Lyroda

Lyroda subita female with prey, copyright S. R. Johnson.

Belongs within: Crabronidae.

Lyroda is a genus of slender, ground-nesting wasps that provision their nests with crickets of the families Gryllidae and Tetrigidae (Bohart & Menke 1976).

Characters (from Bohart & Menke 1976): Inner orbits essentially parallel or converging above; antenna elongate, scape usually short and stout, flagellomeres longer than broad, those of male unmodified; clypeal free margin with several teeth laterally (and sometimes mesally) in female, margin with a median truncate or weakly toothed lobe in male; labrum quadrangular, apex sometimes visible; inner margin of mandible with two subbasal teeth in female, none in male, externoventral margin notched or angulate in both sexes; mouthparts short; occipital carina incomplete, disappearing short of hypostomal carina; collar moderately long, usually trituberculate dorsally; admedian lines of scutum nearly contiguous or widely separated, propodeum long, dorsal enclosure not defined, dorsum usually with a median longitudinal carina, lateral carina sometimes present; mesopleuron with scrobal sulcus; gaster usually sessile, rarely pedunculate, both sexes with a pygidial plate defined by carinae, surface densely or sparsely covered with short setae; legs long, slender, foreleg with a weak tarsal rake in both sexes, spines few; midcoxae separated, hindcoxae contiguous; hindtibia with two parallel rows of spines, the uppermost borne on a weak ridge, arolia large in both sexes; marginal cell of forewing truncate, appendiculate; three submarginal cells, second submarginal cell four- to six-sided (rarely three-sided), not petiolate, receiving both recurrent veins or first recurrent interstitial; length of hindwing jugal lobe equal to or less than half of anal area; male with volsella well differentiated.

<==Lyroda Say 1837 C85 (see below for synonymy)
    |--*L. subita (Say 1837) C85 (see below for synonymy)
    |--L. aethiopica Kohl 1894 BM76
    |--L. argenteofacialis (Cameron 1889) [=Astata argenteofacialis] BM76
    |--L. concinna (Smith 1856) [=Morphota concinna] BM76
    |--L. errans (Turner 1936) [=Gastrosericus errans] C85
    |--L. fasciata (Smith 1856) [=*Morphota fasciata] BM76
    |--L. formosa (Smith 1859) [=Morphota formosa; incl. *Odontolarra rufiventris Cameron 1900] BM76
    |--L. harpactoides (Smith 1856) [=Morphota harpactoides] BM76
    |--L. japonica Iwata 1933 BM76
    |    |--L. j. japonica BM76
    |    `--L. j. takasago Tsuneki 1967 BM76
    |--L. madecassa Arnold 1945 BM76
    |--L. michaelseni Schulz 1908 C85
    |    |--L. m. michaelseni C85
    |    `--L. m. tasmanica Turner 1914 C85
    |--L. minima Turner 1936 C85
    |--L. nigra (Cameron 1904) [=Odontolarra nigra] BM76
    |--L. queenslandensis Turner 1916 C85
    |--L. salai Giner Marí 1945 BM76
    |--L. taiwana Tsuneki 1967 BM76
    |--L. tridens (Taschenberg 1870) [=Morphota tridens] BM76
    |--L. triloba (Say 1837) [=Lyrops trilobus; incl. Lyrops calipterus Say 1837 (n. n.)] BM76
    `--L. venusta Bingham 1897 BM76

Lyroda Say 1837 C85 [=Lyrodon (l. c.) non Goldfuss 1837 BM76; incl. Morphota Smith 1856 BM76, Odontolarra Cameron 1900 BM76]

*Lyroda subita (Say 1837) C85 [=Lyrops (*Lyroda) subita C85; incl. Larrada arcuata Smith 1856 BM76, Lyroda cockerelli Rohwer 1909 BM76]

*Type species of generic name indicated

REFERENCES

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[C85] Cardale, J. C. 1985. Vespoidea and Sphecoidea. In: Walton, D. W. (ed.) Zoological Catalogue of Australia vol. 2. Hymenoptera: Formicoidea, Vespoidea and Sphecoidea pp. 150–303. Australian Government Publishing Service: Canberra.

Nitela

Nitela sp., copyright Christophe Quintin.

Belongs within: Crabronidae.

Nitela is a widespread genus of very small, mostly black crabronid wasps with markedly reduced wing venation.

Characters (from Bohart & Menke 1976): Inner orbits converging above: inner and/or outer orbits sometimes margined by a carina; eyes sometimes with a fine pubescence; antenna variable, pedicel, and flagellomeres usually longer than broad: antennal socket very low on face, surrounded by a basin which may be carinate above; frontal carina usually weak or absent, sometimes bifurcate dorsad, or lamelliform; clypeus usually very narrow, mesally tumid and usually bearing a vertical carina which may be continuous with frontal carina, clypeal margin usually with a median truncate or sinuate lobe; labrum quadrate to subtriangular, its free margin sometimes visible beyond clypeal margin; inner margin of mandible usually with a subapical tooth in female, apex blunt or weakly bifid in some females, inner margin usually toothless in male (exceptionally subapically dentate or with a large subbasal tooth), externoventral margin without a notch; malar space usually present in both sexes: mouthparts short, labial palpus very short, first segment much longer than remaining three; occipital carina incomplete below, usually ending far short of hypostomal carina; mandibular socket open; pronotal collar moderately long and often with a transverse sulcus which is usually interrupted at midline by a carina or triangular prominence, front margin of collar sometimes carinate, humeral angles rounded or angulate; admedian lines of scutum short and widely separated when present; scutellum simple or with a narrow sulcus along anterior margin; propodeum long, dorsum without enclosure but usually longitudinally ridged, coarsely reticulate or areolate, posterior face nearly vertical, flat, and sometimes margined by a carina; propleuron usually simple; mesopleuron with an episternal sulcus and a hypersternaulus; metapleuron sometimes poorly defined beneath upper metapleural pit; pygidial plate absent, segment VI conical in female and sometimes laterally compressed; legs moderately long, foretarsal rake absent; midcoxae separated, occasionally narrowly so, hindcoxae contiguous; hindtibia spineless; tarsomere V not swollen, arolia small; forewing marginal cell truncate apically (sometimes narrowly so) and appendiculate, forewing with only one submarginal and discoidal cell, and subdiscoidal cell open posteriorly because A₁ is evanescent beyond cu-a, recurrent vein usually received by submarginal I, or received beyond in several exceptional species; hindwing venation greatly reduced and jugal lobe small; male sternum VIII variable; volsella at most weakly differentiated from gonostyle.

<==Nitela Latreille 1809 C85
    |  i. s.: N. apoensis Tsuneki et al. 1992 LL10
    |         N. borealis Valkeila 1974 BM76
    |         N. obliquipropodea Li & Li 2010 LL10
    |         N. philippinica Tsuneki et al. 1992 LL10
    |         N. protuberancis Li & Li 2010 LL10
    |--N. (Nitela) [incl. Rhinonitela Williams 1928] BM76
    |    |--*N. (N.) spinolae Latreille 1809 C85 [=N. spinolai Valkeila 1974 BM76]
    |    |--N. (N.) australiensis Schulz 1908 BM76 [incl. N. nigricans Turner 1910 C85]
    |    |--N. (N.) austrocaledonica Williams 1945 BM76
    |    |--N. (N.) bicornis Williams 1928 BM76
    |    |--N. (N.) braunsii Arnold 1940 BM76
    |    |--N. (N.) capicola Brauns 1911 BM76
    |    |--N. (N.) carinifrons Menke 1968 BM76
    |    |--N. (N.) cerasicola Pate 1937 BM76
    |    |--N. (N.) collaris Turner 1926 BM76
    |    |--N. (N.) costaricensis Brauns 1911 BM76
    |    |--N. (N.) darwini Turner 1916 BM76
    |    |--N. (N.) domestica (Williams 1928) [=*Rhinonitela domestica] BM76
    |    |--N. (N.) fallax Kohl 1884 BM76
    |    |--N. (N.) floridana Pate 1934 BM76
    |    |--N. (N.) henrici Turner 1926 BM76
    |    |--N. (N.) kurandae Turner 1908 BM76
    |    |--N. (N.) leoni Krombein 1968 BM76
    |    |--N. (N.) lubutuana Arnold 1929 BM76
    |    |    |--N. l. lubutuana BM76
    |    |    `--N. l. nyasae Arnold 1947 BM76
    |    |--N. (N.) luzonensis Williams 1928 BM76
    |    |--N. (N.) maxima Maidl 1925 BM76
    |    |--N. (N.) merceti Brauns 1911 [incl. N. transvaalensis Brauns 1911] BM76
    |    |--N. (N.) mochii Arnold 1940 BM76
    |    |--N. (N.) ochripes Arnold 1940 BM76
    |    |--N. (N.) ohgushii Tsuneki 1956 BM76
    |    |--N. (N.) oxiana Gussakovskij 1945 BM76
    |    |--N. (N.) parallela Arnold 1929 BM76
    |    |--N. (N.) pendleburyi Turner 1926 BM76
    |    |--N. (N.) promontorii Brauns 1911 BM76
    |    |--N. (N.) reticulata Ducke 1908 BM76
    |    |--N. (N.) retifera Arnold 1940 BM76
    |    |--N. (N.) rhodesiae Arnold 1940 BM76
    |    |--N. (N.) rufiventris Turner 1916 BM76
    |    |--N. (N.) rugosa Williams 1928 BM76
    |    |--N. (N.) rugosissima Arnold 1940 BM76
    |    |--N. (N.) schmidti Brauns 1911 BM76
    |    |--N. (N.) sculpturata Turner 1916 BM76 [=N. reticulata Turner 1908 non Ducke 1908 C85]
    |    |--N. (N.) townesorum Krombein 1950 BM76
    |    |--N. (N.) virginiensis Rohwer 1923 BM76
    |    |--N. (N.) williamsi Menke 1968 BM76
    |    `--N. (N.) yasumatsui Tsuneki 1956 BM76
    |         |--N. y. yasumatsui LL10
    |         `--N. y. taiwana Tsuneki 1956 LL10
    `--N. (Tenila Brèthes 1913) BM76
         |--N. (*T.) amazonica Ducke 1903 BM76
         |--N. (T.) bifida Menke 1969 BM76
         `--N. (T.) gulana (Williams 1928) [=Rhinonitela gulana] BM76

