Atriplex

Sprawling saltbush Atriplex suberecta, copyright Forest & Kim Starr.


Belongs within: Chenopodiaceae.
Contains: Atriplex subsection Arenariae.

Atriplex, saltbushes, is a genus of herbs and shrubs found in temperate to subtropical regions worldwide, commonly growing in alkaline or saline soils (Hickman 1993).

Characters (from Hickman 1993): Annual (generally monoecious) to shrub (generally dioecious), often scaly. Leaves generally alternate, gen entire; lower leaves generally more or less short-petioled; upper leaves generally sessile, more or less reduced. Staminate inflorescence a spike or spheric cluster; bracts absent. Pistillate inflorescence clusters to spike- or panicle-like; bracts 2 per fruit, free to fused, generally compressed, generally sessile. Staminate flower with three to five calyx lobes; three to five stamens. Pistillate flower with calyx more or less absent; ovary ovoid to spheric, two style branches. Seed generally erect.

<==Atriplex L. 1753 A61
    |--A. sect. Obione (Gaertner) Reichenb. 1828 FO03
    |    |  i. s.: A. (sect. *Obione) sibirica FO03
    |    |         A. hillmanii (Jones) Standley 1916 FO03 [=A. argentea var. hillmanii H93]
    |    |--A. subsect. Arenariae FO03
    |    |--A. subsect. Argenteae (Standley) Welsh 2000 [incl. Atriplex subsect. Saccariae Standley 1916] FO03
    |    |    |--A. (subsect. *Argenteae) argentea Nuttall 1818 FO03
    |    |    |    |--A. a. var. argentea H93
    |    |    |    `--A. a. var. mohavensis [incl. A. argentea ssp. expansa] H93
    |    |    `--A. saccaria Watson 1874 [=A. (subsect. *Saccariae) saccaria] FO03
    |    |--A. subsect. Graciliflorae (Standley) Welsh 2000 FO03
    |    |    `--A. (subsect. *Graciliflorae) graciliflora Jones 1895 FO03
    |    |--A. subsect. Leucophyllae (Standley) Welsh 2000 FO03
    |    |    `--A. (subsect. *Leucophyllae) leucophylla (Moquin-Tandon) Dietrich 1852 FO03
    |    |--A. subsect. Powellianae (Standley) Flores Olvera 2003 FO03
    |    |    `--A. (subsect. *Powellianae) powellii Watson 1874 FO03
    |    |--A. subsect. Pusillae (Standley) Welsh 2000 FO03
    |    |    |--A. (subsect. *Pusillae) pusilla (Torrey) Watson 1874 FO03
    |    |    |--A. cordulata Jeps. 1892 FO03
    |    |    |--A. depressa Jeps. 1892 FO03
    |    |    |--A. parishii Watson 1882 FO03
    |    |    |--A. tularensis Coville 1893 FO03
    |    |    `--A. wolfii Watson 1874 FO03
    |    `--A. subsect. Truncatae (Standley) Welsh 2000 FO03
    |         `--A. (subsect. *Truncatae) truncata (Torrey ex Watson) Gray 1872 FO03
    |--A. sect. Pleianthae Welsh 2000 FO03
    |    `--A. (sect. *Pleianthae) pleiantha FO03
    `--A. (sect. Semibaccatae) suberecta DH98 [incl. A. muelleri GR98]

