Nagurus cristatus, copyright Arp Kruihof.

Belongs within: Oniscidea.
Contains: Oniscidae, Philosciidae, Armadillidae, Porcellio.

The Crinocheta are a group of terrestrial isopods characterised by an 'embolophore' type of male genitalia with the two vasa deferentia opening by separate pores on either side of the penis towards its apex (Hatch 1947, as Embolophora). They are divided between the Armadilloidea, in which the pleopods carry respiratory pseudotracheae, and the Oniscoidea in which such structures are absent.

<==Crinocheta [Embolophora] MD01
    |--Oniscoidea MD01
    |    |  i. s.: Tasmanoniscus Vandel 1973 V73
    |    |           `--*T. evansi Vandel 1973 V73
    |    |--Berytoniscidae MD01
    |    |--Olibrinus Budde-Lund 1913 JP13 [Olibrinidae MD01]
    |    |--Pudeoniscidae MD01
    |    |--Speleoniscidae MD01
    |    |--Sphaeroniscidae MD01
    |    |--Stenoniscidae MD01
    |    |--Tendosphaeridae MD01
    |    |--Oniscidae MD01
    |    |--Philosciidae MD01
    |    |--Halophiloscia PP64 [Halophilosciidae MD01]
    |    |    `--H. couchii PP64
    |    |--Australoniscus Vandel 1973 V73 [Bathytropidae MD01]
    |    |    `--*A. springetti Vandel 1973 V73
    |    |--Deto V73 [Detonidae MD01]
    |    |    `--D. marina (Chilton 1884) V73
    |    |--Rhyscotus M90 [Rhyscotidae MD01]
    |    |    `--R. texensis M90
    |    |--Platyarthrus M90 [Platyarthridae MD01]
    |    |    |--P. hoffmannseggi M90
    |    |    `--P. schoebli M90
    |    `--Scyphacidae MD01
    |         |--Scyphacella arenicola M90
    |         |--Detonella [Scyphacinae] H47
    |         |    |--D. papillicornis [=Trichoniscus papillicornis; incl. D. lohmanderi] H47
    |         |    `--D. sachalina H47
    |         `--Armadilloniscus [Armadilloniscinae] H47
    |              |--A. ellipticus M90
    |              |--A. holmesi M90
    |              |--A. lindahli M90
    |              `--A. tuberculatus [=Actoniscus tuberculatus] H47
    `--Armadilloidea MD01
         |--Atlantididae MD01
         |--Balloniscidae MD01
         |--Eubelidae MD01
         |--Periscyphicidae MD01
         |--Armadillidae MD01
         |--Cylisticus M90 [Cylisticidae MD01]
         |    |--C. convexus M90 [incl. C. convexus var. confluentus H47, Porcellio glaber H47]
         |    `--C. rotabilis D01
         |--Armadillidium M90 [Armadillidiidae MD01]
         |    |--A. (Armadillidium) vulgare [incl. Armadillo pilularis Say 1818] H47
         |    `--A. (Pseudosphaerium) nasatum H47
         |--Actaecia Dana 1853 V73 [incl. Cylloma Budde-Lund 1885 V73; Actaeciidae MD01]
         |    |--A. euchroa Chilton 1901 NB26
         |    |--A. opihensis Chilton 1901 NB26
         |    `--A. pallida Nicholls & Barnes 1926 V73
         |--Porcellionidae MD01
         |    |--Porcellio S77
         |    |--Agabiformius lentus (Budde-Lund 1885) V73
         |    `--Porcellionides Miers 1877 JP13
         |         `--P. pruinosus (Brandt 1833) JP13 [=Metoponorthus pruinosus M90; incl. P. immaculatus H47]
         `--Trachelopodidae [Trachelipidae] MD01
              |--Trachelipus [incl. Tracheoniscus] M90
              |    `--T. rathkei M90 (see below for synonymy)
              `--Nagurus Holthuis 1949 [incl. Nagara Budde-Lund 1908 (preoc.), Nagaroides Wahrberg 1922] V73
                   |--N. aegaeus Schmalfuss 1977 S77
                   |--N. carinatus [incl. N. hermonensis Vandel 1955] S77
                   |--N. cristatus (Dollfus 1889) V73
                   |--N. declivus (Wahrberg 1922) V73
                   |--N. gracillimus (Wahrberg 1922) V73
                   |--N. longiflagellatus (Wahrberg 1922) V73
                   |--N. rhodiensis S77
                   `--N. tumidus (Wahrberg 1922) V73

Nomen nudum: Australoniscus alticolus Vandel 1973 V73

Trachelipus rathkei M90 [incl. Porcellio dorsalis H47, P. lateralis H47, P. limatus H47, P. limbalis H47, P. marginatus H47, P. mixtus H47, P. multiguttatus H47, T. rathkei var. ochraceus H47, Porcellio variegatus H47, T. rathkei var. varius H47]

*Type species of generic name indicated


[D01] Dollfus, A. 1901. Isopodák [Isopodes]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 371–374. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[H47] Hatch, M. H. 1947. The Chelifera and Isopoda of Washington and adjacent regions. University of Washington Publications in Biology 10 (5): 155–274.

[JP13] Judd, S., & G. Perina. 2013. An illustrated key to the morphospecies of terrestrial isopods (Crustacea: Oniscidea) of Barrow Island, Western Australia. Records of the Western Australian Museum Supplement 83: 185–207.

[MD01] Martin, J. W., & G. E. Davis. 2001. An updated classification of the Recent Crustacea. Natural History Museum Los Angeles County, Science Series 39: 1–124.

[M90] Muchmore, W. B. 1990. Terrestrial Isopoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 805–817. John Wiley & Sones: New York.

[NB26] Nicholls, G. E., & H. M. Barnes. 1926. A description of two new terrestrial isopods from Western Australia. Journal of the Royal Society of Western Australia 12 (17): 149–159.

[PP64] Peres, J. M., & J. Picard. 1964. Nouveau manuel de bionomie benthique de la mer Mediterranee. Recueil des Travaux de la Station Marine d'Endoume, Bulletin 31 (27): 5–137.

[S77] Schmalfuss, H. 1977. Eine neue Nagurus-Art aus der Süd-Agäis (Crustacea: Isopoda: Oniscoidea: Trachelipidae). Senckenbergiana Biologica 57 (4–6): 359–365.

[V73] Vandel, A. 1973. Les isopodes terrestres de l'Australie. Étude systématique et biogéographique. Mémoires du Muséum National d'Histoire Naturelle, Série A, Zoologie 82: 1–171.

Last updated: 29 October 2017.

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