Nurus

Nurus imperialis, from Mal et al. (2012).

Belongs within: Pterostichitae.

Nurus is a genus of very large, heavy-bodied ground beetles found in eastern Australia.

Nurus Motschulsky 1865 [incl. Nuridius Sloane 1890, Pachymelas Tschitschérine 1902] LM87
    |--*N. brevis Motschulsky 1865 [incl. Feronia solandersii Castelnau 1867] LM87
    |--N. atlas (Castelnau 1867) (see below for synonymy) LM87
    |--N. curtus (Chaudoir 1865) [=Feronia (Trichosternus) curta, *Pachymelas curtus] LM87
    |--N. fortis (Sloane 1890) [=*Nuridius fortis] LM87
    |--N. grandis (Sloane 1910) [=Nuridius grandis] LM87
    |--N. imperialis (Sloane 1895) [=Homalosoma imperiale] LM87
    |--N. latipennis (Sloane 1903) [=Castelnaudia latipennis] LM87
    |--N. medius Darlington 1961 LM87
    |--N. niger Chaudoir 1878 LM87
    |--N. nox Darlington 1961 LM87
    `--N. rex Darlington 1961 LM87

Nurus atlas (Castelnau 1867) [=Feronia atlas; incl. Homalosoma crassiforme Sloane 1900, F. obesa Castelnau 1867] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Prosopogmus

Prosopogmus sp., copyright Kipling Will.

Belongs within: Pterostichitae.

Prosopogmus is an Australasian genus of ground beetles with deeply impressed clypeo-ocular sulci on the head, a subcordiform to quadrate pronotum with two pairs of marginal setae, and transverse sulci on abdominal ventrites 3–6.

Prosopogmus Chaudoir 1865 [incl. Ceneus Chaudoir 1865, Hormochilus Chaudoir 1874, Ophryosternus Chaudoir 1873] LM87
    |--*P. impressifrons (Chaudoir 1865) [=Feronia (*Prosopogmus) impressifrons] LM87
    |--P. austrinus Sloane 1895 LM87
    |--P. boisduvalii (Castelnau 1867) [=Feronia boisduvalii] LM87
    |--‘Pterostichus (Poecilus)’ coracinus Erichson 1842 (see below for synonymy) LM87
    |--‘Feronia (*Ceneus) coracina’ Chaudoir 1865 non Pterostichus (Poecilus) coracinus Erichson 1842 LM87
    |--P. delicatulus Tschitschérine 1898 LM87
    |--P. foveipennis (Macleay 1871) [=Argutor foveipennis] LM87
    |--P. harpaloides (Chaudoir 1874) [=Feronia (Notonomus) harpaloides] LM87
    |--P. insperatus Sloane 1896 LM87
    |--P. interstitialis Sloane 1923 LM87
    |--P. leai Sloane 1920 LM87
    |--P. monochrous (Chaudoir 1865) (see below for synonymy) LM87
    |--P. namoyensis Sloane 1895 LM87
    |--P. occidentalis (Macleay 1888) [=Simodontus occidentalis] LM87
    |--P. oodiformis (Macleay 1871) [=Argutor oodiformis] LM87
    |--P. opacidermis Sloane 1923 LM87
    |--P. punctifer Sloane 1920 [=P. punctiferus] LM87
    |--P. reichei (Castelnau 1867) [=Feronia reichei] LM87
    |--P. rubricornis Sloane 1895 LM87
    |--P. sulcatulus (Chaudoir 1873) [=*Ophryosternus sulcatulus] LM87
    |--P. suspectus (Chaudoir 1878) [=Feronia (Ceneus) suspecta] LM87
    |--P. tasmanicus Sloane 1920 LM87
    `--P. yarrensis Sloane 1915 LM87

Prosopogmus monochrous (Chaudoir 1865) [=Feronia (Ceneus) monochroa, *Hormochilus monochrous; incl. Eccoptogenius feronoides Castelnau 1867, Feronia inedita Castelnau 1867, Harpalis quadraticollis Castelnau 1867] LM87

‘Pterostichus (Poecilus)’ coracinus Erichson 1842 [incl. Hypherpes chalybeipennis Chaudoir 1843, Prosopogmus chalybeipennis, Tibarisus niger Macleay 1871, Feronia vilis Castelnau 1867] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Sarticus

Sarticus cyaneocinctus, copyright Kipling Will.