*Type species of generic name indicated

REFERENCES

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[C85] Cardale, J. C. 1985. Vespoidea and Sphecoidea. In: Walton, D. W. (ed.) Zoological Catalogue of Australia vol. 2. Hymenoptera: Formicoidea, Vespoidea and Sphecoidea pp. 150–303. Australian Government Publishing Service: Canberra.

[LL10] Li, T., & Q. Li. 2010. Key to the Oriental species of Nitela Latreille (Hymenoptera: Crabronidae), with descriptions of two new species from China. Oriental Insects 44: 337–344.

Trypoxylini

Pison koreense, from here.

Belongs within: Crabronidae.
Contains: Trypoxylon (Trypoxylon), Trypoxylon (Trypargilum), Pison (Pison).

The Trypoxylini are a group of apoid wasps characterised by notched inner orbits on the compound eyes and often reduced venation in the fore wings (Bohart & Menke 1976).

Characters (from Bohart & Menke 1976, as Trypoxylonini): Inner orbits angularly emarginate (notched), emargination rarely weak; ocelli normal. Antennal sockets low on face, oftne contiguous with frontoclypeal suture; male with elevenb flagellomeres (exceptionally ten). Clypeus variable, usually with median projection or lobe; labrum small. Externoventral margin of mandible entire (occasionally notched), inner margin simple (occasionally a single median tooth or apically bidentate); mouthparts short; (c) mandibular socket open. Pronotal collar high, usually narrow and not closely appressed to scutum; posterolateral margin of propleuron usually lamellate, posterolateral angle usually declivous and set off from rest of propleuron by inner ridge or hump. Scutum with or without notauli. Episternal sulcus present, omaulus and subomaulus rarely present. Female foreleg without a tarsal rake; hind femur simple apically; mid coxae widely separated to contiguous, arolia moderate to large, equal on all legs; male fore coxa and trochanter unmodified. Propodeum short to long; enclosure often undefined; propodeal sternite sometimes present. One, two, or three submarginal cells, cell II petiolate when three are present; two recurrent veins except one in wings with only one submarginal cell, end point of veins variable; marginal cell often acuminate. Jugal lobe small, less than half length of anal area; hamuli usually divided into two groups. Gaster sessile or petiolate; pygidial plate usually not defined. Volsella present; gonostyle simple or bifurcate apically.

<==Trypoxylini [Trypoxyloninae] GOY03
    |--Pisoxylon Menke 1968 BM76
    |    `--*P. xanthosoma Menke 1968 BM76
    |--Aulacophilus Smith 1869 BM76
    |    |--*A. vespoides Smith 1869 BM76
    |    |--A. eumenoides Ducke 1904 BM76
    |    `--A. jansoni Turner 1916 BM76
    |--Trypoxylon Latreille 1796 C85 (see below for synonymy)
    |    |  i. s.: T. appendiculatum Tsuneki 1974 BM76
    |    |         T. deceptorium Antropov 1991 GOY03
    |    |         T. latilobatum Antropov 1991 GOY03
    |    |         T. monstruosum Tsuneki 1974 BM76
    |    |         T. pappi Tsuneki 1974 BM76
    |    |--T. (Trypoxylon) BM76
    |    `--T. (Trypargilum) BM76
    |--Pisonopsis Fox 1893 BM76
    |    |--*P. clypeata Fox 1893 BM76
    |    |    |--P. c. clypeata BM76
    |    |    `--P. c. occidentalis Williams 1954 BM76
    |    |--P. areolata (Spinola 1851) (see below for synonymy) BM76
    |    |--P. australis Fritz 1965 BM76
    |    |--P. birkmanni Rohwer 1909 BM76
    |    `--P. triangularis Ashmead 1899 BM76
    |         |--P. t. triangularis BM76
    |         `--P. t. californica Williams 1954 BM76
    `--Pison Jurine in Spinola 1808 [=Pisum Agassiz 1847 non Megerle 1811] BM76
         |  i. s.: P. antiquum SN18
         |         P. cockerellae Rohwer 1908 BM76
         |         P. oligocenum Cockerell 1908 [=P. oligocaenum Cockerell 1909] BM76
         |--P. (Pison) C85
         |--P. (Entomopison Menke 1968) BM76
         |    |--P. (*E.) pilosum Smith 1873 BM76
         |    |--P. (E.) aureofaciale Strand 1910 BM76
         |    `--P. (E.) convexifrons Taschenberg 1870 BM76
         |--P. (Krombeiniellum Richards 1962) [=Paraceramius Radoszkowski 1887 non Saussure 1854] BM76
         |    |--P. (*K.) koreense (Radoszkowski 1887) [=*Paraceramius koreensis] BM76
         |    |--P. (K.) browni (Ashmead 1905) [=Pisonoides browni] BM76
         |    |--P. (K.) differens Turner 1916 BM76
         |    |--P. (K.) duckei Menke 1968 BM76
         |    |--P. (K.) krombeini Menke 1968 BM76
         |    |--P. (K.) neotropicum Menke 1968 BM76
         |    |--P. (K.) plaumanni Menke 1968 BM76
         |    `--P. (K.) stangei Menke 1968 BM76
         `--P. (Pisonoides Smith 1858) C85 [=Parapison Smith 1869 BM76]
              |--P. (*P.) obliteratum Smith 1858 C85 [=*Parapison obliteratum BM76]
              |--P. (P.) difficile Turner 1908 [=P. (Aulacophilus) difficile] C85
              `--P. (P.) icarioides Turner 1908 [=P. (Aulacophilus) icarioides] C85

Nomina nuda: Trypoxylon annulatum Cameron in Richards 1934 non Brèthes 1913 BM76
             Trypoxylon aureosericeum Schrottky 1913 BM76
             Trypoxylon hispanicum Mercet in Richards 1934 BM76
             Trypoxylon meridionale Gussakovskij 1936 non Richards 1934 BM76
             Trypoxylon simlaense Cameron in Richards 1934 BM76

Pisonopsis areolata (Spinola 1851) [=Pison areolatum; incl. Pisonopsis anomala Mantero 1901, Pison variicornis Reed 1894] BM76

Trypoxylon Latreille 1796 C85 [=Apius Panzer 1806 BM76, Tripoxilon Spinola 1806 BM76, Trypoxilon Jurine 1807 BM76, Trypoxylum Schulz 1906 BM76]

*Type species of generic name indicated

REFERENCES

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[C85] Cardale, J. C. 1985. Vespoidea and Sphecoidea. In: Walton, D. W. (ed.) Zoological Catalogue of Australia vol. 2. Hymenoptera: Formicoidea, Vespoidea and Sphecoidea pp. 150–303. Australian Government Publishing Service: Canberra.

[GOY03] Gayubo, S. F., H. Özbek & E. Yıldırım. 2003. A contribution to the knowledge of Spheciformes of Turkey: tribes Astatini, Dinetini, Larrini, Palarini and Trypoxylini (Hymenoptera: Apoidea: Crabronidae). Zoology in the Middle East 29: 83–92.

[SN18] Sann, M., O. Niehuis, R. S. Peters, C. Mayer, A. Kozlov, L. Podsiadlowski, S. Bank, K. Meusemann, B. Misof, C. Bleidorn & M. Ohl. 2018. Phylogenomic analysis of Apoidea sheds new light on the sister group of bees. BMC Evolutionary Biology 18: 71.

