Strombus

Fighting conch Strombus pugilis, copyright H. Zell.

Belongs within: Strombidae.

Strombus, the conches, is a pantropical genus of large marine gastropods characterised by a shell with a large, flaring outer lip bearing a U-shaped notch near the anterior end.

<==Strombus Linnaeus 1758 [Strombinae, Strombini] BR05
    |--*S. (Strombus) pugilis Linnaeus 1758 BR17
    |--S. (Dolomena Iredale 1931) WG71
    |    |--S. (D.) dilatatus Swainson 1821 WG71
    |    |    |--S. d. dilatatus WG71
    |    |    `--S. d. swainsoni Reeve 1850 WG71
    |    |--S. (D.) plicatus (Röding 1798) WG71 [=Lambis plicata H09, S. columba (nom. inv.) H08]
    |    |    |--S. p. plicatus WG71
    |    |    `--S. p. pulchellus Reeve 1851 WG71
    |    `--S. (D.) variabilis Swainson 1820 WG71
    |--S. (Doxander Iredale 1931) WG71
    |    |--S. (D.) campbelli Griffith & Pidgeon 1834 WG71 [incl. Alaba sulcata H08]
    |    `--S. (D.) vittatus Linnaeus 1758 WG71
    |--S. (Euprotomus Gill 1870) WG71
    |    |--S. (E.) aurisdianae Linnaeus 1758 WG71
    |    |    |--S. a. aurisdianae WG71
    |    |    `--S. a. aratrum Röding 1798 WG71
    |    `--S. (E.) vomer Röding 1798 WG71
    |         |--S. v. vomer WG71
    |         `--S. v. iredalei Abbott 1960 WG71
    |--S. (Gibberulus Jousseaume 1888) WG71
    |    `--S. (G.) gibberulus Linnaeus 1758 WG71
    |         |--S. g. gibberulus WG71
    |         `--S. g. gibbosus Röding 1798 WG71
    |--S. (Labiostrombus Oostingh 1925) WG71
    |    `--S. (L.) epidromis Linnaeus 1758 WG71
    `--S. (Lentigo Jousseaume 1866) WG71
         |--S. (L.) lentiginosus Linnaeus 1758 WG71
         `--S. (L.) pipus Röding 1798 WG71

Strombus incertae sedis:
  S. alatus Gmelin 1790 S11
  S. australis Gray 1827 H09
  S. belutschiensis M01
  S. bituberculatus [incl. S. peruvianus] C64
  S. bubonius P75
  S. canalis Coquebert & Brongniart 1793 CB93
  S. cancellatus Valenciennes 1833 C64
  S. chiragra Linnaeus 1758 L58
  S. decorus (Roeding 1798) BD86
    |--S. d. decorus BD86
    |--S. d. persicus Swainson 1821 BD86
    `--S. d. raybaudi Nicolay & Manoja 1983 BD86
  S. dentatus Linnaeus 1758 H09
  S. elegans Sowerby 1842 H09
  S. fissura Coquebert & Brongniart 1793 CB93
  S. fragilis (Röding 1798) BW09
  S. galeatus C64
  S. gallus Linnaeus 1758 L58
  S. gigas Linnaeus 1758 L58 [incl. S. goliath G79]
  S. gracilior Sowerby 1825 S11
  S. granulatus C64
  S. laciniatus Dillwyn 1817 H09
  S. latissimus Linnaeus 1758 BW09
  S. lucifer Linnaeus 1758 L58
  S. marginatus Linnaeus 1758 L58
  S. melanostomus Swainson 1822 H09
  S. microurceus Kira 1959 BW09
  S. millepeda Linnaeus 1758 L58
  S. papilio Dillwyn 1817 H09
  S. preoccupatus Finlay 1927 [=S. spinosus Martin 1899 non Linnaeus 1767] F27
  S. raninus KB99
  S. scorpius Linnaeus 1758 L58
  S. sibbaldii Sowerby 1842 H09
  S. spinosus Linnaeus 1767 F27
  S. tricornis P75
  S. wilsoni Abbott 1967 WG71

*Type species of generic name indicated

REFERENCES

[BD86] Barash, A., & Z. Danin. 1986. Further additions to the knowledge of Indo-Pacific Mollusca in the Mediterranean Sea (Lessepsian migrants). Spixiana 9 (2): 117–141.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BW09] Bryce, C., & C. Whisson. 2009. The macromolluscs of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, Western Australia. Records of the Western Australian Museum Supplement 77: 177–208.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[CB93] Coquebert, R., & A. Brongniart. 1793. Extrait d'un mémoire sur la formation de la coquille du Strombus fissurella, et sur deux espèces analogues à celle-ci. Bulletin de la Société Philomatique, a ses Correspondans 25: 1–2.

[F27] Finlay, H. J. 1927. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488–533.

[G79] Guppy, R. J. L. 1879. First sketch of a marine invertebrate fauna of the Gulf of Paria and its neighbourhood. Part I.—Mollusca. Journal of Conchology 2: 151–172.

[H08] Hedley, C. 1908. Studies on Australian Mollusca. Part X. Proceedings of the Linnean Society of New South Wales 33: 456–489, pls 7–10.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[KB99] Kiel, S., & K. Bandel. 1999. The Pugnellidae, a new stromboidean family (Gastropoda) from the Upper Cretaceous. Paläontologische Zeitschrift 73 (1-2): 47–58.

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[M01] Melville, J. C. 1901. A few further remarks upon the Erythrean molluscan fauna, with descriptions of seven species from Aden, in the collection of Commander E. R. Shopland, R.I.M. Annals and Magazine of Natural History, series 7, 7: 550–556.

[P75] Por, F. D. 1975. Pleistocene pulsation and preadaptation of biotas in Mediterranean seas: consequences for Lessepsian migration. Systematic Zoology 24 (1): 72–78.

[S11] Simone, L. R. L. 2011. Phylogeny of the Caenogastropoda (Mollusca), based on comparative morphology. Arquivos de Zoologia 42 (4): 161–323.

[WG71] Wilson, B. R., & K. Gillett. 1971. Australian Shells: illustrating and describing 600 species of marine gastropods found in Australian waters. A. H. & A. W. Reed: Sydney.