Belongs within: Pterostichitae.

Sarticus is an Australian genus of flightless ground beetles with filiform antennae, a rounded prothorax and wide elytra with an abbreviated subscutellar stria.

Sarticus Motschulsky 1865 LM87
    |--S. (Sarticus) LM87
    |    |--*S. (S.) obesulus (Chaudoir 1865) (see below for synonymy) LM87
    |    |--S. (S.) aubei (Castelnau 1867) [=Feronia aubei] LM87
    |    |--S. (S.) blackburni (Sloane 1895) [=Pterostichus blackburni] LM87
    |    |--S. (S.) brevicornis Blackburn 1892 LM87
    |    |--S. (S.) civilis (Germar 1848) [=Pterostichus civilis] LM87
    |    |--S. (S.) cooki Sloane 1903 LM87
    |    |--S. (S.) coradgeri Sloane 1903 LM87
    |    |--S. (S.) cyaneocinctus (Chaudoir 1865) (see below for synonymy) LM87
    |    |--S. (S.) cycloderus (Chaudoir 1865) (see below for synonymy) LM87
    |    |--S. (S.) dampieri Sloane 1903 LM87
    |    |--S. (S.) discopunctatus (Chaudoir 1865) (see below for synonymy) LM87
    |    |--S. (S.) dixoni Sloane 1915 LM87
    |    |--S. (S.) elegantulus (Castelnau 1867) [=Feronia elegantula; incl. F. (Sarticus) ischna Chaudoir 1878] LM87
    |    |--S. (S.) esmeraldipennis (Castelnau 1867) [=Feronia esmeraldipennis] LM87
    |    |--S. (S.) freyi Straneo 1960 LM87
    |    |--S. (S.) habitans Sloane 1889 LM87
    |    |--S. (S.) impar Sloane 1903 LM87
    |    |--S. (S.) iriditinctus (Chaudoir 1865) [=Feronia (Steropus) iriditincta] LM87
    |    |--S. (S.) macleayi Sloane 1889 LM87
    |    |--S. (S.) monarensis Sloane 1889 LM87
    |    |--S. (S.) obscurus Blackburn 1892 LM87
    |    `--S. (S.) rockhamptoniensis (Castelnau 1867) [=Feronia rockhamptoniensis] LM87
    `--S. (Coronacanthus Macleay 1878) LM87
         |--S. (*C.) sulcatus (Macleay 1878) [=*Coronacanthus sulcatus] LM87
         `--S. (C.) obsolescens Moore 1963 LM87
              |--S. o. obsolescens LM87
              `--S. o. substriatus Moore 1963 LM87

Sarticus (Sarticus) cyaneocinctus (Chaudoir 1865) [=Feronia (Steropus) cyaneocincta; incl. F. azureomarginata Castelnau 1867, F. saphyreomarginata Castelnau 1867] LM87

Sarticus (Sarticus) cycloderus (Chaudoir 1865) [=Feronia (Steropus) cyclodera; incl. F. blagravii Castelnau 1867, F. mastersii Castelnau 1867, F. waterhousii Castelnau 1867] LM87

Sarticus (Sarticus) discopunctatus (Chaudoir 1865) [=Feronia (Steropus) discopunctata; incl. F. bonvouloirei Castelnau 1867, F. germari Castelnau 1867, Sarticus ovicollis Motschulsky 1865] LM87

*Sarticus (Sarticus) obesulus (Chaudoir 1865) [=Feronia (Steropus) obesula; incl. F. olivieri Castelnau 1867, *Sarticus orbicollis Motschulsky 1865, F. saphyripennis Castelnau 1867] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Rhytisternus

Rhytisternus liopleurus, copyright Birgit E. Rhode.

Belongs within: Pterostichitae.

Rhytisternus is a genus of ground beetles found in Australia and New Zealand, characterised by the presence of functional wings, the lack of a scrobal seta on the mandibles, antennae with the basal three segments glabrous, elytra with the third interstice impunctate, and striolate pro-episterna.