Sericophorus

Sericophorus sp., copyright Jean and Fred Hort.

Belongs within: Crabronidae.

Sericophorus, policemen flies, is an Australian genus of bee-like, ground-nesting wasps that stock their nests with muscoid flies (Bohart & Menke 1976). Historically, the genus has been divided between the subgenera Sericophorus and Zoyphidium based on whether the marginal cell of the fore wing is narrowly truncate or acuminate, respectively, but Lomholdt & Pulawski (2010) did not regard this distinction as significant.

Characters (from Lomholdt & Pulawski 2010): Occipital carina usually reaching hypostomal carina; hind ocelli round, non-modified; propodeal spiracle usually elongate, extending nearly to propodeal foremargin; posterior propodeal surface bearing well defined median carina on approximately ventral third; fore wing usually with three submarginal cells (occasionally two), second submarginal cell usually markedly narrowing anteriorly (except when secondarily fused with third submarginal cell); hind femur simply attenuate; female with setose pygidial plate; male with only ten flagellomeres, volsella absent.

<==Sericophorus Smith 1851 (see below for synonymy) LP10
    |  i. s.: S. centralis Pulawski in Lomholdt & Pulawski 2010 LP10
    |         S. halli Lomholdt in Lomholdt & Pulawski 2010 LP10
    |         S. kurandae Pulawski in Lomholdt & Pulawski 2010 LP10
    |         S. mandibularis Lomholdt in Lomholdt & Pulawski 2010 LP10
    |         S. minutus Rayment 1955 LO10
    |         S. minys Pulawski in Lomholdt & Pulawski 2010 LP10
    |         S. punctum Lomholdt in Lomholdt & Pulawski 2010 LP10
    |--S. asperithorax (Rayment 1955) LP10 [=Anacrucis asperithorax LP10, S. (Zoyphidium) asperithorax C85]
    |--S. aurifrons Lomholdt in Lomholdt & Pulawski 2010 LP10
    |--S. chrysophorus Pulawski in Lomholdt & Pulawski 2010 LP10
    |--S. doddi (Turner 1912) LP10 [=Zoyphium doddi LP10, S. (Zoyphidium) doddi C85]
    |--S. erythrosoma (Turner 1908) LP10 [=Zoyphium erythrosoma LP10, S. (Zoyphidium) erythrosoma C85]
    |--S. laticeps Lomholdt in Lomholdt & Pulawski 2010 LP10
    |--S. ornatus (Hacker & Cockerell 1922) LP10 [=Zoyphium ornatum LP10, S. (Zoyphidium) ornatus C85]
    |--S. papuanus Pulawski in Lomholdt & Pulawski 2010 LP10
    |--S. trichocephalus Pulawski in Lomholdt & Pulawski 2010 LP10
    |--+--S. ambonensis Pulawski in Lomholdt & Pulawski 2010 LP10
    |  `--S. laevigatus (Rayment 1955) LP10 [=*Anacrucis laevigata LP10, S. (Zoyphidium) laevigatus C85]
    |--+--S. crassicornis (Cockerell 1914) LP10 [=Zoyphium crassicorne LP10, S. (Zoyphidium) crassicornis C85]
    |  `--S. pusillus (Hacker & Cockerell 1922) LP10 [=Zoyphium pusillum LP10, S. (Zoyphidium) pusillus C85]
    |--+--S. iridipennis (Turner 1914) LP10 (see below for synonymy)
    |  `--S. vicarius Lomholdt in Lomholdt & Pulawski 2010 LP10
    |--+--S. kohlii (Turner 1908) LP10 [=Zoyphium kohlii LP10, S. (Zoyphidium) kohlii C85]
    |  `--S. riparius Pulawski in Lomholdt & Pulawski 2010 LP10
    |--+--S. froggatti Rayment 1955 LP10 (see below for synonymy)
    |  `--S. funebris Turner 1907 LP10 (see below for synonymy)
    |--+--S. striatulus (Rayment 1955) LP10 [=Anacrucis striatula LP10, S. (Zoyphidium) striatulus C85]
    |  `--S. subalaris Lomholdt in Lomholdt & Pulawski 2010 LP10
    |--+--S. albitarsis Lomholdt in Lomholdt & Pulawski 2010 LP10
    |  |--S. argyreus (Hacker & Cockerell 1922) LP10 [=Zoyphium argyreum LP10, S. (Zoyphidium) argyreus C85]
    |  `--S. tibialis Pulawski in Lomholdt & Pulawski 2010 LP10
    |--+--S. emarginatus (Hacker & Cockerell 1922) LP10 [=Zoyphium emarginatum LP10, S. (Zoyphidium) emarginatus C85]
    |  |--S. lissonotus Lomholdt in Lomholdt & Pulawski 2010 LP10
    |  `--S. niger (Hacker & Cockerell 1922) LP10 [=Zoyphium nigrum LP10, S. (Zoyphidium) niger C85]
    |--+--S. affinis (Hacker & Cockerell 1922) LP10 [=Zoyphium affine LP10, S. (Zoyphidium) affinis C85]
    |  |--S. bicellularis Pulawski in Lomholdt & Pulawski 2010 LP10
    |  |--S. confusus Lomholdt in Lomholdt & Pulawski 2010 LP10
    |  `--S. glossatus Lomholdt in Lomholdt & Pulawski 2010 LP10
    |--+--S. mimus Pulawski in Lomholdt & Pulawski 2010 LP10
    |  |--S. rufoniger (Turner 1908) LP10 (see below for synonymy)
    |  `--+--S. flavorufoniger Pulawski in Lomholdt & Pulawski 2010 LP10
    |     `--S. frontalis (Turner 1908) LP10 [=Zoyphium frontale LP10, S. (Zoyphidium) frontalis C85]
    |--+--S. nigror Rayment 1955 LP10
    |  `--+--S. clypeatus (Rayment 1955) LP10 [=Anacrucis clypeata LP10, S. (Zoyphidium) clypeatus C85]
    |     `--+--S. fissus Lomholdt in Lomholdt & Pulawski 2010 LP10
    |        `--S. nasutus Pulawski in Lomholdt & Pulawski 2010 LP10
    |--+--S. relucens Smith 1856 (see below for synonymy) LP10
    |  `--+--S. collaris (Hacker & Cockerell 1922) LP10 [=Zoyphium collare LP10, S. (Zoyphidium) collaris C85]
    |     |--S. maculatus Pulawski in Lomholdt & Pulawski 2010 LP10
    |     `--+--S. dipteroides Turner 1907 LP10 (see below for synonymy)
    |        `--S. rufithorax Lomholdt in Lomholdt & Pulawski 2010 LP10
    |--+--S. flavipes Pulawski in Lomholdt & Pulawski 2010 LP10
    |  |--S. xanthomelas Pulawski in Lomholdt & Pulawski 2010 LP10
    |  `--+--S. deserticola Pulawski in Lomholdt & Pulawski 2010 LP10
    |     `--+--S. genalis Pulawski in Lomholdt & Pulawski 2010 LP10
    |        `--+--S. karrathensis Pulawski in Lomholdt & Pulawski 2010 LP10
    |           `--+--S. arrernte Pulawski in Lomholdt & Pulawski 2010 LP10
    |              `--S. axillaris Lomholdt in Lomholdt & Pulawski 2010 LP10
    |--+--S. splendidus (Hacker & Cockerell 1922) LP10 (see below for synonymy)
    |  |--+--S. cardaleae Lomholdt in Lomholdt & Pulawski 2010 LP10
    |  |  `--S. multipictus Pulawski in Lomholdt & Pulawski 2010 LP10
    |  `--+--S. cornutus Lomholdt in Lomholdt & Pulawski 2010 LP10
    |     |--S. flavofasciatus (Turner 1916) LP10 [=Zoyphium flavofasciatum LP10, S. (Zoyphidium) flavofasciatum C85]
    |     |--S. nasicornis Lomholdt in Lomholdt & Pulawski 2010 LP10
    |     |--S. sericeus (Kohl 1893) LP10 [=*Zoyphium sericeum LP10, S. (*Zoyphidium) sericeus C85]
    |     |--S. tuberculatus (Turner 1936) LP10 [=Zoyphium tuberculatum C85, S. (Zoyphidium) tuberculatus C85]
    |     `--+--S. rhamphigaster Menke 1989 LP10
    |        `--+--S. pulawskii Ohl 2009 LP10
    |           `--S. rhinoceros Pulawski 1989 LP10
    `--+--S. evansi Lomholdt in Lomholdt & Pulawski 2010 LP10
       |--+--S. nigrescens Lomholdt in Lomholdt & Pulawski 2010 LP10
       |  `--S. protrudens Lomholdt in Lomholdt & Pulawski 2010 LP10
       `--+--S. argentifacies Lomholdt in Lomholdt & Pulawski 2010 LP10
          |--+--S. littoralis Rayment 1955 LP10
          |  `--S. niveifrons Rayment 1955 LP10
          |--+--S. nigripes Lomholdt in Lomholdt & Pulawski 2010 LP10
          |  |--S. wheeleri Lomholdt in Lomholdt & Pulawski 2010 LP10
          |  `--+--*S. chalybeus Smith 1851 (see below for synonymy) LP10
          |     |--S. cliffordi Rayment 1955 (see below for synonymy) LP10
          |     `--S. glaucus Pulawski in Lomholdt & Pulawski 2010 LP10
          `--+--S. viridis (de Saussure 1854) (see below for synonymy) LP10
             `--+--+--+--S. aliceae Turner 1936 LP10
                |  |  `--S. viridescens Lomholdt in Lomholdt & Pulawski 2010 LP10
                |  |--+--S. bicolor Smith 1873 (see below for synonymy) LP10
                |  |  `--S. scintillans Pulawski in Lomholdt & Pulawski 2010 LP10
                |  `--+--S. metallescens Rayment 1955 [incl. S. patongensis Rayment 1955, S. rugosus Rayment 1955] LP10
                |     `--S. sabulosus Lomholdt in Lomholdt & Pulawski 2010 LP10
                `--+--S. setulosus Lomholdt in Lomholdt & Pulawski 2010 LP10
                   |--+--S. houstoni Lomholdt in Lomholdt & Pulawski 2010 LP10
                   |  `--+--S. barakula Pulawski in Lomholdt & Pulawski 2010 LP10
                   |     `--S. rufus Pulawski in Lomholdt & Pulawski 2010 LP10
                   |--+--S. mesopleuralis Pulawski in Lomholdt & Pulawski 2010 LP10
                   |  |--S. ruficollis Lomholdt in Lomholdt & Pulawski 2010 LP10
                   |  `--S. sanguineus Lomholdt in Lomholdt & Pulawski 2010 LP10
                   `--+--S. triangularis Lomholdt in Lomholdt & Pulawski 2010 LP10
                      `--+--S. xanthogaster Lomholdt in Lomholdt & Pulawski 2010 LP10
                         `--+--S. glyptus Pulawski in Lomholdt & Pulawski 2010 LP10
                            `--+--S. sparsus Pulawski in Lomholdt & Pulawski 2010 LP10
                               `--+--S. nitidus Pulawski in Lomholdt & Pulawski 2010 LP10
                                  `--+--S. pectinatus Lomholdt in Lomholdt & Pulawski 2010 LP10
                                     `--S. politus Lomholdt in Lomholdt & Pulawski 2010 LP10

