarziella interrupta (Carter 1936) [=Pedaria interrupta, *Notopedaria interrupta; incl. Demarziella setosa Balthasar 1967] CW92

*Type species of generic name indicated

REFERENCES

[CW92] Cassis, G., & T. A. Weir. 1992. Scarabaeinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 106–173. AGPS Press: Canberra.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part III. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 21–126.

Matthews, E. G. 1976. A revision of the scarabaeine dung beetles of Australia. III. Tribe Coprini. Australian Journal of Zoology, Supplementary Series 38: 1–52.