Atriplex incertae sedis:
  A. acanthocarpa BTA75
  A. acutibractea H90
  A. acutiloba H90
  A. amnicola KM08
  A. angulata H90
  A. australasica B00
  A. buchananii (Kirk) Cheesem. 1906 [=Chenopodium buchanani Kirk 1898] A61
    |--A. b. var. buchananii A61
    `--A. b. var. tenuicaulis Petrie 1915 A61
  A. californica H93
  A. canescens [incl. A. canescens var. laciniata] H93
    |--A. c. ssp. canescens [incl. A. canescens var. macilenta] H93
    `--A. c. ssp. linearis H93
  A. cinerea FS54
  A. conduplicata [incl. A. lindleyi ssp. conduplicata] H90
  A. confertifolia C90
  A. crassipes H90
  A. eardleyae H90
  A. eichleri B00
  A. elachophylla B00
  A. fissivalvis H90
  A. gardneri CL86
    |--A. g. var. gardneri H93
    `--A. g. var. falcata [=A. nuttallii var. falcata] H93
  A. halimus C55
  A. hastata G60
  A. heterosperma H93
  A. holocarpa GR98
  A. hortensis V09
    |--A. h. var. hortensis V09
    `--A. h. var. rubra V09
  A. hymenelytra H93
  A. inflata [=A. lindleyi ssp. inflata] H06
  A. infrequens Wilson 1984 [incl. A. microcarpa (nom. illeg.)] DH98
  A. intermedia H90
  A. joaquiniana [incl. A. patula ssp. spicata] H93
  A. lampa RJ11
  A. lasiantha DH98
  A. lentiformis BT72
    |--A. l. ssp. lentiformis [incl. A. lentiformis ssp. breweri] H93
    `--A. l. ssp. torreyi H93
  A. leptocarpa M99
  A. leucoclada DH98
  A. limbata B00
  A. lindleyi [incl. A. halimoides, A. lindleyi ssp. quadripartita] H90
  A. lobativalvis H90
  A. macropterocarpa H90
  A. minuscula H93
  A. morrisii H90
  A. nessorhina H90
  A. nitens C55
  A. novae-zelandiae Aellen 1940 [=A. hastata ssp. novae-zelandiae ms; incl. A. hastata var. angusta ms] A61
  A. nummularia [incl. A. nummularia ssp. omissa] H90
  A. obconica B00
  A. paludosa KM08
    |--A. p. ssp. paludosa KM08
    `--A. p. ssp. moquiniana KM08
  A. papillata H90
  A. parryi H93
  A. patula WH02
    |--A. p. var. patula H93
    `--A. p. var. obtusa H93
  A. pendunculata CS77
  A. phyllostegia H93
  A. polycarpa BT72
  A. portulacoides Y98
  A. pseudocampanulata [incl. A. campanulata var. inappendiculata] H90
  A. pterocarpa H90
  A. pumilio [incl. A. prostrata] H90
  A. quinii B00
  A. rhagodioides B00
  A. rosea H93
  A. sagittata K09
  A. semibaccata GR98
  A. semilunaris KM08
  A. spinibractea H90
  A. spinifera H93
  A. spongiosa B00
  A. stipitata H90
  A. sturtii H90
  A. subcordata F92
  A. subspicata H93
  A. tornabensis F92
  A. triangularis H93
  A. turbinata [=A. leptocarpa f. turbinata] H90
  A. vallicola H93
  A. velutinella B00
  A. vesicaria KM08
    |--A. v. ssp. vesicaria H90
    |--A. v. ssp. calcicola H90
    |--A. v. ssp. macrocystidia H90
    |--A. v. ssp. minor H90
    `--A. v. ssp. sphaerocarpa H90
  A. watsonii H93

*Type species of generic name indicated

REFERENCES

[A61] Allan, H. H. 1961. Flora of New Zealand vol. 1. Indigenous Tracheophyta: Psilopsida, Lycopsida, Filicopsida, Gymnospermae, Dicotyledones. R. E. Owen, Government Printer: Wellington (New Zealand).

[BT72] Baker, E. W., & D. M. Tuttle. 1972. New species and further notes on the Tetranychoidea mostly from the southwestern United States (Acarina: Tetranychidae and Tenuipalpidae). Smithsonian Contributions to Zoology 116: 1–37.

[BTA75] Baker, E. W., D. M. Tuttle & M. J. Abbatiello. 1975. The false spider mites of northwestern and north central Mexico (Acarina: Tenuipalpidae). Smithsonian Contributions to Zoology 194: 1–23.

