Male Calotarsa pallipes, copyright William Donald Newton.

Belongs within: Cyclorrhapha.
Contains: Phoridae.

The Platypezoidea are a clade of small, humpbacked flies supported by molecular phylogenetic analyses. Members of the Platypezidae, flat-footed flies, often have the hind tarsi modified, such as being enlarged and flattened in females or bearing processes in males (Chandler 2010). The Ironomyiidae are small flies restricted in the modern fauna to Australia though fossil representatives are more widespread in the Cretaceous (Grimaldi & Engel 2005).

<==Platypezoidea [Clythioidea, Phoroidea] WT11
    |  i. s.: Opetiala shatalkani GE05
    |--Opetiidae [Opetiinae] WT11
    |    |--Lithopetia hirsuta GE05
    |    `--Opetia nigra WT11
    `--+--Phoridea GC99
       |    |--Phoridae WT11
       |    `--Ironomyiidae WT11
       |         |--Palaeopetia laiyangensis Zhang 1987 GE05, RJ93
       |         |--Hermaeomyia GE05
       |         |--Eridomyia GE05
       |         |--Cretonomyia pristina GE05
       |         |--Sinolesta [Sinolestinae] GE05
       |         `--Ironomyia [Ironomyiinae] WT11
       |              `--I. nigromaculata WT11
       `--Platypezidae [Clythiidae] WT11
            |  i. s.: Maritulus sospes GC99
            |         Parnasos firmipes GC99
            |         Proplatypeza GE05
            |           |--P. amabilis GC99
            |           `--P. parva GC99
            |         Lebambromyia Grimaldi & Cumming 1999 GE05, GC99
            |           `--*L. acrai Grimaldi & Cumming 1999 GC99
            |         Promittor GE05
            |         Oloplatypeza GE05
            |         Bolopus C10
            |         Pamelamyia K87
            |         Orthovena K87
            |         Mesopetia GC99
            |         Oppenheimiella baltica P92
            |--Electrosania Grimaldi & Cumming 1999 GC99
            |    `--*E. cretica Grimaldi & Cumming 1999 GC99
            `--+--Melanderomyia [Melanderomyiinae] C10
               |    `--M. kahli K87
               |--Platypezininae K87
               |    |--Platypezina diversa K87
               |    `--Grossoseta pacifica K87
               |--Microsania [Microsaniinae] C10
               |    |--M. australis CM91
               |    |--M. imperfecta C10
               |    `--M. stigmaticalis C10
               |--Callomyiinae C10
               |    |--Bertamyia notata C10
               |    |--Chydaeopeza C10
               |    |--Callomyia C10
               |    |    |--C. calla C10
               |    |    `--C. gilloglyorum K87
               |    `--Agathomyia C10
               |         |--A. bella (Williston 1903) [=Callomyia bella] C10
               |         `--A. canadensis C10
               `--Platypezinae C10
                    |--Metaclythia currani K87
                    |--Polyporivora polypori K87, C10
                    |--Seri dymka K87
                    |--Paraplatypeza C10
                    |    |--P. coraxa K87
                    |    `--P. velutina WT11
                    |--Protoclythia K87
                    |    |--P. califormica K87
                    |    `--P. modesta K87
                    |--Platypeza C10
                    |    |--P. agarici RD77
                    |    |--P. brunnescens C10
                    |    `--P. consobrina K87
                    |--Lindneromyia [incl. Grossovena, Penesymmetria, Plesioclythia, Symmetricella] C10
                    |    |--L. agarici [=Plesioclythia agarici] C10
                    |    |--‘Grossovena’ carbonaria K87
                    |    |--‘Symmetricella’ mogollonensis K87
                    |    `--‘Penesymmetria’ umbrosa K87
                    `--Calotarsa K87
                         |--C. durangoensis Kessel & Young 1974 C10
                         |--C. insignis C10
                         |--C. mexicana Kessel & Young 1974 C10
                         `--C. pallipes C10

*Type species of generic name indicated


[C10] Chandler, P. J. 2010. Platypezidae (flat-footed flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 715–719. NRC Research Press: Ottawa.

[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).

[GC99] Grimaldi, D., & J. Cumming. 1999. Brachyceran Diptera in Cretaceous ambers and Mesozoic diversification of the Eremoneura. Bulletin of the American Museum of Natural History 239: 1–124.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[K87] Kessel, E. L. 1987. Platypezidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 681–688. Research Branch, Agriculture Canada.

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms' General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.

[WT11] Wiegmann, B. M., M. D. Trautwein, I. S. Winkler, N. B. Barr, J.-W. Kim, C. Lambkin, M. A. Bertone, B. K. Cassel, K. M. Bayless, A. M. Heimberg, B. M. Wheeler, K. J. Peterson, T. Pape, B. J. Sinclair, J. H. Skevington, V. Blagoderov, J. Caravas, S. N. Kutty, U. Schmidt-Ott, G. E. Kampmeier, F. C. Thompson, D. A. Grimaldi, A. T. Beckenbach, G. W. Courtney, M. Friedrich, R. Meier & D. K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the USA 108 (14): 5690–5695.

Last updated: 13 July 2021.

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