Belongs within: Schizophora.
The Milichiidae are a group of small flies that are mostly saprophages or coprophages though a number of species are associated with social Hymenoptera as inquilines or kleptoparasites (Swann 2010). Members are divided between the subfamilies Madizinae, with a prominent vibrissal angle, the vibrissa below the lower margin of the eye, and without a prominent anal area on the wing, and the Milichiinae, with the vibrissal angle scarcely evident and the vibrissa above the level of the lower margin of the eye, and a conspicuously broad anal area on the wing (Sabrosky 1987).
Characters (from Swann 2010): Small (body length 1–7 mm), often largely brown or black, occasionally orange or yellow; males of some genera with silvery microtomentum on abdomen. Usually with two or three pairs of medioclinate frontal bristles and two or three pairs of lateroclinate to proclinate orbital bristles; rarely with uniform row of 8–10 pairs of reclinate fronto-orbital bristles or frontal and orbital setae greatly reduced to absent. Frons usually with two rows of interfrontal setae, sometimes on distinctly shining stripes; lunule often with one or two pairs of setae; proboscis often elongate, geniculate. Wing with humeral and subcostal break; region of latter often modified into costal lappet; cell cup closed, small. Male terminalia with pregonites fused with hypandrium and postgonites absent.
<==Milichiidae [Phyllomyzidae] S10
| i. s.: Costalima myrmicola S10
| Pseudomilichia S10
| Ulia poecilogastra S10
| Microsimus S10
| Prosaetomilichia O98
|--Milichiinae S10
| |--Eusiphona S87
| | |--E. mira S10
| | `--E. vittata S10
| |--Milichiella S87 [incl. Eccoptomma S10]
| | |--M. bisignata S10
| | `--M. lacteipennis S10
| |--Milichia S10
| | |--M. aethiops S10
| | |--M. albomaculata (Strobl 1900) C-T92
| | |--M. piscivora CM91
| | `--M. speciosa Meigen 1830 C-T92
| `--Pholeomyia S10
| |--P. comans S87
| |--P. dampfi S10
| |--P. indecora BW09
| `--P. leucozona S10
`--Madizinae [Phyllomyzinae] S10
|--Madiza glabra Fallén 1820 C-T92
|--Paramyia nitens S87
|--Aldrichiomyza agromyzina S87
|--Stomosis S87
| |--S. innominata S10
| `--S. rufula S10
|--Desmometopa S87
| |--D. m-nigrum (Zetterstedt 1848) C-T92
| |--D. sordida (Fallén 1820) C-T92
| `--D. varipalpis Malloch 1927 C-T92
|--Leptometopa S87
| |--L. halteralis S10
| `--L. latipes (Meigen 1830) C-T92 [=Desmometopa latipes S10]
|--Phyllomyza S10
| |--P. hurdi S87
| |--P. jaegeri S87
| `--P. securicornis S10
`--Neophyllomyza Melander 1913 S87, C-T92
|--N. acyglossa (Villeneuve 1920) C-T92
|--N. magnipalpus S10
|--N. quadricornis S10
`--N. wulpi S10
*Type species of generic name indicated
REFERENCES
[BW09] Buck, M., N. E. Woodley, A. Borkent, D. M. Wood, T. Pape, J. R. Vockeroth, V. Michelsen & S. A. Marshall. 2009. Key to Diptera families—adults. In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 95–156. NRC Research Press: Ottawa.
[C-T92] Carles-Tolrá, M. 1992. New and interesting records of Diptera Acalyptrata from Spain. Part I: Acartophthalmidae, Opomyzidae, Anthomyzidae, Asteiidae, Carnidae, Tethinidae, Milichiidae and Cryptochetidae. Bull. Annls Soc. R. Belge Ent. 128: 343–353.
[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).
[O98] Oosterbroek, P. 1998. The Families of Diptera of the Malay Archipelago. Brill: Leiden.
[S87] Sabrosky, C. W. 1987. Milichiidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 903–908. Research Branch, Agriculture Canada.
[S10] Swann, J. E. 2010. Milichiidae (milichiid flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 1125–1136. NRC Research Press: Ottawa.
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