Strombidae

Elegant conch Canarium erythrinum, copyright H. Zell.

Belongs within: Stromboidea.
Contains: Lambis, Strombus.

The Strombidae are a group of marine herbivorous gastropods characterised by a flared or thickened outer lip on the adult shell, and a sickle-shaped operculum that is used as a lever by the animal when moving (Wilson & Gillett 1971).

Characters (from Wilson & Gillett 1971): Shell bearing a flaring or thickened outer lip when mature, with deep U-shaped notch near anterior end. Columella smooth. Eyes large, located on short branches near end of usually long eye stalks; right eye extended through anterior notch on lip. Operculum long, sharp, sickle-shaped, usually serrated on one side; located on posterior protuberance of foot.

<==Strombidae [Alatidae, Strombeae, Strombia, Strombina, Strombusidae]
    |  i. s.: Pyramis Röding 1798 BR05
    |         Laevistrombus Kira 1955 WG71
    |           |--L. canarium (Linnaeus 1758) [=Strombus canarium] WG71
    |           `--L. guidoi (Man in’t Veld & De Turck 1998) PK11
    |         Pterocera Lamarck 1799 [Pteroceridae] BR17
    |           |--*P. lambis (Linnaeus 1758) [=Strombus lambis] BR17
    |           `--P. bryonia (Gmelin 1791) [=Strombus bryonius] H09
    |         Dientomochilus stueri Cossmann 1904 TTE93
    |--Canarium Schumacher 1817 S11, WG71 [Canariini]
    |    |--*C. ustulatum Schumacher 1817 BR17 [=Strombus ustulatus H09]
    |    |--C. erythrinum (Dillwyn 1817) [=Strombus (Canarium) erythrinus] WG71
    |    |--C. labiatum (Röding 1798) [=Strombus (Canarium) labiatus] WG71
    |    |--C. mutabile (Swainson 1821) WG71 [=Strombus (Canarium) mutabilis BC01]
    |    |    |--C. m. mutabile BC01
    |    |    `--C. m. ochroglottis (Abbott 1960) [=Strombus (Canarium) mutabilis ochroglottis] BC01
    |    `--C. urceus (Linnaeus 1758) S11 [=Strombus urceus BC01]
    |         |--C. u. urceus BC01
    |         `--C. u. orrae (Abbott 1960) [=Strombus (Canarium) urceus orrae] BC01
    `--+--Conomurex Fischer 1884 S11, WG71
       |    `--C. luhuanus (Linnaeus 1758) S11 [=Strombus luhuanus WG71]
       `--+--Tricornis Jousseaume 1886 S11, WG71
          |    |--T. oldi (Emerson 1965) [=Strombus (Tricornis) oldi] BC01
          |    |--T. raninus (Gmelin 1791) S11
          |    `--T. sinuatus (Humphrey 1786) [=Strombus (Tricornis) sinuatus] WG71
          `--+--Lambis S11
             `--+--Strombus S11
                `--+--Aliger S11
                   |    |--A. costatus (Gmelin 1791) S11
                   |    `--A. gallus (Linné 1758) S11
                   `--Eustrombus S11
                        |--E. gigas (Schröter 1805) S11
                        |--E. goliath (Schroter 1805) S11
                        `--E. mayacensis (Tucker & Wilson 1933) [=Strombus (Eustrombus) macayensis] P89

*Type species of generic name indicated

REFERENCES

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[P89] Petuch, E. J. 1989. New species of Malea (Gastropoda: Tonnidae) from the Pleistocene of southern Florida. Nautilus 103 (3): 92–95.

[PK11] Puillandre, N., Y. I. Kantor, A. Sysoev, A. Couloux, C. Meyer, T. Rawlings, J. A. Todd & P. Bouchet. 2011. The dragon tamed? A molecular phylogeny of the Conoidea (Gastropoda). Journal of Molluscan Studies 77: 259–272.

[S11] Simone, L. R. L. 2011. Phylogeny of the Caenogastropoda (Mollusca), based on comparative morphology. Arquivos de Zoologia 42 (4): 161–323.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

[WG71] Wilson, B. R., & K. Gillett. 1971. Australian Shells: illustrating and describing 600 species of marine gastropods found in Australian waters. A. H. & A. W. Reed: Sydney.