Asellidae

One-spotted waterlouse Proasellus meridianus, copyright Keith Lugg.


Belongs within: Asellota.
Contains: Asellus (Asellus).

The Asellidae are a group of freshwater crustacean of the Holarctic region (Henry et al. 1986).

Characters (from Henry et al. 1986): Antenna bearing a squama; maxillipedal oostegite reduced to a "Wasserstrudelapparat" (water swirling apparatus); pleonites 1 and 2 reduced both in length and width, forming a stalk between pereion and pleotelson; male copulatory organ (endopodite of pleopods 2) secondarily uniarticulate, but complex.

<==Asellidae
    |--Baicalasellus HLM86
    |--Gallasellus Henry & Magniez 1977 HLM86
    |    `--G. heilyi (Legrand 1956) HLM86
    |--Stygasellus Chappuis 1943 HLM86
    |    `--S. phreaticus (Chappuis 1943) [=Stygonectes phreaticus] HLM86
    |--Uenasellus Matsumoto 1962 HLM86
    |    `--U. iyoensis (Matsumoto 1960) HLM86
    |--Salmasellus Bowman 1975 HLM86
    |    `--S. steganothrix Bowman 1975 HLM86
    |--Psammasellus Braga 1968 HLM86
    |    `--P. capitatus Braga 1968 HLM86
    |--Calasellus Bowman 1981 HLM86
    |    |--C. californicus (Miller 1933) HLM86
    |    `--C. longus Bowman 1981 HLM86
    |--Lirceolus Bowman & Longley 1976 HLM86
    |    |--L. pilus (Steeves 1968) HLM86
    |    `--L. smithii (Ulrich 1902) HLM86
    |--Lirceus Rafinesque 1820 HLM86
    |    |--L. culveri Holsinger & Bowman 1973 HLM86
    |    |--L. fontinalis HLM86
    |    `--L. usdagalum Estes & Holsinger 1976 HLM86
    |--Nipponasellus Matsumoto 1962 HLM86
    |    |--N. aioii (Chappuis 1955) HLM86
    |    |--N. hubrichti (Matsumoto 1956) HLM86
    |    |--N. kagaensis (Msatsumoto 1958) HLM86
    |    |--N. takefuensis (Matsumoto 1961) HLM86
    |    `--N. tonensis (Matsumoto 1961) HLM86
    |--Bragasellus Henry & Magniez 1968 HLM86
    |    |--B. aireyi Henry & Magniez 1980 HLM86
    |    |--B. boui Henry & Magniez 1974 HLM86
    |    |--B. comasi Henry & Magniez 1976 HLM86
    |    |--B. comasioides Magniez & Brehier 2004 F05
    |    |--B. conimbricensis (Braga 1946) HLM86
    |    |--B. escolai Henry & Magniez 1978 HLM86
    |    |--B. frontellum (Braga 1964) HLM86
    |    |--B. incurvatus Afonso 1984 HLM86
    |    |--B. lagari Henry & Magniez 1973 HLM86
    |    |--B. pauloae (Braga 1958) HLM86
    |    `--B. seabrai (Braga 1943) HLM86
    |--Asellus Geoffroy 1762 HLM86
    |    |  i. s.: A. cavaticus T30
    |    |         A. meridianus N29
    |    |         A. nipponensis Nicholls 1929 N29
    |    |--A. (Asellus) HLM86
    |    |--A. (Conasellus) communis H47
    |    |--A. (Mesoasellus Birstein 1939) HLM86
    |    |    |--A. (M.) dentifer Birstein & Levanidov 1952 HLM86
    |    |    `--A. (M.) strinatii Chappuis 1955 HLM86
    |    `--A. (Phreatosellus Matsumoto 1962) HLM86
    |         |--A. (P.) akiyoshiensis (Ueno 1927) HLM86
    |         |--A. (P.) higoensis Matsumoto 1960 HLM86
    |         |--A. (P.) iriei Matsumoto 1978 HLM86
    |         |--A. (P.) kawamurai (Tattersall 1921) HLM86
    |         |--A. (P.) minatoi Matsumoto 1978 HLM86
    |         |--A. (P.) miurai Chappuis 1955 HLM86
    |         |--A. (P.) uenoi Matsumoto 1978 HLM86
    |         `--A. (P.) yoshinoensis Matsumoto 1960 HLM86
    |--Synasellus Braga 1944 HLM86
    |    |--S. albicastrensis Braga 1960 HLM86
    |    |--S. barcelensis Noodt & Galhano 1969 HLM86
    |    |--S. brigantinus Braga 1959 HLM86
    |    |--S. exiguus Braga 1944 HLM86
    |    |--S. fragilis Braga 1946 HLM86
