Heptageniomorpha

Female (above) and male of Ecdyonurus torrentis, copyright crnabambula.


Belongs within: Pisciformia.

The Heptageniomorpha is a group of mayflies in which the nymphs bear filamentous gills on the sides of the abdomen.

Synapomorphies (from Kluge & Sinitshenkova 2002): Posterior arms of mesothoracic prealar bridge reduced in adults; eggs with anchors of peculiar structure; nymphal maxillae with ventral longitudinal row of setae; nymphal labial palp with segments 2 and 3 fused; nymphal tergaliae with gill filaments.

<==Heptageniomorpha [Branchitergaliae]
    |--Oligoneurioidea [Eusetisura] KS02
    |    |--Isonychiidae KS02
    |    |    |  i. s.: Siphlurus sanukensis Takahashi 1929 TYM08
    |    |    |--Rallidentinae D70b
    |    |    `--Isonychiinae D70b
    |    |--Oligoneuriidae KS02
    |    |    |  i. s.: Colocrurus KS02
    |    |    |--Chromarcys KS02
    |    |    `--Oligoneuriinae KS02
    |    `--Coloburiscidae [Coloburiscinae] KS02
    |         |--Coloburiscus humeralis PC66
    |         |--Murphyella needhami D70b
    |         |--Coloburiscoides D70b
    |         `--Cronicus Eaton 1871 D70a
    |              `--C. major Demoulin 1968 D70a
    `--Heptagenioidea [Heptagennota] KS02
         |--Pseudiron KS02
         `--Pentamerotarsata KS02
              |--Arthroplea KS02
              `--Heptageniidae [Radulapalpata] KS02
                   |  i. s.: Miocenogenia gorbunovi KS02, S02
                   |         Pseudokageronia KS02
                   |         Trichogenia Braasch & Soldán 1988 B06
                   |           `--T. maxillaris Braasch & Soldán 1988 B06
                   |         Cinygmina Kimmins 1937 B06
                   |         Epeorus Eaton 1881 B06
                   |         Rhithrogeniella Ulmer 1939 B06
                   |         Rhithrogena Eaton 1881 B06
                   |           `--R. diaphana Navás 1917 M86
                   |         Dacnogenia Kluge 1988 B06
                   |         Parafronurus Zhou & Braasch 2003 B06
                   |         Electrogena Zurwerra & Tomka 1985 B06
                   |         Ecdyurus TYM08
                   |           |--E. flavus Takahashi 1929 TYM08
                   |           `--E. tobiironis Takahashi 1929 TYM08
                   |         Iron pellucidus Takahashi 1931 TYM08
                   |--Heptagenia Walsh 1863 [Heptageniinae] D70a
                   |    |--H. bachofeni Demoulin 1968 D70a
                   |    |--H. flava Rostock 1877 M86
                   |    |--H. fuscogrisea KS02
                   |    `--H. sulphurea CCG01
                   `--Ecdyonurinae B06
                        |  i. s.: Compsoneuria Eaton 1881 [incl. Compsoneuriella Ulmer 1939] B06
                        |           |--*C. (Compsoneuria) spectabilis Eaton 1881 B06
                        |           `--C. (Siamoneuria Braasch 2006) B06
                        |                `--C. (*S.) kovaci Braasch 2006 B06
                        |--Leucrocutini B06
                        |--Notacanthurus Tshernova 1972 [Notacanthurini] B06
                        |--Atopopini B06
                        |    |--Afronurus Lestage 1924 B06
                        |    |--Asionurus Braasch & Soldán 1986 B06
                        |    |--Atopopus Eaton 1881 B06
                        |    `--Thalerosphyrus Eaton 1881 B06
                        `--Ecdyonurini B06
                             |--Nixe Flowers 1980 B06
                             `--Ecdyonurus Eaton 1868 B06
                                  |--E. affinis (Eaton 1885) M86
                                  |--E. aurantiacus (Burneister 1839) M86
                                  |--E. baeticus Alba-Tercedor & Derka 2004 F05
                                  |--E. dispar (Curtis 1834) M86
                                  |--E. lateralis (Curtis 1834) M86
                                  |--E. olgae Alba-Tercedor & Derka 2004 F05
                                  |--E. starmachi Sowa 1971 M86
                                  |--E. submontanus M86
                                  `--E. torrentis Kimmins 1942 M86

*Type species of generic name indicated

REFERENCES

[B06] Braasch, D. 2006. Compsoneuria (Siamoneuria) kovaci subg. n., sp. n., a new mayfly from northern Thailand (Insecta, Ephemeroptera, Heptageniidae). Senckenbergiana Biologica 86 (1): 47-53.

[CCG01] Canivet, V., P. Chambon & J. Gibert. 2001. Toxicity and bioaccumulation of arsenic and chromium in epigean and hypogean freshwater macroinvertebrates. Archives of Environmental Contamination and Toxicology 40: 345-354.

[D70a] Demoulin, G. 1970a. Troisieme contribution a la connaisance des ephemeropteres d l'ambre Oligocene de la Baltique. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique 46 (2): 1-11.

[D70b] Demoulin, G. 1970b. Contribution a la connaisance de ephemeropteres du Miocene. I. Siphlurites explanatus Cockerell. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique 46 (5): 1-4.

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261-282.

[KS02] Kluge, N. Yu., & N. D. Sinitshenkova. 2002. Order Ephemerida Latreille, 1810. The true mayflies (=Ephemeroptera Hyatt et Arms, 1891 (s. l.); =Euephemeroptera Kluge, 2000. In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 89-97. Kluwer Academic Publishers: Dordrecht.

[M86] Marten, M. 1986. Drei für Deutschland neue und weitere, selten gefundene Eintagsfliegen aus der Fulda (Insecta, Ephemeroptera). Spixiana 9 (2): 169-173.

[PC66] Pendergrast, J. G., & D. R. Cowley. 1966. An Introduction to New Zealand Freshwater Insects. Collins: Auckland.

[S02] Sinitshenkova, N. D. 2002. Ecological history of the aquatic insects. In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 388-426. Kluwer Academic Publishers: Dordrecht.

[TYM08] Tennent, W. J., M. Yasuda & K. Morimoto. 2008. Lansania Journal of arachnology and zoology—a rare and obscure Japanese natural history journal. Archives of Natural History 35 (2): 252-280.

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