Rhytisternus Chaudoir 1865 [=Amastus Motschulsky 1865] LM87
    |--*R. liopleurus (Chaudoir 1865) (see below for synonymy) LM87
    |--R. angustulus Macleay 1888 LM87
    |--R. arnheimensis (Castelnau 1867) [=Feronia arnheimensis] LM87
    |--R. bovilli Blackburn 1890 LM87
    |--R. callabonnensis Sloane 1895 LM87
    |--R. cardwellensis Blackburn 1892 LM87
    |--R. carpentarius Sloane 1895 LM87
    |--R. cyathoderus (Chaudoir 1865) [=Feronia (Rhytisternus) cyathodera; incl. F. lachlandiensis Castelnau 1867] LM87
    |--R. froggatti (Macleay 1888) [=Omaseus froggatti] LM87
    |--R. gigas Sloane 1895 LM87
    |--R. laevis (Macleay 1883) [=Poecilus laevis] LM87
    |--R. limbatus Macleay 1888 LM87
    |--R. mastersii (Macleay 1871) [=Omaseus mastersii] LM87
    |--R. miser (Chaudoir 1865) [=Feronia (Rhytisternus) misera; incl. F. subcarbonaria Castelnau 1867] LM87
    |--R. nigellus Sloane 1895 LM87
    |--R. obtusus Sloane 1895 LM87
    |--R. plebeius (Chaudoir 1874) [=Feronia (Rhytisternus) plebeia] LM87
    |--R. puellus (Chaudoir 1865) [=Feronia (Rhytisternus) puella; incl. F. clarenciensis Castelnau 1867] LM87
    |--R. solidus Sloane 1895 LM87
    |--R. splendens Blackburn 1892 LM87
    |--R. stuarti Sloane 1895 LM87
    `--R. sulcatipes Blackburn 1888 LM87

*Rhytisternus liopleurus (Chaudoir 1865) [=Feronia (*Rhytisternus) liopleura; incl. *Amastus nigricolor Motschulsky 1865] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Simodontus

Simodontus australis, copyright Kipling Will.

Belongs within: Pterostichitae.

Simodontus is a genus of ground beetles found in southern Australia, characterised by the lack of a scrobal seta on the mandible, antennae with the three basal segments glabrous, transverse sulci on abdominal ventrites 4–6, and elytra with three punctures on the third interstice.

Simodontus Chaudoir 1843 LM87
    |--S. (Simodontus) LM87
    |    |--*S. (S.) aeneipennis Chaudoir 1843 LM87
    |    |--S. (S.) australis (Dejean 1828) (see below for synonymy) LM87
    |    |--S. (S.) brunneus (Castelnau 1867) [=Harpalus brunneus] LM87
    |    |--S. (S.) convexus Chaudoir 1873 LM87
    |    |--S. (S.) curtulus Chaudor 1873 LM87
    |    |--S. (S.) fortnumi (Castelnau 1867) [=Harpalus fortnumi] LM87
    |    |--S. (S.) grandiceps Sloane 1900 LM87
    |    |--S. (S.) holomelanus (Germar 1848) [=Pterostichus holomelanus; incl. S. orthomoides Chaudoir 1873] LM87
    |    |--S. (S.) laeviceps Sloane 1900 LM87
    |    |--S. (S.) leai Sloane 1898 LM87
    |    |--S. (S.) murrayensis Blackburn 1890 LM87
    |    |--S. (S.) occultus Sloane 1898 LM87
    |    |--S. (S.) picescens Chaudoir 1873 LM87
    |    |--S. (S.) rotundipennis (Castelnau 1867) (see below for synonymy) LM87
    |    |--S. (S.) sexfoveatus (Chaudoir 1878) [=Feronia (S.) sexfoveata] LM87
    |    `--S. (S.) transfuga Chaudoir 1873 LM87
    `--S. (Trochoglymmus Straneo 1937) LM87
         |--S. (*T.) rufipalpis (Sloane 1902) (see below for synonymy) LM87
         `--S. (T.) clermonti (Straneo 1937) [=Prosopogmus clermonti] LM87

Simodontus (Simodontus) australis (Dejean 1828) [=Feronia australis; incl. Orthomus antipodus Motschulsky 1865, Simodontus elongatus Chaudoir 1873] LM87

Simodontus (Simodontus) rotundipennis (Castelnau 1867) [=Phorticosomus rotundipennis; incl. S. mandibularis Sloane 1900] LM87

Simodontus (*Trochoglymmus) rufipalpis (Sloane 1902) [=Feronia rufipalpis, Notonomus rufipalpis, Prosopogmus (Trochoglymmus) rufipalpis] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Sphodrina

Laemostenus complanatus, copyright Udo Schmidt.