    |    |--S. insignis Afonso 1984 HLM86
    |    |--S. lafonensis Braga 1959 HLM86
    |    |--S. longicauda Braga 1959 HLM86
    |    |--S. mariae (Braga 1942) HLM86
    |    |    |--S. m. mariae HLM86
    |    |    `--S. m. favaiensis Eiras 1974 HLM86
    |    |--S. mateusi Braga 1954 HLM86
    |    |--S. meirelesi Braga 1959 HLM86
    |    |--S. minutus Braga 1967 HLM86
    |    |--S. nobrei Braga 1967 HLM86
    |    |--S. pireslimai Braga 1959 HLM86
    |    |--S. serranus Braga 1967 HLM86
    |    `--S. transmontanus Braga 1959 HLM86
    |--Caecidotea Packard 1871 HLM86
    |    |--C. acuticarpa Mackin & Hubricht 1940 HLM86
    |    |--C. adenta Mackin & Hubricht 1940 HLM86
    |    |--C. alabamensis Stafford 1911 HLM86
    |    |--C. ancyla (Fleming 1972) HLM86
    |    |--C. antricola (Creaser 1931) HLM86
    |    |--C. barri (Steeves 1965) HLM86
    |    |--C. beattyi Lewis & Bowman 1981 HLM86
    |    |--C. bicrenata (Steeves 1963) HLM86
    |    |    |--C. b. bicrenata HLM86
    |    |    `--C. b. whitei Lewis & Bowman 1981 HLM86
    |    |--C. bisetus (Steeves 1968) HLM86
    |    |--C. bowmani Lewis 1980 HLM86
    |    |--C. brevicauda HLM86
    |    |--C. cannula (Steeves 1963) HLM86
    |    |--C. carolinensis Lewis & Bowman 1977 HLM86
    |    |--C. catachaetus (Fleming & Steeves 1972) HLM86
    |    |--C. chiapas Bowman 1975 HLM86
    |    |--C. circulus (Steeves 1968) HLM86
    |    |--C. cyrtorhynchus (Fleming & Steeves 1972) HLM86
    |    |--C. dentadactyla HLM86
    |    |--C. dimorpha Mackin & Hubricht 1940 HLM86
    |    |--C. extensolingualus (Fleming 1972) HLM86
    |    |--C. filicispeluncae Bowman & Hobbs 1983 HLM86
    |    |--C. franzi Lewis 1981 HLM86
    |    |--C. fustis Lewis 1981 HLM86
    |    |--C. henroti (Bresson 1955) HLM86
    |    |--C. hobbsi Maloney 1939 HLM86
    |    |--C. holsingeri (Steeves 1963) HLM86
    |    |--C. incurva (Steeves 1968) HLM86
    |    |--C. jordani (Eberly 1965) HLM86
    |    |--C. kendeighi (Steeves & Seidenbergh 1971) HLM86
    |    |--C. lesliei Lewis & Bowman 1981 HLM86
    |    |--C. macropropoda Chase & Blair 1937 [incl. C. ozarkana Chase & Blair 1937] HLM86
    |    |--C. metcalfi (Fleming 1972) HLM86
    |    |--C. mitchelli Argano 1972 HLM86
    |    |--C. nickajackensis Packard 1881 HLM86
    |    |--C. nortoni (Steeves 1966) HLM86
    |    |--C. oculata Mackin & Hubricht 1940 HLM86
    |    |--C. packardi Mackin & Hubricht 1940 HLM86
    |    |--C. parva (Steeves 1964) HLM86
    |    |--C. pasquinii Argano 1972 HLM86
    |    |--C. paurotrigonus (Fleming 1972) HLM86
    |    |--C. pricei Levi 1949 HLM86
    |    |--C. recurvata (Steeves 1963) HLM86
    |    |--C. reddelli (Steeves 1968) HLM86
    |    |--C. richardsonae Hay 1901 HLM86
    |    |--C. rotunda Bowman & Lewis 1984 HLM86
    |    |--C. salemensis Lewis 1981 HLM86
    |    |--C. scrupulosa HLM86
    |    |--C. scyphus (Steeves 1968) HLM86
    |    |--C. sequoiae Bowman 1975 HLM86
    |    |--C. serrata (Fleming 1972) HLM86
    |    |--C. simonini (Bresson 1955) HLM86
    |    |--C. sinuncus (Steeves 1965) HLM86
    |    |--C. spatulata Mackin & Hubricht 1940 HLM86
    |    |--C. steevesi (Fleming 1972) HLM86
    |    |--C. stiladactyla Mackin & Hubricht 1940 HLM86
    |    |--C. stygia Packard 1871 HLM86
    |    |--C. teresae Lewis 1982 HLM86
    |    |--C. tridentata Hungerford 1922 HLM86
    |    |--C. vandeli (Bresson 1955) HLM86
    |    |--C. vomeroi Argano 1972 HLM86
    |    `--C. zullinii Argano 1972 HLM86
