Diastata fuscula, copyright Dick Belgers.

Belongs within: Ephydroidea.

The Diastatidae are a group of small, drosophilid-like flies found primarily among lush, low vegetation, often in relatively damp locations (Mathis 2010).

Characters (from Mathis 2010): Adults small, greyish-brown, body length 2.5–4 mm; usually with patterned wings and weakly spinose costa. Postocellar setae convergent; both inner and outer verticals present; ocellar and vibrissal setae well developed; two well-developed fronto-orbital setae present, proclinate seta laterad of reclinate seta present. Antenna decumbent; arista elongate, macropubescent or short plumose. Anepisternum setose or bare; katepisternum bearing twelve setae, posterior seta stronger. C extending to M; C with either subcostal break only or both humeral and subcostal breaks; Sc incomplete, not reaching C, often fusing with R1 distally; R1 bare; both CuA2 and A1 present; cell cup closed, crossvein bm-cu present. Legs moderatly elongate, slender. Abdominal segments 1–5 unmodified; male with segments 6–8 reduced, although tergite 7 well developed, sternite 7 absent; abdominal spiracles (6 or 7) in membrane.

<==Diastatidae [Campichoetidae] M10
    |--Euthychaeta M10
    |--Pareuthychaeta M10
    |    |--P. electrica M87
    |    `--P. minuta M87
    |--Campichoeta M10
    |    |--C. basalis [incl. Thryptocheta tristis] M10
    |    |--C. latigena M10
    |    `--C. punctum WT11
    `--Diastata M10
         |--D. argentina M10
         |--D. costaricensis M10
         |--D. dimidiata M10
         |--D. fuscula WT11
         `--D. vagans M10

*Type species of generic name indicated


[M10] Mathis, W. N. 2010. Diastatidae (diastatid flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 1207–1210. NRC Research Press: Ottawa.

[M87] McAlpine, J. F. 1987. Diastatidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 1019–1022. Research Branch, Agriculture Canada.

[WT11] Wiegmann, B. M., M. D. Trautwein, I. S. Winkler, N. B. Barr, J.-W. Kim, C. Lambkin, M. A. Bertone, B. K. Cassel, K. M. Bayless, A. M. Heimberg, B. M. Wheeler, K. J. Peterson, T. Pape, B. J. Sinclair, J. H. Skevington, V. Blagoderov, J. Caravas, S. N. Kutty, U. Schmidt-Ott, G. E. Kampmeier, F. C. Thompson, D. A. Grimaldi, A. T. Beckenbach, G. W. Courtney, M. Friedrich, R. Meier & D. K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the USA 108 (14): 5690–5695.

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