Belongs within: Platynini.

The Sphodrina are a primarily Palaearctic group of ground beetles characterised by features of the female genitalia: the gonostyli lack a fringe of setae on the penultimate segment, have the dorsal seta on the apical segment moved to the inner lateral margin, and possess preapical sensory setae on the apical segment.

Sphodrina [Sphodrini]
    |--Pristonychus Dejean 1828 R-JC01, LM87 [=Prystonychus (l. c.) LM87]
    |    |--*P. tericola (Herbst 1783) [=Carabus tericola] LM87
    |    |--P. barbarus Lucas 1846 E12
    |    |--P. complanatus R-JC01
    |    `--P. sardous Lucas 1846 E12
    `--Laemostenus Bonelli 1810 E04, LM87 [=Laemosthenus (l. c.) LM87]
         |--*L. venustus (Dejean 1828) [=Carabus venustus] LM87
         |--L. cavicola [=Antisphodrus cavicola] PN02
         |    |--L. c. cavicola PN02
         |    `--L. c. aeacus (Miller 1861) PN02
         `--L. complanatus (Dejean 1828) [=Pristonychus complanatus; incl. P. australis Blackburn 1888] LM87

*Type species of generic name indicated

REFERENCES

[E04] Emberson, R. M. 2004. A European ground beetle, Harpalus tardus (Panzer) (Coleoptera: Carabidae), in Canterbury, and the predisposition of European carabids to establish in exotic temperate locations. New Zealand Entomologist 27: 39–42.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[PN02] Poggi, R., G. Nonveiller, A. Colla, D. Pavićević & T. Rađa. 2001–2002. Thaumastocephalini, a new tribe of Pselaphinae for Thaumastocephalus folliculipalpus n. gen., n. sp., an interesting new troglobious species from central Dalmatia (Croatia) (Coleoptera, Staphylinidae, Pselaphinae). Annali del Museo Civico di Storia Naturale “Giacomo Doria” 94: 1–20.

[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.

Morion

Morion japonicum, copyright Katsuro Yahiro.

Belongs within: Harpalinae.

Morion is a cosmopolitan genus of ground beetles found in rotten logs.

Characters (from Bell 1990): Mandible long, scrobal seta absent; two orbital setae over each eye; antennal segments 5–10 submoniliform, short, slightly compressed; tibiae distinctly flattened.

<==Morion Latreille 1810 [=Morio (l. c.)] LM87
    |--*M. monilicornis (Latreille 1810) LM87 [=Harpalus monilicornis LM87, Carabus monilicornis L02]
    |--M. aridus OM08
    |--M. australis Castelnau 1867 [=Morio australis] LM87
    |--M. crassipes Sloane 1904 [=Morio crassipes] LM87
    |--M. dalbertisi Chaudoir 1880 [=Morio dalbertisi] LM87
    |--M. germanus Chaudoir 1880 [=Morio germanus] LM87
    |--M. japonicum (Bates 1883) I92
    |--M. longicollis Macleay 1871 LM87
    |--M. longipennis Putzeys 1875 LM87
    |--M. novaehollandiae Castelnau 1867 (see below for synonymy) LM87
    |--M. orientalis B89
    |--M. pachysomus Chaudoir 1880 LM87
    |--M. piceus Castelnau 1867 LM87
    |--M. stolidus M86
    `--M. victoriae Castelnau 1867 LM87

Morion novaehollandiae Castelnau 1867 [=Morio novaehollandiae; incl. Morio australasiae Chaudoir 1869, Morio seticollis Macleay 1871] LM87

*Type species of generic name indicated

REFERENCES

[B89] Bates, H. W. 1889. Viaggio di Leonardo Fea in Birmani e regioni vicine. XVI.—On some Carabidae from Burma collected by Mr. L. Fea. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 100–111.

Bell, R. T. 1990. Insecta: Coleoptera Carabidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1053–1092. John Wiley & Sones: New York.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[M86] Macleay, W. 1886. The insects of the Fly River, New Guinea, "Coleoptera". Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 136–157.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

Abacetus

Abacetus salzmanni, copyright Fred Chevaillot.

Belongs within: Pterostichitae.