Sericophorus Smith 1851 [incl. Anacrucis Rayment 1955, Tachyrrhostus de Saussure 1854, Zoyphidium Pate 1937, Zoyphium Kohl 1893 non Agassiz 1847] LP10

Sericophorus bicolor Smith 1873 [=Tachyrrhostus bicolor; incl. S. brisbanensis Rayment 1955, S. castaneus Rayment 1955, S. cockerelli Menke in Bohart & Menke 1976, S. hackeri Rayment 1955 non Zoyphium hackeri Cockerell 1932] LP10

*Sericophorus chalybeus Smith 1851 [=S. chalybaeus (l. c.), Tachyrhostus chalybeus; incl. *Tachyrrhostus cyaneus de Saussure 1854, S. inornatus Rayment 1955, S. lilacinus Rayment 1955, S. raymenti Menke in Bohart & Menke 1976, S. rufipes Rayment 1955 non Zoyphium rufipes Rohwer 1911, S. sculpturatus Rayment 1955, S. subviridis Rayment 1955, S. sydneyi Rayment 1955, S. victoriensis Rayment 1955] LP10

Sericophorus cliffordi Rayment 1955 [incl. S. cyanophilus Rayment 1955, S. chalybaeus fulleri Rayment 1955, S. gracilis Rayment 1955, S. rufotibialis Rayment 1955] LP10

Sericophorus dipteroides Turner 1907 LP10 [=S. (Zoyphidium) dipteroides C85, Zoyphium dipteroides LP10; incl. Z. hackeri Cockerell 1932 LP10, S. (Zoyphidium) hackeri C85]

Sericophorus froggatti Rayment 1955 LP10 [incl. S. pescotti Rayment 1955 LP10, Anacrucis punctuosa Rayment 1955 LP10, S. (Zoyphidium) punctuosus C85]

Sericophorus funebris Turner 1907 LP10 [=Zoyphium funebre LP10; incl. Z. humile Cockerell 1932 LP10, S. (Zoyphidium) humilis C85]

Sericophorus iridipennis (Turner 1914) LP10 [=Zoyphium iridipenne LP10, S. (Zoyphidium) iridipennis C85; incl. Zoyphium fuscipenne Hacker & Cockerell 1922 LP10, S. (Z.) fuscipennis C85]

Sericophorus relucens Smith 1856 [incl. S. carinatus Rayment 1955, S. relucens nigricornis Rayment 1955, S. relucens ruficornis Rayment 1955, Zoyphidium rufipes Rohwer 1911, S. rufobasalis Rayment 1955] LP10

Sericophorus rufoniger (Turner 1908) LP10 [=Zoyphium rufonigrum LP10, S. (Zoyphidium) rufoniger C85, S. rufonigrus (l. c.) LP10]

Sericophorus splendidus (Hacker & Cockerell 1922) LP10 [=Zoyphium splendidum LP10, S. (Zoyphidium) splendidus C85; incl. Anacrucis cingulata Rayment 1955 LP10, S. (Z.) cingulatus C85, A. ferruginea Rayment 1955 LP10, S. (Z.) ferrugineus C85]

Sericophorus viridis (de Saussure 1854) [=Tachyrrhostus viridis; incl. S. claviger burnsiellus Rayment 1955, Tachyrhostus (l. c.) claviger Kohl 1892, Sericophorus claviger, S. elegantior Rayment 1955, S. occidentalis Rayment 1955, S. teliferopodus okiellus Rayment 1955, S. viridis roddi Rayment 1955, S. spryi Rayment 1955, S. tallongensis Rayment 1955, S. teliferopodus Rayment 1955, S. violaceus Rayment 1955] LP10

*Type species of generic name indicated

REFERENCES

[C85] Cardale, J. C. 1985. Vespoidea and Sphecoidea. In: Walton, D. W. (ed.) Zoological Catalogue of Australia vol. 2. Hymenoptera: Formicoidea, Vespoidea and Sphecoidea pp. 150–303. Australian Government Publishing Service: Canberra.

[LP10] Lomholdt, O., & W. J. Pulawski. 2010. A revision of the wasp genus Sericophorus F. Smith, 1851 (Hymenoptera: Apoidea: Crabronidae). Proceedings of the California Academy of Sciences, series 4, 61 (1): 1–234.

Last updated: 27 August 2020.

Pison (Pison)

Pison spinolae, from Insects of Tasmania.

Belongs within: Trypoxylini.

The type subgenus of the genus Pison is a cosmopolitan group of wasps nesting in cells constructed in hollows in plant stems or beetle burrows, or in mud cells in sheltered locations. Nests are provisioned with spiders (Bohart & Menke 1976).

Characters (from Bohart & Menke 1976): Eyes bare. Mandible entire. Gaster sessile, not pedunculate, segment I bell-shaped in dorsal view , width of segment I much more than two-thirds greatest width of II and usually subequal to II. Fore wing with two or three submarginal cells.

