Sacoglossa

Cylindrobulla sp., from Sea Slugs of Hawai'i.

Belongs within: Heterobranchia.
Contains: Plakobranchidae.

The Sacoglossa are a group of algal-feeding gastropods characterised by a muscular, suctorial pharynx and uniseriate radula.

<==Sacoglossa [Sacoglossida] BR17
    |--Oxynooidea [Conchoidea, Julioidea, Oxynoacea, Oxynoiformes] BR05
    |    |--Cylindrobulla Fischer 1857 BR05 (see below for synonymy)
    |    |    |--*C. beauii Fischer 1857 BR17
    |    |    |--C. fischeri Adams & Angas 1865 H09
    |    |    |--C. gigas Mikkelsen 1998 BC01
    |    |    |--C. phuketi Jensen 1989 J96
    |    |    `--C. pusilla Nevill 1869 H09
    |    |--Oxynoidae [Oxynoeidae] BR05
    |    |    |--Oxynoe Rafinesque 1814 BR05
    |    |    |    `--*O. olivacea Rafinesque 1814 BR17
    |    |    |--Icarus Forbes 1844 [Icaridae, Icarina, Icarinae] BR05
    |    |    |    `--*I. gravesi Forbes 1844 BR17
    |    |    |--Lophocercus Krohn 1847 [=Lephocercus (l. c.); Lephocercina, Lophocercidae, Lophocercinae] BR05
    |    |    |    `--*L. sieboldii Krohn 1847 BR17
    |    |    `--Lobiger Krohn 1847 [Lobigeridae, Lobigerina] BR05
    |    |         `--*L. philippii Krohn 1847 BR17
    |    |--Volvatellidae [Arthessidae, Arthessoidea, Volvatellacea, Volvatellina, Volvatelloidea, Volvatelloidei] BR05
    |    |    |--Arthessa Evans 1950 BR05
    |    |    |    `--*A. cincta (Nevill & Nevill 1869) [=Volvatella cincta] BR17
    |    |    |--Volvatella Pease 1860 [Volvatellinae] BR05
    |    |    |    |--*V. fragilis Pease 1860 BR17
    |    |    |    `--V. pyriformis Pease 1860 H09
    |    |    `--Ascobulla Marcus 1972 [Ascobullacea, Ascobullidae] BR05
    |    |         |--*A. ulla (Marcus & Marcus 1970) [=Cylindrobulla ulla] BR17
    |    |         `--A. fischeri (Adams & Angas 1864) J96
    |    `--Juliidae [Juliacea] BR05
    |         |--Gougerotia Le Renard 1980 [Gougerotiinae] BR05
    |         |    `--*G. orthodonta Le Renard 1980 BR17
    |         |--Berthelinia Crosse 1875 (see below for synonymy) BR05
    |         |    |--*B. elegans Crosse 1875 BR17
    |         |    `--‘*Tamanovalva’ limax Kawaguti & Baba 1959 BR17
    |         `--Juliinae BR05
    |              |--Prasina Deshayes 1863 [Prasinidae] BR05
    |              |    `--*P. borbonica Deshayes 1863 BR17
    |              `--Julia Gould 1862 BR05
    |                   `--*J. exquisita Gould 1862 BR17
    |--Plakobranchoidea (see below for synonymy) BR05
    |    |--Jenseneria Ortea & Moro 2015 [Jenseneriidae] BR17
    |    |    `--*J. borgninii (Trinchese 1896) [=Phyllobranchus borgninii] BR17
    |    |--Costasiella Pruvot-Fol 1951 BR05 [Costasiellidae BR17]
    |    |    `--*C. virescens Pruvot-Fol 1951 BR17
    |    |--Hermaeidae [Caliphyllacea, Caliphyllidae, Caliphyllina, Phyllobranchidae, Phyllobranchillidae] BR17
    |    |    |--Caliphylla Costa 1867 BR17, BR05
    |    |    |    `--*C. mediterranea Costa 1867 BR17
    |    |    |--Phyllobranchillus Pruvot-Fol 1933 [=Phyllobranchus Alder & Hancock 1864 non Girard 1851] BR05
    |    |    |    `--*P. orientalis (Kelaart 1858) [=Proctonotus orientalis, *Phyllobranchus orientalis] BR17
    |    |    |--Polybranchia Pease 1860 (see below for synonymy) BR05
    |    |    |    |--*P. pellucida Pease 1860 [=*Lobifera pellucida] BR17
    |    |    |    `--P. westralis Jensen 1993 BW09
    |    |    `--Hermaea Lovén 1844 [Hermaeinae] BR05
    |    |         |--*H. bifida (Montagu 1815) [=Doris bifida] BR17
    |    |         |--H. aoteana Powell 1937 P61
    |    |         `--H. dendritica P61
    |    |--Limapontiidae [Limapontiadae, Stiligeridae, Stiligerinae, Stiligeroidea] BR05
    |    |    |--Calliopaea d’Orbigny 1837 APS-P86
    |    |    |--Limapontia Johnston 1836 BR05 [=Pontolimax Creplin 1848 BR17; Limapontiinae, Pontolimacidae]
    |    |    |    |--*L. nigra Johnston 1836 BR17
    |    |    |    `--L. depressa B64
    |    |    |--Olea Agersborg 1923 [Oleidae] BR05
    |    |    |    `--*O. hansineensis Agersborg 1923 BR17
    |    |    |--Alderia Allmann 1845 [Alderiidae] BR05
    |    |    |    `--*A. modesta (Lovén 1844) [=Stiliger modestus] BR17
    |    |    |--Ercolania Trinchese 1872 [Ercolaniinae] BR05
    |    |    |    |--*E. siottii Trinchese 1872 BR17
    |    |    |    `--E. funerea (Costa 1867) H01
    |    |    `--Stiliger Ehrenberg 1828 BR05
    |    |         |--*S. ornatus Ehrenberg 1828 BR17
    |    |         |--S. bellulus [=Calliopaea bellula] P61
    |    |         |--S. felinus (Hutton 1882) P61
    |    |         `--S. vossi CG99
    |    `--Plakobranchidae BR05
    `--Platyhedylidae [Platyhedylacea, Platyhedyloidea] BR17
         |--Gascoignella Jensen 1985 [Gascoignellidae] BR05
         |    `--*G. aprica Jensen 1985 BR17
         `--Platyhedyle Salvini-Plawen 1973 BR05
              `--*P. denudata Salvini-Plawen 1973 BR17

