Greenleaf desmodium Desmodium intortum, copyright Anita Gould.

Belongs within: Desmodieae.

Desmodium, tick clovers, is a genus of leguminous herbs and shrubs found in warmer parts of the world, a number of which have seeds covered in tiny hooked hairs that stick to animal skins or clothes for dispersal.

Characters (from Flora of China): Herbs, subshrubs, or shrubs. Leaves pinnately 3-foliolate or 1-foliolate by reduction of lateral leaflets, stipulate, stipellate. Flow­ers usually smaller, in racemes or panicles, axillary or terminal, rarely solitary or binate in leaf axils. Bracts persistent or early decid­uous; bracteoles present or absent. Calyx campanulate, 4- or 5-lobed, bilabiate; upper 2 lobes connate for all or most of length, entire or 2-toothed at apex, lower lobes free, longest. Corolla white, green-white, pink, purple, or violet; standard elliptic or obovate to nearly orbicular; wings more or less adherent to keel, wholly clawed. Stamens diadelphous (9+1), rarely monadelphous. Ovary usually sessile, with several to many ovules. Legume compressed, usually indehiscent, transversely segmented, upper (adaxial) suture slightly con­stricted or straight, lower (abaxial) one constricted; articles mostly ellipsoidal to quadrangular. Cotyledons epigeous.

    |--D. axillare J87
    |--D. barbatum CP13
    |--D. brownii LK14
    |--D. diffusum P03
    |--D. filiforme LK14
    |--D. gangeticum P03
    |--D. glareosum LK14
    |--D. hannii LK14
    |--D. heterocarpon LK14
    |    |--D. h. var. heterocarpon LK14
    |    `--D. h. var. strigosum LK14
    |--D. incanum C55
    |--D. intortum MM96
    |--D. muelleri LK14
    |--D. ormocarpoides B00
    |--D. ovalifolium S00
    |--D. paniculatum W10
    |--D. pulchellum P03
    |--D. pullenii LK14
    |--D. pycnotrichum LK14
    |--D. repandum BMM99
    |--D. scorpiurus (Swartz) Desvaux 1813 [=Hedysarum scorpiurus Swartz 1788] GB02
    |--D. tortuosum ACW01
    |--D. trichostachyum LK14
    |--D. triflorum P03 [=Hedysarum triflorum C55; incl. Nicolsonia reptans C55]
    |--D. umbellatum P88
    |--D. velutinum P03
    `--D. zonatum Miq. 1855 [=Meiobomia zonatum (Miq.) Kuntze 1891] SR07

*Type species of generic name indicated


[ACW01] Aguilar, H., C. C. Childers & W. C. Welbourn. 2001. Relative abundance and seasonal occurrence of mites in the family Tydeidae on citrus in Florida. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 376–380. CSIRO Publishing: Melbourne.

[B00] Braby, M. F. 2000. Butterflies of Australia: their identification, biology and distribution vol. 2. CSIRO Publishing: Collingwood (Victoria).

[BMM99] Braun, U., J. Mouchacca & E. H. C. McKenzie. 1999. Cercosporoid hyphomycetes from New Caledonia and some other South Pacific islands. New Zealand Journal of Botany 37: 297–327.

[C55] Candolle, A. de. 1855. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 2. Librairie de Victor Masson: Paris.

[CP13] Cardoso, D., R. T. Pennington, L. P. de Queiroz, J. S. Boatwright, B.-E. Van Wyk, M. F. Wojciechowski & M. Lavin. 2013. Reconstructing the deep-branching relationships of the papilionoid legumes. South African Journal of Botany 89: 58–75.

[GB02] Gopalakrishna Bhat, K. 2002. Additions to the flora of Karnataka. Journal of the Bombay Natural History Society 99 (3): 566–567.

[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum—Biological Sciences 32 (1): 1–136.

[LK14] Lyons, M. N., G. J. Keighery, L. A. Gibson & T. Handasyde. 2014. Flora and vegetation communities of selected islands off the Kimberley coast of Western Australia. Records of the Western Australian Museum Supplement 81: 205–244.

[MM96] Mound, L. A., & R. Marullo. 1996. The thrips of Central and South America: an introduction (Insecta: Thysanoptera). Memoirs on Entomology, International 6: 1–487.

[P03] Paul, T. K. 2003. Botanical observations on the Purulia pumped storage hydropower project area, Bagmundi Hills, Purulia district, West Bengal. Bulletin of the Botanical Survey of India 45: 121–142.

[P88] Polunin, I. 1988. Plants and Flowers of Malaysia. Times Editions: Singapore.

[SR07] Sankar, R. V., K. Ravikumar, N. M. Ganesh Babu & D. K. Ved. 2007. Botany of Anapady MPCA, Palghat district, Kerala with special emphasis on species of conservation concern. Bulletin of the Botanical Survey of India 49: 165–172.

[S00] Siddiqi, M. R. 2000. Tylenchida: Parasites of plants and insects 2nd ed. CABI Publishing: Wallingford (UK).

[W10] Whittington, A. E. 2010. Platystomatidae (signal flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 903–907. NRC Research Press: Ottawa.

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