<==Pison Jurine in Spinola 1808 (Pison) C85 (see below for synonymy)
    |--P. (P.) atrum (Spinola 1808) (see below for synonymy) BM76
    |--P. (P.) aberrans Turner 1908 [=P. (Parapison) aberrans] C85
    |--P. (P.) agile (Smith 1869) [=Parapison agile] BM76
    |--P. (P.) algiricum Kohl 1898 BM76
    |--P. (P.) allonymum Schulz 1906 (see below for synonymy) BM76
    |--P. (P.) areniferum Evans 1981 C85
    |--P. (P.) argentatum Shuckard 1838 (see below for synonymy) BM76
    |--P. (P.) ashmeadi Turner 1916 [incl. P. punctulatum Ashmead 1905 non Kohl 1883] BM76
    |--P. (P.) assimile Sickmann 1894 BM76
    |--P. (P.) atripenne Gussakovskij 1938 BM76
    |--P. (P.) auratum Shuckard 1838 [=P. auratus] C85
    |--P. (P.) aureosericeum Rohwer 1915 C85
    |--P. (P.) aurifex Smith 1869 C85
    |--P. (P.) auriventre Turner 1908 C85
    |--P. (P.) barbatum Evans 1981 C85
    |--P. (P.) basale Smith 1869 [=P. basalis] C85
    |--P. (P.) caliginosum Turner 1908 [=P. (Parapison) caliginosum] C85
    |--P. (P.) cameronii Kohl 1893 [incl. P. fasciatum Kohl 1883 non Pseudonysson fasciatus Radoszkowski 1876] BM76
    |--P. (P.) carinatum Turner 1917 [incl. P. cyprium Gussakovskij 1937] BM76
    |--P. (P.) chilense Spinola 1851 BM76
    |--P. (P.) ciliatum Evans 1981 C85
    |--P. (P.) collare Kohl 1887 BM76
    |--P. (P.) conforme Smith 1869 BM76
    |--P. (P.) congenerum Turner 1916 [=P. congener] C85
    |--P. (P.) cressoni Rohwer 1911 BM76
    |--P. (P.) decipiens Smith 1869 C85
    |--P. (P.) denticeps Cameron 1910 BM76
    |--P. (P.) deperditum Turner 1917 [=P. (Pisonitus) deperditum] C85
    |--P. (P.) dimidiatum Smith 1869 [=Pison dimidiatus] C85
    |--P. (P.) dives Turner 1916 C85
    |--P. (P.) eremnon Menke 1968 BM76
    |--P. (P.) erythrocerum Kohl 1884 [=Parapison ruficornis Smith 1869 non Pison ruficornis Smith 1856] C85
    |--P. (P.) erythrogastrum Rohwer 1915 [=P. (Parapison) erythrogastrum] C85
    |--P. (P.) erythropus Kohl 1884 [incl. Parapison rufipes Smith 1869 non Pison rufipes Shuckard 1838] BM76
    |--P. (P.) esakii Yasumatsu 1937 BM76
    |--P. (P.) exclusum Turner 1916 [=P. (Parapison) exclusum] C85
    |--P. (P.) exornatum Turner 1916 C85
    |--P. (P.) exultans Turner 1916 C85
    |--P. (P.) fasciatum (Radoszkowski 1876) [=*Pseudonysson fasciatus] BM76
    |--P. (P.) fenestratum Smith 1869 [=P. fenestratus, P. nitidus Smith 1868 non Smith 1858] C85
    |--P. (P.) festivum Smith 1869 [=P. festivus] C85
    |--P. (P.) flavolimbatum Turner 1917 BM76
    |--P. (P.) fraterculus Turner 1916 C85
    |--P. (P.) fuscipenne Smith 1869 [=P. fuscipennis] C85
    |--P. (P.) glabrum Kohl 1908 BM76
    |--P. (P.) hanedai Tsuneki 1973 BM76
    |--P. (P.) hissaricum Gussakovskij 1937 BM76
    |--P. (P.) hospes Smith 1879 [incl. P. palauense Yasumatsu 1937 (n. n.)] BM76
    |--P. (P.) humile Arnold 1945 BM76
    |--P. (P.) ignavum Turner 1908 C85
    |--P. (P.) impunctatum Turner 1912 BM76
    |--P. (P.) inaequale Turner 1916 BM76
    |--P. (P.) inconspicuum Turner 1916 C85
    |--P. (P.) infumatum Turner 1908 C85
    |--P. (P.) insigne Sickmann 1894 BM76
    |--P. (P.) insulare Smith 1869 BM76
    |--P. (P.) iridipenne Smith 1879 C85
    |--P. (P.) isolitum Turner 1911 BM76
    |--P. (P.) kohlii Bingham 1897 [incl. P. aureopilosum Cameron 1909] BM76
    |--P. (P.) korrorense Yasumatsu 1937 BM76
    |--P. (P.) laeve Smith 1856 BM76
    |--P. (P.) lobiferum Arnold 1945 BM76
    |--P. (P.) lutescens Turner 1916 C85
    |--P. (P.) maculipenne Smith 1860 BM76
    |--P. (P.) mandibulatum Turner 1916 C85
    |--P. (P.) marginatum Smith 1856 [=P. marginatus] C85
    |--P. (P.) mariannense Yasumatsu 1953 BM76
    |--P. (P.) melanocephalum Turner 1908 C85
    |--P. (P.) meridionale Turner 1916 C85
    |--P. (P.) minicum Arnold 1945 BM76
    |--P. (P.) montanum Cameron 1910 BM76
    |--P. (P.) morosum Smith 1856 BM76
    |--P. (P.) multistrigatum Turner 1917 BM76
    |--P. (P.) nigellum Krombein 1949 BM76
    |--P. (P.) nitidum Smith 1858 BM76
    |--P. (P.) noctulum Turner 1908 [=P. (Parapison) noctulum] C85
    |--P. (P.) novocaledonicum Krombein 1949 BM76
    |--P. (P.) oakleyi Krombein 1949 BM76
    |    |--P. o. oakleyi BM76
    |    |--P. o. boninense Tsuneki 1973 BM76
    |    `--P. o. rotaense Tsuneki 1968 BM76
    |--P. (P.) obesum Arnold 1958 BM76
    |--P. (P.) orientale Cameron 1897 BM76
    |--P. (P.) pallidipalpe Smith 1863 BM76
    |--P. (P.) papuanum Schulz 1904 [incl. P. constrictum Turner 1912, P. morosum Smith 1864 non Smith 1856] BM76
    |--P. (P.) pasteelsi Leclercq 1965 BM76
    |--P. (P.) peletieri Le Guillou 1841 C85 [=P. pelletieri Le Guillou 1841 BM76]
    |--P. (P.) perplexum Smith 1856 [=P. perplexus] C85
    |--P. (P.) pertinax Turner 1908 [=P. (Parapison) pertinax] C85
    |--P. (P.) petularum Leclercq 1965 BM76
    |--P. (P.) ponape Krombein 1949 BM76
    |--P. (P.) pregustum Leclercq 1965 BM76
    |--P. (P.) premunitum Leclercq 1965 BM76
    |--P. (P.) priscum Turner 1908 [=P. insulare priscum] C85
    |--P. (P.) pulchrinum Turner 1916 C85
    |--P. (P.) punctifrons Shuckard 1838 (see below for synonymy) BM76
    |--P. (P.) punctulatum Kohl 1884 C85
    |--P. (P.) regale Smith 1852 BM76
    |--P. (P.) repentinum Arnold 1940 BM76
    |--P. (P.) rothneyi Cameron 1896 [incl. P. crassicorne Cameron 1897] BM76
    |--P. (P.) ruficorne Smith 1856 [=P. (Pisonitus) ruficornis] C85
    |--P. (P.) rufipes Shuckard 1838 [=P. (Pisonitus) rufipes] C85
    |--P. (P.) rufitarse Arnold 1944 BM76
    |--P. (P.) rugosum Smith 1856 [incl. P. appendiculatum Cameron 1897] BM76
    |--P. (P.) sarawakense Cameron 1903 BM76
    |--P. (P.) scabrum Turner 1908 C85
    |--P. (P.) scruposum Arnold 1955 BM76
    |--P. (P.) separatum Smith 1869 [=P. separatus] C85
    |--P. (P.) sericeum Kohl 1888 BM76
    |--P. (P.) seyrigi Arnold 1945 BM76
    |--P. (P.) simillimum Smith 1869 [=P. simillimus] C85
    |--P. (P.) simulans Turner 1915 [=P. (Parapison) simulans] C85
    |--P. (P.) sogdianum Gussakovskij 1937 BM76
    |--P. (P.) speculare Turner 1911 BM76
    |--P. (P.) spinolae Shuckard 1838 C85 (see below for synonymy)
    |--P. (P.) strandi Yasumatsu 1935 [incl. P. iwatai Yasumatsu 1935] BM76
    |--P. (P.) strenuum Turner 1916 C85
    |--P. (P.) strictifrons Vachal 1907 BM76
    |--P. (P.) strigulosum Turner 1917 BM76
    |--P. (P.) susanae Cheesman 1955 BM76
    |--P. (P.) suspicax Kokujev 1912 BM76
    |--P. (P.) tahitense Saussure 1867 [incl. P. rechingeri Kohl 1908] BM76
    |--P. (P.) tenebrosum Turner 1908 [=P. (Parapison) tenebrosum] C85
    |--P. (P.) testaceipes Turner 1916 BM76
    |--P. (P.) tibiale Smith 1869 [=Pison tibialis] C85
    |--P. (P.) tosawai Yasumatsu 1935 BM76
    |--P. (P.) transvaalense Cameron 1910 [incl. P. clypeatum Cameron 1910] BM76
    |--P. (P.) trukense Yasumatsu 1953 BM76
    |--P. (P.) tuberculatum Smith 1869 BM76
    |--P. (P.) ugandense Arnold 1955 BM76
    |--P. (P.) ussuriense Gussakovskij 1937 BM76
    |--P. (P.) vestitum Smith 1856 [=P. vestitus] C85
    |--P. (P.) virosum Turner 1908 C85
    |--P. (P.) wagneri Arnold 1932 BM76
    |--P. (P.) westwoodii Shuckard 1838 [incl. P. obliquus Smith 1856] C85
    |--P. (P.) wollastoni Turner 1916 BM76
    `--P. (P.) xanthopus (Brullé 1833) [=*Nephridia xanthopus; incl. P. obscurum Shuckard 1838] BM76

