Female Methocha japonica, from here.

Belongs within: Aculeata.
Contains: Apoidea, Formicidae, Scoliidae, Pompiloidea, Tiphiidae, Thynninae, Vespidae.

The Euaculeata are a major clade of stinging wasps, united by sexually dimorphic antennae with thirteen antennomeres in males and twelve in males, and a hidden and largely desclerotised seventh metasomal tergum in females (Grimaldi & Engel 2005).

<==Euaculeata [Formicina, Heterogyna, Melliniores, Mutillariae, Praedones, Scolietae, Sphegimae] GE05
    |  i. s.: Falsiformicidae GE05
    |           |--Falsiformica cretacea Rasnitsyn 1975 P92
    |           `--Taimyrisphex pristinus Evans 1973 P92
    |--+--+--Apoidea BD17
    |  |  `--+--Formicidae BD17
    |  |     `--Armania [Armaniidae] GE05
    |  |          `--A. robusta GE05
    |  `--Scolioidea BD17
    |       |--Scoliidae JB13
    |       `--Bradynobaenidae JB13
    |            |--Typhoctes [Typhoctinae] GE05
    |            `--Apterogyna [Apterogyninae] JB13
    |                 `--A. olivieri [=Mutilla (Apterogyna) olivieri] G20
    `--+--+--Sierolomorpha GE05 [Sierolomorphidae, Sierolomorphoidea BD17]
       |  |    |--S. bicolor B11
       |  |    `--S. canadensis B11
       |  `--+--Pompiloidea PK17
       |     `--+--Tiphiidae JB13
       |        `--Thynnoidea BD17
       |             |--Chyphotes JB13 [Chyphotidae BD17, Chyphotinae]
       |             |    `--C. mellipes RBV08
       |             `--Thynnidae BD17
       |                  |--Myzinum [Myzininae] BD17
       |                  |    |--M. frontale S94
       |                  |    `--M. quinquecinctum (Fabricius 1775) [=Tiphia quinquecincta, Sphex quinquecinctus] BM76
       |                  `--+--Thynninae BD17
       |                     `--Methocha Latreille 1804 BD17, LB08 [Methochinae N91, Methocidae]
       |                          |--M. articulata PK17
       |                          |--M. formicaria (Linnaeus 1758) G20, L58 [=Mutilla formicaria G20]
       |                          |--M. ichneumonides A71
       |                          |--M. japonica S94
       |                          `--M. yasumatsui S94
       `--Vespoidea BD17
             |--Vespidae JB13
             `--+--Rhopalosomatidae JB13
                |    |--Rhopalosoma nearcticum HH07
                |    |--Mesorhopalosoma cearae Darling in Darling & Sharkey 1990 RJ93
                |    |--Propalosoma GE05
                |    |--Olixon [incl. Psyllosphex Arnold 1935] S94
                |    `--Harpagocryptus australiae R70
                `--Scolebythidae HR11
                     |  i. s.: Libanobythus Prentice & Poinar in Prentice, Poinar & Milki 1996 PPM96
                     |           `--*L. milkii Prentice & Poinar in Prentice, Poinar & Milki 1996 PPM96
                     |         Dominibythus Prentice & Poinar in Prentice, Poinar & Milki 1996 PPM96
                     |           `--*D. inopinatus Prentice & Poinar in Prentice, Poinar & Milki 1996 PPM96
                     |         Boreobythus Engel & Grimaldi 2007 B11
                     |         Cretabythus sibiricus Evans 1973 P92
                     |--+--Clystopsenella Kieffer 1911 B11
                     |  |    `--C. longiventris PPM96
                     |  `--Scolebythus Evans 1963 B11
                     |       `--S. madecassus HR11
                     `--+--Ycaploca Nagy 1975 B11
                        |    `--Y. evansi GE05
                        `--Pristapenesia Brues 1933 B11
                             |--P. inopinata B11
                             `--P. primaeva B11

*Type species of generic name indicated


[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[BD17] Branstetter, M. G., B. N. Danforth, J. P. Pitts, B. C. Faircloth, P. S. Ward, M. L. Buffington, M. W. Gates, R. R. Kula & S. G. Brady. 2017. Phylogenomic insights into the evolution of stinging wasps and the origins of ants and bees. Current Biology 27: 1019–1025.

[B11] Brothers, D. J. 2011. A new Late Cretaceous family of Hymenoptera, and phylogeny of the Plumariidae and Chrysidoidea (Aculeata). ZooKeys 130: 515–542.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: N├╝rnberg.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[HR11] Heraty, J., F. Ronquist, J. M. Carpenter, D. Hawks, S. Schulmeister, A. P. Dowling, D. Murray, J. Munro, W. C. Wheeler, N. Schiff & M. Sharkey. 2011. Evolution of the hymenopteran megaradiation. Molecular Phylogenetics and Evolution 60: 73–88.

[HH07] Hines, H. M., J. H. Hunt, T. K. O'Connor, J. J. Gillespie & S. A. Cameron. 2007. Multigene phylogeny reveals eusociality evolved twice in vespid wasps. Proceedings of the National Academy of Sciences of the USA 104 (9): 3295–3299.

[JB13] Johnson, B. R., M. L. Borowiec, J. C. Chiu, E. K. Lee, J. Atallah & P. S. Ward. 2013. Phylogenomics resolves evolutionary relationships among ants, bees, and wasps. Current Biology 23: 2058–2062.

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[N91] Naumann, I. D. 1991. Hymenoptera (wasps, bees, ants, sawflies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 916–1000. Melbourne University Press: Carlton (Victoria).

[PK17] Peters, R. S., L. Krogmann, C. Mayer, A. Donath, S. Gunkel, K. Meusemann, A. Kozlov, L. Podsiadlowski, M. Petersen, R. Lanfear, P. A. Diez, J. Heraty, K. M. Kjer, S. Klopfstein, R. Meier, C. Polidori, T. Schmitt, S. Liu, X. Zhou, T. Wappler, J. Rust, B. Misof & O. Niehuis. 2017. Evolutionary history of the Hymenoptera. Current Biology 27 (7): 1013–1018.

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

[PPM96] Prentice, M. A., G. O. Poinar, Jr & R. Milki. 1996. Fossil scolebythids (Hymenoptera: Scolebythidae) from Lebanese and Dominican amber. Proceedings of the Entomological Society of Washington 98 (4): 801–811.

[RBV08] Rabeling, C., J. M. Brown & M. Verhaagh. 2008. Newly discovered sister lineage sheds light on early ant evolution. Proceedings of the National Academy of Sciences of the USA 105 (39): 14913–14917.

[R70] Riek, E. F. 1970. Hymenoptera (wasps, bees, ants). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 867–959. Melbourne University Press.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.

[S94] Shimizu, A. 1994. Phylogeny and classification of the family Pompilidae (Hymenoptera). Tokyo Metropolitan University Bulletin of Natural History 2: 1–142.

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