    `--Proasellus Dudich 1925 HLM86
         |--P. acutianus Argano & Henry 1972 HLM86
         |--P. adriaticus Argano & Pesce 1979 HLM86
         |--P. albigensis (Magniez 1965) HLM86
         |--P. ambracicus Pesce & Argano 1980 HLM86
         |--P. amiterninus Argano & Pesce 1979 HLM86
         |--P. anophtalmus (Karaman 1934) HLM86
         |    |--P. a. anophtalmus HLM86
         |    |--P. a. bosnicus (Sket 1965) HLM86
         |    |--P. a. dalmatinus (Karaman 1955) HLM86
         |    |--P. a. longisetus (Remy 1941) HLM86
         |    |--P. a. radikanus (Karaman 1955) HLM86
         |    |--P. a. rhausinus (Remy 1941) HLM86
         |    `--P. a. serbicus (Karaman 1955) HLM86
         |--P. aquaecalidae (Racovitza 1922) HLM86
         |--P. arnautovici HLM86
         |    |--P. a. arnautovici HLM86
         |    `--P. a. elongatus (Karaman 1953) HLM86
         |--P. arthrodilus (Braga 1945) HLM86
         |--P. bagradicus Henry & Magniez 1972 HLM86
         |--P. bardaunii Alouf, Henry & Magniez 1982 HLM86
         |--P. basnosanui (Codreanu & Codreanu 1962) HLM86
         |--P. bellesi Henry & Magniez 1982 HLM86
         |--P. beroni Henry & Magniez 1968 HLM86
         |--P. boui Henry & Magniez 1969 HLM86
         |--P. burgundus Henry & Magniez 1969 HLM86
         |--P. cantabricus Henry & Magniez 1968 HLM86
         |--P. cavaticus (Leydig 1871) HLM86
         |--P. chappuisi Henry & Magniez 1968 HLM86
         |--P. chauvini Henry & Magniez 1978 HLM86
         |--P. coiffaiti Henry & Magniez 1972 HLM86
         |--P. comasi Henry & Magniez 1982 HLM86
         |--P. coxalis (Dollfus 1892) HLM86
         |    |--P. c. coxalis HLM86
         |    |--P. c. africanus (Monod 1924) HLM86
         |    |--P. c. banyulensis (Racovitza 1919) HLM86
         |    |--P. c. cephallenus (Strouhal 1942) HLM86
         |    |--P. c. cyrenaicus (Arcangeli 1942) HLM86
         |    |--P. c. gabriellae (Margalef 1950) HLM86
         |    |--P. c. leucadius (Strouhal 1942) HLM86
         |    |--P. c. lucifugus (Deeleman-Reinhold 1965) HLM86
         |    |--P. c. perarmatus (Remy 1936) HLM86
         |    `--P. c. peyerimhoffi (Racovitza 1919) HLM86
         |--P. cretensis Pesce & Argano 1980 HLM86
         |--P. danubialis (Codreanu & Codreanu 1962) HLM86
         |--P. delhezi Henry & Magniez 1973 HLM86
         |--P. deminutus (Sket 1959) HLM86
         |    |--P. d. deminutus HLM86
         |    |--P. d. intermedius (Sket 1965) HLM86
         |    `--P. d. vulgaris (Sket 1965) HLM86
         |--P. elegans (Codreanu & Codreanu 1962) HLM86
         |--P. escolai Henry & Magniez 1982 HLM86
         |--P. espanoli Henry & Magniez 1982 HLM86
         |--P. exiguus Afonso 1983 HLM86
         |--P. franciscoloi (Chappuis 1955) HLM86
         |--P. gardinii (Arcangeli 1942) HLM86
         |--P. gauthieri (Monod 1924) HLM86
         |    |--P. g. gauthieri HLM86
         |    `--P. g. theodori (Birstein 1951) [incl. P. gauthieri numidicus Monod in Gauthier 1928 (n. n.)] HLM86
         |--P. gjorgjevici (Karaman 1933) HLM86
         |--P. gourbaultae Henry & Magniez 1981 HLM86
         |--P. hercegovinensis (Karaman 1933) HLM86
         |--P. hermallensis (Arcangeli 1938) HLM86
         |--P. hussoni (Remy 1941) HLM86
         |    |--P. h. hussoni HLM86
         |    `--P. h. curvifrons (Remy 1941) HLM86
         |--P. hypogeus (Racovitza 1922) HLM86
         |--P. infirmus (Birstein 1936) HLM86
         |--P. istrianus (Stammer 1932) HLM86
