Female spoonwing Chasmoptera hutti, copyright Jean & Fred Hort.

Belongs within: Neuroptera.
Contains: Ascalaphidae, Myrmeleontidae, Kalligrammatidae.

The Myrmeleontiformia are a lineage of lacewings including the ant-lions, owlflies and related taxa, united by the presence in larvae of a modified prementum resembling a labial palp segment, and by features of the adult genitalia (Grimaldi & Engel 2005).

<==Myrmeleontiformia [Myrmeleontoidea] GE05
    |--+--Nymphitidae GE05
    |  |--Solenoptilidae GE05
    |  |--Brongniartiellidae GE05
    |  |--Petropsychops [Osmylopsychopidae] GE05
    |  |    `--P. superba GE05
    |  `--+--+--Babinskaiidae GE05
    |     |  |--Ascalaphidae GE05
    |     |  `--Myrmeleontidae GE05
    |     `--Nymphidae GE05
    |          |--Mesonymphes rohdendorfi P02
    |          |--Nymphes GE05
    |          |--Myiodactylus [Myiodactylidae] GE05
    |          `--Osmylops GE05
    `--+--Kalligrammatidae GE05
       `--+--Psychopsidae GE05
          |    |--Calopsychops extinctus P02
          |    |--Kagapsychops continentalis P02
          |    |--Notopsychops zebra (Brauer 1889) (see below for synonymy) HM55
          |    `--Psychopsis insolens ZS10
          `--Nemopteridae GE05
               |  i. s.: Marquettia americana GE05
               |         Nemopistha imperatrix (Westwood 1867) (see below for synonymy) HM55
               |--Laurhervasia [Crocinae] GE05
               |    `--L. setacea GE05
               `--Nemopterinae GE05
                    |--Nemoptera bipennis GE05
                    `--Chasmoptera Kirby 1900 K67
                         |--*C. hutti (Westwood 1848) [=Nemoptera huttii] K67
                         |--C. mathewsi Koch 1967 K67
                         `--C. superba Tillyard 1925 K67

Myrmeleontiformia incertae sedis:
  Siniphes Ren & Yin 2002 RY02
    `--*S. delicatus Ren & Yin 2002 RY02
  Panfilovia Makarkin 1980 [Panfilovidae] R02
  Grammolingiidae R02
    |--Grammolingia Ren 2002 R02
    |    `--*G. boi Ren 2002 R02
    |--Leptolingia Ren 2002 R02
    |    |--*L. jurassica Ren 2002 R02
    |    `--L. tianyiensis Ren 2002 R02
    `--Litholingia Ren 2002 R02
         |--*L. rhora Ren 2002 R02
         |--L. eumorpha Ren 2002 R02
         `--L. polychotoma Ren 2002 R02

Nemopistha imperatrix (Westwood 1867) [=Nemoptera imperatrix; incl. Halter remipennis Kirby 1900] HM55

Notopsychops zebra (Brauer 1889) [=Psychopsis zebra, Psychomorphe zebra, Zygophlebius zebra; incl. P. felina Navas 1912, Z. leoninus Navas 1910] HM55

*Type species of generic name indicated


[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[HM55] Handschin, E. & W. Markl. 1955. Neuropteren aus Angola. Diamang Publicações Culturais 27: 65–82.

[K67] Koch, L. E. 1967. The genus Chasmoptera (Neuroptera: Nemopteridae), with the description of a new species from Western Australia. Proceedings of the Royal Entomological Society, London (B) 36 (9-10): 137–146.

[P02] Ponomarenko, A. G. 2002. Superorder Myrmeleontidea Latreille, 1802 (=Neuropteroidea Handlirsch, 1903). In: Rasnitsyn, A. P. & D. L. J. Quicke (eds) History of Insects pp. 176–189. Kluwer Academic Publishers: Dordrecht.

[R02] Ren D. 2002. A new lacewing family (Neuroptera) from the Middle Jurassic of Inner Mongolia, China. Entomologica Sinica 9 (4): 53–67.

[RY02] Ren D. & Yin J.-C. 2002. A new genus and new species of lacewings in the Jurassic of China (Neuroptera: Myrmeleontoidea). Acta Zootaxonomica Sinica 27 (2): 269–273.

[ZS10] Zborowski, P. & R. Storey. 2010. A Field Guide to Insects in Australia. Reed New Holland: Sydney.

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