Berthelinia Crosse 1875 [incl. Tamanovalva Kawaguti & Baba 1959; Bertheliniidae, Bertheliniinae, Tamanovalvacea, Tamanovalvidae, Tamanovalvoidei] BR05

Cylindrobulla Fischer 1857 BR05 [Cylindrobullacea, Cylindrobullida, Cylindrobullidae BR17, Cylindrobullina, Cylindrobullinae, Cylindrobulloidea]

Plakobranchoidea [Aconchoidea, Ascoglossa, Cirrobranchia, Elysiacea, Elysiina, Elysiomorpha, Limapontioidea, Pellibranchiata, Phyllobranchiacea, Placobranchacea, Placobranchoidea, Plakobranchacea, Stiligerida, Stiligerina] BR05

Polybranchia Pease 1860 [=Lobifera Pease 1860; Lobiferidae, Polybranchidae, Polybranchiidae, Polybranchiinae, Polybranchioidea] BR05

*Type species of generic name indicated

REFERENCES

[APS-P86] Arnaud, P. M., C. Poizat & L. v. Salvini-Plawen. 1986. Marine-interstitial Gastropoda (including one freshwater interstitial species). In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 153–176. E. J. Brill/Dr W. Backhuys: Leiden.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[H01] Huys, R. 2001. Splanchnotrophid systematics: a case of polyphyly and taxonomic myopia. Journal of Crustacean Biology 21 (1): 106–156.

[J96] Jensen, K. R. 1996. The Diaphanidae as a possible sister group of the Sacoglossa (Gastropoda, Opisthobranchia). In: Taylor, J. D. (ed.) Origin and Evolutionary Radiation of the Mollusca pp. 231–247. Oxford University Press: Oxford.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4^th ed. Whitcombe and Tombs Limited: Christchurch.