[B00] Braby, M. F. 2000. Butterflies of Australia: their identification, biology and distribution vol. 2. CSIRO Publishing: Collingwood (Victoria).

[C55] Candolle, A. de. 1855. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 2. Librairie de Victor Masson: Paris.

[CL86] Collins, E. I., & R. W. Lichvar. 1986. Vegetation inventory of current and historic black-footed ferret habitat in Wyoming. Great Basin Naturalist Memoirs 8: 85–93.

[CS77] Cramp, S., & K. E. L. Simmons (eds) 1977. Handbook of the Birds of Europe, the Middle East and North Africa: The Birds of the Western Palaearctic vol. 1. Ostrich to Ducks. Oxford University Press: Oxford.

[C90] Crawford, C. S. 1990. Scorpiones, Solifugae, and associated desert taxa. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 421–475. John Wiley & Sones: New York.

[DH98] Danin, A., & I. C. Hedge. 1998. Contributions to the flora of Jordan 2. A new species of Satureja (Labiatae) and some new records. Willdenowia 28: 135–142.

[FS54] Fairbridge, R. W., & V. N. Serventy. 1954. Physiography. In: Willis, J. H. (ed.) The Archipelago of the Recherche pp. 9–28. Australian Geographical Society: Melbourne.

[F92] Fan Z. 1992. Key to the Common Flies of China 2nd ed. Science Press: Beijing.

[FO03] Flores Olvera, H. 2003. Classification of North American species of Atriplex section Obione (Chenopodiaceae) based on numerical taxonomic analysis. Taxon 52: 247–260.

[G60] Gillham, M. E. 1960. Vegetation of Little Brother Island, Cook Strait, in relation to spray-bearing winds, soil salinity, and pH. Transactions of the Royal Society of New Zealand 88 (3): 405–424.

[GR98] Greuter, W., & T. Raus (eds.) 1998. Med-Checklist Notulae, 17. Willdenowia 28: 163–174.

[H90] Harden, G. J. (ed.) 1990. Flora of New South Wales vol. 1. New South Wales University Press.

[H06] Henderson, L. 2006. Comparisons of invasive plants in southern Africa originating from southern temperate, northern temperate and tropical regions. Bothalia 36 (2): 201–222.

[H93] Hickman, J. C. (ed.) 1993. The Jepson Manual: Higher Plants of California. University of California Press: Berkeley (California).

[KM08] Keighery, G. J., & W. Muir. 2008. Vegetation and vascular flora of Faure Island, Shark Bay, Western Australia. Records of the Western Australian Museum Supplement 75: 11–19.

[K09] Köhler, G. 2009. Heuschrecken (Saltatoria) und Ohrwürmner (Dermaptera) im Immissionsgebiet des Düngemittelwerkes Steudnitz/Thüringen—eine Langzeitstudie (1978–2001). Mauritiana 20 (3): 601–646.

[M99] Matthews, M. 1999. Heliothine Moths of Australia: A guide to bollworms and related noctuid groups. CSIRO Publishing.

[RJ11] Rising, J. D., A. Jaramillo, J. L. Copete, P. G. Ryan & S. C. Madge. 2011. Family Emberizidae (buntings and New World sparrows). In: Hoyo, J. del, A. Elliott & D. A. Christie (eds) Handbook of the Birds of the World vol. 16. Tanagers to New World Blackbirds pp. 428–683. Lynx Edicions: Barcelona.

[V09] Verdcourt, B. (ed.) 2009. Additions to the Wild Fauna and Flora of the Royal Botanic Gardens, Kew. XXVI. Miscellaneous records. Kew Bulletin 64 (1): 183–194.

[WH02] Wolfert, H. P., P. W. F. M. Hommel, A. H. Prins & M. H. Stern. 2002. The formation of natural levees as a disturbance process significant to the conservation of riverine pastures. Landscape Ecology 17 (Suppl. 1): 47–57.

[Y98] Yannitsaros, A. 1998. Additions to the flora of Kithira (Greece) I. Willdenowia 28: 77–94.

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