Pison Jurine in Spinola 1808 (Pison) C85 [incl. Nephridia Brullé 1833 BM76, Pisonitus Shuckard 1838 BM76, Pseudonysson Radoszkowski 1876 BM76, Tachybulus Latreille 1809 BM76, Taranga Kirby 1883 BM76]

Pison (Pison) allonymum Schulz 1906 [incl. P. iridipenne Cameron 1905 non Smith 1879, P. karrooense Arnold 1924, P. rhodesianum Bischoff 1913] BM76

Pison (Pison) argentatum Shuckard 1838 [=*Pisonitus argentatus; incl. Pisonitus argenteus Ashmead 1904, Pison fuscipalpe Cameron 1901] BM76

Pison (Pison) atrum (Spinola 1808) [=Alyson ater; incl. *Pison jurinei Spinola 1808, P. jurini, *Tachybulus niger Latreille 1809] BM76

Pison (Pison) punctifrons Shuckard 1838 [incl. P. fabricator Smith 1869, P. japonicum Gussakovskij 1937, P. javanum Cameron 1905, P. lagunae Ashmead 1904, P. striolatum Cameron 1896, P. suspiciosum Smith 1858] BM76

Pison (Pison) spinolae Shuckard 1838 C85 [incl. P. australis Saussure 1853 C85, *Taranga dubia Kirby 1883 BM76, P. pruinosum Cameron 1898 BM76, P. tasmanicus Smith 1856 C85]

*Type species of generic name indicated

REFERENCES

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[C85] Cardale, J. C. 1985. Vespoidea and Sphecoidea. In: Walton, D. W. (ed.) Zoological Catalogue of Australia vol. 2. Hymenoptera: Formicoidea, Vespoidea and Sphecoidea pp. 150–303. Australian Government Publishing Service: Canberra.

Trypoxylon (Trypoxylon)

Trypoxylon cf. scutatum, copyright Hectonichus.

Belongs within: Trypoxylini.

The type subgenus of the genus Trypoxylon is a cosmopolitan group of slender-bodied wasps, species of which seem to take a variety of prey species (Bohart & Menke 1976).

Characters (from Bohart & Menke 1976): Transverse (may be angulate) interantennal carina present, or if not, then frons just above or between sockets usually strongly or abruptly elevated; frontal carina weak or strong, often bifurcating above, the arms sometimes rejoining to enclose a shield-shaped area on frons; dorsal margin of petiole socket convex, narrowly bandlike or merely sharp edged, often with a backward projecting lamella; omaulus absent: propodeal side often limited dorsally by a carina or foveate sulcus between spiracle and petiole socket; propodeal sternite often present; two groups of hind wing hamuli usually separated by more than length of apical row. Male genitalia with gonostyle usually divided apically into dorsal and ventral lobe ("squama"); sternum VIII variable (apical processes short if present).