         |--P. jaloniacus Henry & Magniez 1977 HLM86
         |--P. karamani (Remy 1934) HLM86
         |--P. lagari Henry & Magniez 1982 HLM86
         |--P. lescherae Henry & Magniez 1977 HLM86
         |--P. linearis (Birstein 1967) HLM86
         |--P. ljovuschkini (Birstein 1967) HLM86
         |--P. lusitanicus (Frade 1938) HLM86
         |--P. lykaonicus Argano & Pesce 1978 HLM86
         |--P. maleri Henry 1977 HLM86
         |--P. margalefi Henry & Magniez 1982 HLM86
         |--P. mateusorum Afonso 1982 HLM86
         |--P. meridianus (Racovitza 1919) HLM86
         |    |--P. m. meridianus HLM86
         |    `--P. m. belgicus (Arcangeli 1935) HLM86
         |--P. minoicus Pesce & Argano 1980 HLM86
         |--P. monodi (Strouhal 1942) HLM86
         |--P. monsferratus (Braga 1948) HLM86
         |--P. montenigrinus (Karaman 1934) HLM86
         |    |--P. m. montenigrinus HLM86
         |    `--P. m. macedonicus (Karaman 1955) HLM86
         |--P. nolli (Karaman 1952) HLM86
         |--P. notenboomi Henry & Magniez 1982 HLM86
         |--P. parvulus (Sket 1960) HLM86
         |--P. patrizzi (Arcangeli 1952) HLM86
         |--P. pavani (Arcangeli 1942) HLM86
         |    |--P. p. pavani HLM86
         |    `--P. p. orientalis (Sket 1965) HLM86
         |--P. racovitzai Henry & Magniez 1972 HLM86
         |--P. rectangulatus Afonso 1982 HLM86
         |--P. rectus Afonso 1982 HLM86
         |--P. remyi [incl. P. remyi f. nudus (Karaman 1953)] HLM86
         |--P. rouchi Henry 1980 HLM86
         |--P. similis (Birstein 1967) HLM86
         |--P. sketi Henry 1975 HLM86
         |--P. slavus (Remy 1948) HLM86
         |    |--P. s. slavus HLM86
         |    |--P. s. histriae (Sket 1963) HLM86
         |    |--P. s. salisburgensis (Strouhal 1957) HLM86
         |    |--P. s. serbiae (Sket 1963) HLM86
         |    |--P. s. styriacus (Sket 1963) HLM86
         |    |--P. s. vindobonensis (Karaman 1955) HLM86
         |    `--P. s. zeii (Sket 1963) HLM86
         |--P. slovenicus (Sket 1957) HLM86
         |--P. solanasi Henry & Magniez 1972 HLM86
         |--P. spelaeus (Racovitza 1922) HLM86
         |--P. spinipes Afonso 1979 HLM86
         |--P. strouhali (Karaman 1955) HLM86
         |    |--P. s. strouhali HLM86
         |    `--P. s. puteanus (Henry 1966) HLM86
         |--P. synaselloides Henry 1963 HLM86
         |--P. thermonyctophilus (Monod 1924) HLM86
         |--P. valdensis (Chappuis 1948) HLM86
         |--P. vandeli Magniez & Henry 1969 HLM86
         |--P. variegatus Afonso 1982 HLM86
         |--P. vignai Argano & Pesce 1977 HLM86
         `--P. walteri (Chappuis 1948) HLM86

*Type species of generic name indicated

REFERENCES

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[H47] Hatch, M. H. 1947. The Chelifera and Isopoda of Washington and adjacent regions. University of Washington Publications in Biology 10 (5): 155–274.

[HLM86] Henry, J.-P., J. J. Lewis & G. Magniez. 1986. Isopoda: Asellota: Aselloidea, Gnathostenetroidoidea, Stenetrioidea. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 434–464. E. J. Brill/Dr. W. Backhuys: Leiden.

[N29] Nicholls, G. E. 1929. A new species of Asellus (A. nipponensis) from Japan. Archives de Zoologie Expérimentale et Générale 68 (1): 33–38.

[T30] Tattersall, W. M. 1930. Asellus cavaticus, a blind isopod new to the British fauna, from a well in Hampshire. Proceedings of the Linnean Society of London 141: 7.

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