Bucculatrix cristatella, copyright Ilia Ustyantsev.

Belongs within: Heteroneura.
Contains: Tineidae, Apoditrysia, Yponomeutoidea, Gracillariidae, Lyonetiidae, Psychinae.

The Ditrysia are a major clade of Lepidoptera accounting for some 98% of the species of the order. They are characterised by female genitalia with an internal connection between the copulatory opening and ovipore, as well as by large apodemes on the second abdominal sternite and short bands rather than long fibres in the proboscis muscles (Grimaldi & Engel 2005). Basal members of the clade, historically gathered into the superfamily Tineoidea (now recognised as non-monophyletic) are small to medium-sized species with narrow, typically grey fore wings that often retain the M vein in the discal cell (Nielsen & Common 1991). The Simaethistiidae are a small, poorly known group of basal ditrysians found in eastern Asia and Australia

<==Ditrysia [Gracillarioidea, Papilionina, Tineina, Tineites, Tineoidea, Tortrices]
    |--Meessiinae [Meessiidae] KP19
    |    |--Meessia leopoldella YS10
    |    |--Demobrotis NC91
    |    `--Eudarcia simulatricella KP19
    `--+--+--+--Tineidae KP19
       |  |  `--Dryadaula [Dryadaulidae] KP19
       |  |       `--D. visaliella KP19
       |  `--+--Apoditrysia GE05
       |     `--+--+--Yponomeutoidea KP19
       |        |  `--Roeslerstammiidae [Amphitheridae, Amphitherinae] NC91
       |        |       |--Amphithera heteroleuca KP19, NC91
       |        |       |--Enchoptila idiopis P27
       |        |       |--Thereutis NC91
       |        |       |--Macarangela leucochrysa NC91, C91
       |        |       |--Nematobola NC91
       |        |       |--Chalcoteuches phlogera NC91
       |        |       `--Roeslerstammia NC91
       |        `--+--Gracillariidae KP19
       |           `--+--Bucculatricidae NC91
       |              |    |--Ogmograptis scribula NC91
       |              |    `--Bucculatrix KP19
       |              |         |--B. cristatella KP19
       |              |         |--B. gossypii NC91
       |              |         `--B. platani RJ93
       |              `--+--Lyonetiidae KP19
       |                 |--Bedellia NC91 [Bedelliidae M03, Bedelliinae]
       |                 |    |--B. psamminella L27
       |                 |    `--B. somnulentella P27
       |                 `--Heliodinidae M03
       |                      |--Thylacosceles acridomima P27
       |                      |--Heliodines princeps NC91
       |                      |--Epicroesa ambrosia NC91
       |                      |--Baltonides roeselliformis RJ93
       |                      `--Vanicela P27
       |                           |--V. disjunctella P27
       |                           `--V. xenadelpha P27
       `--Psychidae B37
            |  i. s.: Liothula omnivora WFS04
            |         Solenobia triquetrella D51
            |         Iphierga macarista NC91
            |         Epichnopteryx plumella (Denis & Schiffermüller 1775) JP05
            |         Canephora JP05
            |           |--C. hirsuta Poda 1761 JP05
            |           `--C. unicolor RD77
            |         Paramictoides febretta YS10
            |         Pachythelia unicolor YS10
            |         Apterona YS10
            |         Plutorectis G84
            |         Paracharactis MC13
            |         Cryptothelea GE05
            |           |--C. cardiophaga IR05
            |           |--C. gloverii VC07
            |           `--C. junodi F92
            |         Luffia YS10
            |           |--L. ferchaultella BC91
            |           `--L. lapidella YS10
            |         Animula huebneri H01
            |         Cochliotheca RD77
            |           |--C. crenulella RD77
            |           `--C. helix RD77
            |         Acanthopsyche opacella [incl. A. atra] RD77
            |         Amicta RD77
            |         Sterrhopteryx pristinella Rebel 1934 P92
            |--Psychinae NC91
            |--Oiketicinae KP19
            |    |--Thyridopteryx ephemeraeformis KP19
            |    `--Oiketicus G84
            |         |--O. elongatus G84
            |         `--O. kirbyi L81
            `--Taleporiinae NC91
                 |--Ctenocompa C70
                 |--Narycia saxosa C70, P27
                 |--Trigonocyttara clandestina C70
                 |--Lepidoscia arctiella C70
                 |--Cebysa leucotelus C70, NC91
                 `--Taleporia JP05
                      |--T. cawthronella P27
                      `--T. tubulosa (Retzius 1783) JP05

Ditrysia incertae sedis:
  Azinis hilarella WM66
  Apoprogonidae NC91
  Cerostoma Latreille 1802 L02
    |--*C. dorsatum [=Ypsolophus dorsatus] L02
    `--C. persicella F92
  Simaethistiidae GE05
  Pseudarbelidae NC91
  Arrhenophanidae NC91
  Acrolophus D01 [Acrolophidae GE05]
    |--A. boucardi Druce 1901 D01
    |--A. linus Druce 1901 D01
    `--A. underwoodi Druce 1901 D01
  Eucryptogona [Eriocottidae] NC91
    `--E. trichobathra NC91
  Douglasiidae NC91
    |--Douglasia Stainton 1854 [=Douglassia (l. c.) non Bartsch 1934] P66
    |--Tinagma leucanthes NC91
    `--Klimeschia NC91
  Melusina [Compsoctenidae] RD77
    `--M. energa RD77
  Euplocamus guttella G20

*Type species of generic name indicated


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Last updated: 3 March 2021.

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