<==Trypoxylon Latreille 1796 (Trypoxylon) C85 [incl. Asaconoton Arnold 1959 BM76]
    |--*T. (T.) figulus (Linnaeus 1758) C85 (see below for synonymy)
    |    |--T. f. figulus [incl. T. f. major Kohl 1883, T. f. minor Beaumont 1945] BM76
    |    |--T. f. barbarum Beaumont 1957 [incl. T. fieuzeti Giner Marí 1959] BM76
    |    |--T. f. koma Tsuneki 1956 BM76
    |    |--T. f. medium Beaumont 1945 [incl. T. rubi Wolf 1959] BM76
    |    `--T. f. yezo Tsuneki 1956 BM76
    |--T. (T.) abditum Arnold 1924 BM76
    |--T. (T.) accumulator Smith 1875 BM76
    |--T. (T.) acutangulum Arnold 1951 BM76
    |--T. (T.) aegyptium Kohl 1906 [=T. aegyptiacum (l. c.)] BM76
    |--T. (T.) aeneipenne Arnold 1943 BM76
    |--T. (T.) albipes Smith 1856 BM76
    |--T. (T.) albitarsatum Tsuneki 1977 C85
    |    |--T. a. albitarsatum C85
    |    `--T. a. huonense Tsuneki 1977 C85
    |--T. (T.) aldrichi Sandhouse 1940 BM76
    |--T. (T.) algoense Arnold 1924 BM76
    |--T. (T.) ambiguum Tsuneki 1956 BM76
    |--T. (T.) ammophiloides Costa 1871 BM76
    |--T. (T.) arabicum Gussakovskij 1936 BM76
    |--T. (T.) arnoldi Menke in Bohart & Menke 1976 [=T. cognatum Arnold 1924 non Cameron 1897] BM76
    |--T. (T.) arroyense Richards 1934 BM76
    |--T. (T.) arudum Leclercq 1965 BM76
    |--T. (T.) ashmeadi Baltazar 1966 [incl. T. elongatum Ashmead 1905 non Smith 1856] BM76
    |--T. (T.) asinum Leclercq 1965 BM76
    |--T. (T.) asuncicola Strand 1910 BM76
    |    |--T. a. asuncicola BM76
    |    `--T. a. levius Richards 1934 BM76
    |--T. (T.) attenuatum Smith 1851 BM76
    |    |--T. a. attenuatum BM76
    |    `--T. a. hannibalis Gribodo 1896 BM76
    |--T. (T.) backi Sandhouse 1940 BM76
    |--T. (T.) balteatum Cameron 1889 BM76
    |--T. (T.) barberi Richards 1934 BM76
    |--T. (T.) bellardi Richards 1934 BM76
    |--T. (T.) biarti Richards 1934 BM76
    |--T. (T.) bicolor Smith 1856 BM76
    |--T. (T.) bidentatum Fox 1891 [incl. T. morrisoni Richards 1934] BM76
    |--T. (T.) bilobatum Tsuneki 1961 BM76
    |--T. (T.) bituberculatum Tsuneki 1977 C85
    |--T. (T.) bourgeoisi Strand 1911 BM76
    |--T. (T.) brasilianum Saussure 1867 BM76
    |--T. (T.) braunsi Arnold 1924 BM76
    |--T. (T.) brevipenne Saussure 1867 BM76
    |--T. (T.) breviventre Arnold 1959 BM76
    |--T. (T.) bridwelli Sandhouse 1940 BM76
    |--T. (T.) buddha Cameron 1889 BM76
    |--T. (T.) caldesianum Richards 1934 BM76
    |--T. (T.) cameronii Dalla Torre 1897 (see below for synonymy) BM76
    |--T. (T.) canaliculatum Cameron 1889 BM76
    |--T. (T.) capense Cameron 1905 BM76
    |--T. (T.) capitale Richards 1934 BM76
    |--T. (T.) carcinum Leclercq 1965 BM76
    |--T. (T.) carinatum Say 1837 BM76
    |--T. (T.) cariosum Arnold 1959 BM76
    |--T. (T.) carpenteri Richards 1933 BM76
    |--T. (T.) castoris Leclercq 1965 BM76
    |--T. (T.) cataractae Arnold 1924 BM76
    |    |--T. c. cataractae BM76
    |    `--T. c. madecassum Arnold 1945 BM76
    |--T. (T.) catinum Leclercq 1965 BM76
    |--T. (T.) cavallum Leclercq 1965 BM76
    |--T. (T.) centrale Fox 1895 BM76
    |--T. (T.) chichidzimaense Tsuneki 1973 BM76
    |--T. (T.) chichimecum Saussure 1867 BM76
    |--T. (T.) chihpense Tsuneki 1971 BM76
    |--T. (T.) chinense Gussakovskij 1936 BM76
    |--T. (T.) chingi Tsuneki 1971 BM76
    |--T. (T.) chirindense Arnold 1936 BM76
    |--T. (T.) chosenense Tsuneki 1956 BM76
    |--T. (T.) cinctellum Richards 1934 BM76
    |--T. (T.) clarkei Krombein 1962 BM76
    |--T. (T.) clavicerum Lepeletier & Serville 1828 [incl. T. tibiale Zetterstedt 1840] BM76
    |    |--T. c. clavicerum BM76
    |    |--T. c. exiguum Tsuneki 1956 BM76
    |    `--T. c. gussakovskiji Tsuneki 1974 [incl. T. pygmaeum Gussakovskij 1933 non Cameron 1900] BM76
    |--T. (T.) cocorite Richards 1934 BM76
    |--T. (T.) cognatum Cameron 1897 BM76
    |--T. (T.) coloratum Smith 1857 BM76
    |--T. (T.) confrater Kohl 1894 BM76
    |--T. (T.) cornigerum Cameron 1889 BM76
    |--T. (T.) crassifrons Tsuneki 1963 BM76
    |--T. (T.) crassipunctatum Arnold 1946 BM76
    |--T. (T.) cricetum Leclercq 1965 BM76
    |--T. (T.) crudele Richards 1934 BM76
    |--T. (T.) darium Leclercq 1965 BM76
    |--T. (T.) dendrophilum Arnold 1959 BM76
    |--T. (T.) deuterium Leclercq 1965 BM76
    |--T. (T.) dubiosum Tsuneki 1964 BM76
    |--T. (T.) duckei Richards 1934 BM76
    |--T. (T.) dyeri Richards 1934 BM76
    |--T. (T.) ebneri Maidl 1924 BM76
    |--T. (T.) ecuadorium Richards 1934 BM76
    |    |--T. e. ecuadorium BM76
    |    `--T. e. meridionale Richards 1934 BM76
    |--T. (T.) egregium Arnold 1959 [=*Asaconoton egregium] BM76
    |--T. (T.) elegantulum Smith 1860 BM76
    |--T. (T.) elgonense Arnold 1956 BM76
    |--T. (T.) elongatum Smith 1856 BM76
    |--T. (T.) emeritum Leclercq 1965 BM76
    |--T. (T.) errans Saussure 1867 [incl. T. gardneri Cameron 1907] BM76
    |--T. (T.) erythrozonatum Cameorn 1902 BM76
    |--T. (T.) eugeniae Gussakovskij 1936 BM76
    |--T. (T.) excellens Strand 1910 BM76
    |--T. (T.) eximium Smith 1859 C85
    |--T. (T.) eyeni Leclercq 1965 BM76
    |--T. (T.) fabricator Smith 1873 [incl. T. brevicarinatum Cameron 1912, T. gracile Taschenberg 1875] BM76
    |--T. (T.) fastigium Fox 1893 (see below for synonymy) BM76
    |--T. (T.) fenchihuense Tsuneki 1967 BM76
    |--T. (T.) ferox Smith 1860 BM76
    |--T. (T.) fiebrigi Richards 1934 BM76
    |--T. (T.) fitzgeraldi Richards 1934 BM76
    |--T. (T.) flavimanum Arnold 1946 BM76
    |--T. (T.) flavipes Tsuneki 1979 C85
    |--T. (T.) fletcheri Turner 1918 BM76
    |--T. (T.) florale Richards 1934 BM76
    |--T. (T.) formosicola Strand 1922 BM76
    |    |--T. f. formosicola BM76
    |    |--T. f. calcarale Strand 1922 BM76
    |    |--T. f. inornatum Matsumura 1926 [incl. T. amamiense Tsuneki 1964] BM76
    |    `--T. f. kankauense Strand 1922 BM76
    |--T. (T.) foveatum Cameron 1904 BM76
    |--T. (T.) frigidum Smith 1856 [incl. T. plesium Rohwer 1920] BM76
    |    |--T. f. frigidum BM76
    |    |--T. f. chongar Tsuneki 1956 BM76
    |    `--T. f. cornutum Gussakovskij 1933 BM76
    |--T. (T.) frioense Richards 1934 BM76
    |--T. (T.) frontale Smith 1856 BM76
    |--T. (T.) fornticorne Gussakovskij 1936 BM76
    |--T. (T.) fulvocollare Cameron 1904 BM76
    |--T. (T.) funatui Tsuneki 1963 BM76
    |--T. (T.) geniculatum Cameron 1902 BM76
    |--T. (T.) gounellei Richards 1934 BM76
    |--T. (T.) gracilescens Smith 1860 [incl. T. petioloides Strand 1922] BM76
    |--T. (T.) gracilicorne Arnold 1946 BM76
    |--T. (T.) gracillimum Smith 1863 BM76
    |--T. (T.) grenadense Richards 1934 BM76
    |--T. (T.) gussachaos Menke in Bohart & Menke 1976 (see below for synonymy) BM76
    |--T. (T.) gustatum Leclercq 1965 BM76
    |--T. (T.) hova Saussure 1892 BM76
    |--T. (T.) imayoshii Yasumatsu 1938 BM76
    |--T. (T.) inconstans Arnold 1946 BM76
    |--T. (T.) indicum Menke in Bohart & Menke 1976 [=T. ornatipes Cameron 1913 non Fox 1891] BM76
    |--T. (T.) infimum Arnold 1940 BM76
    |--T. (T.) insolitum Fox 1897 BM76
    |--T. (T.) intrudens Smith 1870 BM76
    |--T. (T.) isigakiense Tsuneki 1973 BM76
    |--T. (T.) javanum Taschenberg 1875 BM76
    |--T. (T.) johnsoni Fox 1891 [incl. T. adelphiae Sandhouse 1940, T. ornatipes Fox 1891] BM76
    |--T. (T.) judicum Leclercq 1965 BM76
    |--T. (T.) kabeyae Leclercq 1965 BM76
    |--T. (T.) kalimantan Menke in Bohart & Menke 1976 [=T. annulipes Cameron 1903 non Taschenberg 1875] BM76
    |--T. (T.) kansitakum Tsuneki 1971 BM76
    |--T. (T.) kapiricum Leclercq 1965 BM76
    |--T. (T.) kaszabi Tsuneki 1971 BM76
    |--T. (T.) katangae Leclercq 1965 BM76
    |--T. (T.) khasiae Cameron 1904 BM76
    |--T. (T.) klapperichi Balthasar 1957 BM76
    |--T. (T.) kodamanum Tsuneki 1972 BM76
    |--T. (T.) kohli Arnold 1924 BM76
    |--T. (T.) koikense Tsuneki 1956 BM76
    |--T. (T.) kolazyi Kohl 1893 BM76
    |--T. (T.) kolthoffi Gussakovskij 1938 BM76
    |--T. (T.) konosuense Tsuneki 1968 BM76
    |--T. (T.) koreanum Tsuneki 1956 BM76
    |--T. (T.) koshunicon Strand 1922 BM76
    |--T. (T.) kumaso Tsuneki 1966 BM76
    |--T. (T.) kyotoense Tsuneki 1966 BM76
    |--T. (T.) lacustre Arnold 1956 [incl. T. insulum Arnold 1959] BM76
    |--T. (T.) latiscutatum Arnold 1946 BM76
    |    |--T. l. latiscutatum BM76
    |    `--T. l. rectirugosum Arnold 1959 BM76
    |--T. (T.) layouanum Evans 1972 BM76
    |--T. (T.) leptogaster Kohl 1894 BM76
    |--T. (T.) letiferum Arnold 1946 BM76
    |--T. (T.) leucarthrum Richards 1934 BM76
    |--T. (T.) lissonotum Cameron 1910 BM76
    |--T. (T.) lucidum Arnold 1959 BM76
    |--T. (T.) lusingum Leclercq 1965 BM76
    |--T. (T.) luteosignatum Arnold 1945 BM76
    |--T. (T.) mabwense Leclercq 1965 BM76
    |--T. (T.) magrettii Gribodo 1884 BM76
    |--T. (T.) maidli Richards 1934 BM76
    |    |--T. m. maidli BM76
    |    `--T. m. bodkini Richards 1934 BM76
    |--T. (T.) malaisei Gussakovskij 1933 BM76
    |--T. (T.) mandibulatum Richards 1933 BM76
    |--T. (T.) manni Richards 1934 BM76
    |--T. (T.) marginatum Cameron 1912 BM76
    |--T. (T.) marginifrons Cameron 1912 BM76
    |--T. (T.) massaicum Cameron 1910 BM76
    |--T. (T.) mazaruni Richards 1934 BM76
    |--T. (T.) mediator Nurse 1903 BM76
    |--T. (T.) melanocorne Strand 1922 BM76
    |--T. (T.) melanurum Cameron 1902 BM76
    |--T. (T.) mindanaonis Tsuneki 1976 C85
    |--T. (T.) montanum Schulz 1906 [incl. T. placidum Cameron 1904 non Smith 1863] BM76
    |--T. (T.) monticola Tsuneki 1956 BM76
    |--T. (T.) montivagum Arnold 1940 BM76
    |--T. (T.) moraballi Richards 1934 BM76
    |    |--T. m. moraballi BM76
    |    `--T. m. boliviense Richards 1934 BM76
    |--T. (T.) murotai Tsuneki 1973 BM76
    |--T. (T.) mutilatum Richards 1934 BM76
    |--T. (T.) nagamasae Tsuneki 1963 BM76
    |--T. (T.) nambui Tsuneki 1966 [incl. T. kinkadzanense Tsuneki 1971] BM76
    |--T. (T.) nigrispine Cameron 1905 BM76
    |--T. (T.) nipponicum Tsuneki 1956 BM76
    |--T. (T.) nitidissimum Richards 1934 BM76
    |--T. (T.) nodosicorne Turner 1917 BM76
    |--T. (T.) nodosum Arnold 1944 BM76
    |--T. (T.) obsonator Smith 1873 [incl. T. hyporientale Strand 1922] BM76
    |    |--T. o. obsonator BM76
    |    `--T. o. tropicale Tsuneki 1961 BM76
    |--T. (T.) oculare Menke 1968 BM76
    |--T. (T.) okinawanum Tsuneki 1966 BM76
    |--T. (T.) ordinarium Richards 1934 BM76
    |--T. (T.) orientale Cameron 1904 BM76
    |--T. (T.) orientinum Richards 1969 BM76
    |--T. (T.) pachygaster Richards 1934 BM76
    |--T. (T.) pacificum Gussakovskij 1933 BM76
    |--T. (T.) pan Arnold 1956 BM76
    |--T. (T.) papuanum Tsuneki 1977 C85
    |--T. (T.) parvum Schrottky 1902 BM76
    |--T. (T.) patruele Arnold 1924 BM76
    |--T. (T.) paulisum Leclercq 1965 BM76
    |--T. (T.) peltopse Kohl 1906 BM76
    |--T. (T.) pennsylvanicum Saussure 1867 BM76
    |    |--T. p. pennsylvanicum BM76
    |    `--T. p. japonense Tsuneki 1956 BM76
    |--T. (T.) pentheri Richards 1934 BM76
    |--T. (T.) petiolatum Smith 1857 BM76
    |--T. (T.) philippinense Ashmead 1904 BM76
    |--T. (T.) pileatum Smith 1856 [incl. T. nigricans Cameron 1889 (n. n.)] BM76
    |--T. (T.) placidum Smith 1863 BM76
    |--T. (T.) posterorubrum Richards 1934 BM76
    |--T. (T.) providum Smith 1860 BM76
    |--T. (T.) pulawskii Tsuneki 1956 BM76
    |--T. (T.) puliense Tsuneki 1967 BM76
    |--T. (T.) pumilio Arnold 1959 BM76
    |--T. (T.) punctatissimum Arnold 1924 BM76
    |--T. (T.) punctivertex Richards 1934 BM76
    |--T. (T.) puttalamum Strand 1922 BM76
    |--T. (T.) pygmaeum Cameron 1900 BM76
    |--T. (T.) quadriceps Tsuneki 1971 BM76
    |--T. (T.) regulare Viereck 1906 BM76
    |--T. (T.) rejector Smith 1870 BM76
    |--T. (T.) responsum Nurse 1903 BM76
    |    |--T. r. responsum BM76
    |    |--T. r. hatogayuum Tsuneki 1956 BM76
    |    |--T. r. regium Gussakovskij 1933 BM76
    |    |--T. r. ryukyuense Tsuneki 1966 BM76
    |    `--T. r. taiwanum Tsuneki 1967 BM76
    |--T. (T.) richardsi Sandhouse 1940 BM76
    |--T. (T.) rubiginosum Gussakovskij 1936 BM76
    |--T. (T.) rubrifemoratum Richards 1934 BM76
    |    |--T. r. rubrifemoratum BM76
    |    `--T. r. rubellum Richards 1934 BM76
    |--T. (T.) rufidens Cameron 1905 BM76
    |    |--T. r. rufidens BM76
    |    `--T. r. trinidadianum Richards 1934 BM76
    |--T. (T.) rufimanum Spinola 1851 BM76
    |--T. (T.) rugiceps Dalla Torre 1897 [incl. T. rugifrons Cameorn 1889 non Smith 1873] BM76
    |--T. (T.) rugifrons Smith 1873 BM76
    |--T. (T.) saltamaense Tsuneki 1973 BM76
    |--T. (T.) sanctum Richards 1934 BM76
    |--T. (T.) sapporoense Tsuneki 1960 BM76
    |--T. (T.) schmidti Richards 1936 BM76
    |--T. (T.) schmiedeknechti Kohl 1906 C85
    |    |--T. s. schmiedeknechti C85
    |    `--T. s. connexum Turner 1908 C85
    |--T. (T.) sculleni Sandhouse 1940 BM76
    |--T. (T.) scutatum Chevrier 1867 [incl. T. quartinae Gribodo 1884, T. scutigerum Taschenberg 1881] BM76
    |--T. (T.) scutiferum Taschenberg 1875 BM76
    |--T. (T.) scutifrons Saussure 1892 BM76
    |--T. (T.) segregatum Richards 1934 BM76
    |--T. (T.) senegambicum Kohl 1906 [incl. T. palaeotropicum Schulz 1906] BM76
    |--T. (T.) seyrigi Arnold 1945 BM76
    |--T. (T.) shannoni Richards 1934 BM76
    |--T. (T.) shimoyamai Tsuneki 1958 BM76
    |--T. (T.) shirozui Tsuneki 1966 BM76
    |--T. (T.) silvestre Richards 1934 BM76
    |--T. (T.) sinuosiscutis Arnold 1945 BM76
    |--T. (T.) sogdianum Gussakovskij 1952 BM76
    |--T. (T.) solivagum Arnold 1946 BM76
    |--T. (T.) staudingeri Richards 1934 BM76
    |--T. (T.) stevensoni Arnold 1924 BM76
    |--T. (T.) stieglmayri Richards 1934 BM76
    |--T. (T.) stroudi Gribodo 1884 BM76
    |--T. (T.) subpileatum Strand 1922 BM76
    |--T. (T.) succinctum Cresson 1865 BM76
    |--T. (T.) sulcatoides Richards 1934 BM76
    |--T. (T.) sulcifrons Gussakovskij 1936 BM76
    |--T. (T.) syriacum Mercet 1906 BM76
    |--T. (T.) tainanense Strand 1922 [=T. taiwanense (l. c.)] BM76
    |--T. (T.) takasago Tsuneki 1966 BM76
    |--T. (T.) tanoi Tsuneki 1967 BM76
    |--T. (T.) testaceicorne Cameron 1907 BM76
    |--T. (T.) testaceipes Arnold 1924 BM76
    |--T. (T.) thaianum Tsuneki 1961 BM76
    |--T. (T.) timberlakei Sandhouse 1940 BM76
    |--T. (T.) tinctipenne Cameron 1889 BM76
    |--T. (T.) toltecum Saussure 1867 BM76
    |--T. (T.) tremulum Arnold 1946 BM76
    |--T. (T.) tricolor Sickmann 1894 BM76
    |--T. (T.) trigeminum Richards 1934 BM76
    |--T. (T.) trinidadense Richards 1934 BM76
    |--T. (T.) triste Arnold 1924 BM76
    |--T. (T.) trochanteratum Cameron 1902 BM76
    |--T. (T.) tuberculifrons Arnold 1945 BM76
    |--T. (T.) tubulentum Arnold 1946 BM76
    |--T. (T.) turkestanicum Gussakovskij 1936 BM76
    |--T. (T.) unguicorne Richards 1934 BM76
    |--T. (T.) urichi Richards 1934 BM76
    |--T. (T.) vallicola Tsuneki 1971 BM76
    |--T. (T.) varipes Pérez 1905 BM76
    |    |--T. v. varipes BM76
    |    `--T. v. nasutum Tsuneki 1974 BM76
    |--T. (T.) varipilosum Cameron 1901 BM76
    |--T. (T.) viduum Arnold 1951 BM76
    |--T. (T.) volitans Arnold 1956 BM76
    |--T. (T.) wheeleri Richards 1936 BM76
    |--T. (T.) winthemi Richards 1934 BM76
    |--T. (T.) xenophon Richards 1934 BM76
    |--T. (T.) zikae Arnold 1956 BM76
    `--T. (T.) zurki Leclercq 1965 BM76

Trypoxylon (Trypoxylon) cameronii Dalla Torre 1897 [incl. T. palliditarse Cameron 1889 non Saussure 1867, T. tabascense Strand 1910] BM76

Trypoxylon (Trypoxylon) fastigium Fox 1893 [incl. T. carinifrons Fox 1891 non Cameron 1889, T. nigrellum Rohwer 1909, T. subfrigidum Rohwer 1909] BM76

*Trypoxylon (Trypoxylon) figulus (Linnaeus 1758) C85 [=Sphex figulus C85, *Apius figulus BM76; incl. Trypoxylon apicale Fox 1891 BM76, Sphex fuliginosus Scopoli 1763 BM76]

Trypoxylon (Trypoxylon) gussachaos Menke in Bohart & Menke 1976 [=T. carinifrons Gussakovskij 1936 nec Cameron 1889 nec Fox 1891] BM76

*Type species of generic name indicated

REFERENCES

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[C85] Cardale, J. C. 1985. Vespoidea and Sphecoidea. In: Walton, D. W. (ed.) Zoological Catalogue of Australia vol. 2. Hymenoptera: Formicoidea, Vespoidea and Sphecoidea pp. 150–303. Australian Government Publishing Service: Canberra.