Abacetus is a genus of ground beetles found in warmer regions of the Old World, characterised by filiform, slightly compressed antennae, palpi with an elongate terminal segment, a trapezoid pronotum, and tarsi with the first three segments dilated in the male.

Abacetus Dejean 1828 [Abacetini] LM87
    |--*A. gagates Dejean 1828 LM87
    |--A. aeneus [=Feronia (Argutor) aenea] G89
    |--A. amplicollis Bates 1889 B89
    |--A. angustior Macleay 1871 LM87
    |--A. ater Macleay 1871 LM87
    |--A. australasiae Chaudoir 1878 LM87
    |--A. bisignatus Bates 1889 B89
    |--A. haemorrhous Chaudoir 1878 (see below for synonymy) LM87
    |--A. haplosternus Chaudoir 1878 LM87
    |--A. quadratipennis Macleay 1888 LM87
    |--A. quadriguttatus B89
    |--A. quadripustulatus B89
    |--A. salzmanni B89
    |--A. simplex Blackburn 1890 LM87
    `--A. thouzeti (Castelnau 1867) (see below for synonymy) LM87

Abacetus haemorrhous Chaudoir 1878 [incl. A. crenulatus Blackburn 1890, A. flavipes Macleay 1888, A. macleayi Blackburn 1890] LM87

Abacetus thouzeti (Castelnau 1867) [=Drimostoma thouzeti; incl. A. australis Chaudoir 1870, D. vicina Castelnau 1867] LM87

*Type species of generic name indicated

REFERENCES

[B89] Bates, H. W. 1889. Viaggio di Leonardo Fea in Birmani e regioni vicine. XVI.—On some Carabidae from Burma collected by Mr. L. Fea. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 100–111.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 5–72.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Trechodini

Tasmanitachoides cf. hobarti, from here.

Belongs within: Conjunctae.

The Trechodini are a group of ground beetles characterised by an aedeagus that is dorsally open and gutter-like.

<==Trechodini B09
    |--Trechodes Blackburn 1901 LM87
    |    |--*T. secalioides (Blackburn 1890) [=Bembidium secalioides] LM87
    |    |--T. bipartitus (Macleay 1871) [=Bembidium bipartitum] LM87
    |    |--T. bitinctus (Sloane 1923) [=Trechus bitinctus] LM87
    |    |--T. crypticus Moore 1972 LM87
    |    `--T. lustrans Moore 1972 LM87
    `--Tasmanitachoides Erwin 1972 B09
         |--*T. hobarti (Blackburn 1901) [=Bembidion hobarti] B09
         |--T. arnhemensis Erwin 1972 LM87
         |--T. balli B09
         |--T. bolus (Darlington 1962) [=Tachys bolus] LM87
         |--T. fitzroyi (Darlington 1962) [=Tachys fitzroyi] LM87
         |--T. glabellus B09
         |--T. hackeri B09
         |--T. hendrichi B09
         |--T. katherinei Erwin 1972 LM87
         |--T. kingi (Darlington 1962) [=Tachys kingi] LM87
         |--T. lutus (Darlington 1962) [=Tachys lutus] LM87
         |--T. mandibularis Baehr 2009 B09
         |--T. minor B09
         |--T. murrumbidgensis (Sloane 1895) [=Tachys murrumbidgensis] LM87
         |--T. obliquiceps B09
         `--T. wattsense (Blackburn 1901) [=Bembidium wattsense] LM87

*Type species of generic name indicated

REFERENCES

[B09] Baehr, M. 2009. A new species of the tachyine genus Tasmanitachoides from the Kimberley Division, Western Australia (Coleoptera, Carabidae). Records of the Western Australian Museum 25 (2): 159–164.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Pogonus

Pogonus chalceus, copyright Udo Schmidt.

Belongs within: Conjunctae.

Pogonus is a cosmopolitan genus found in saline habitats such as along coastlines or around salt lakes. They possess a scrobal seta on the mandible, and the head is not constricted behind the eyes.

<==Pogonus Dejean 1821 B10
    |--*P. chalceus (Marsham 1802) [=Carabus chalceus] B10
    |--P. australis Chaudoir 1878 B10
    |--P. cardiotrachelus Chaudoir 1871 B10
    |--P. dichrous Baehr 2010 B10
    |--P. diplochaetoides Baehr 1997 B10
    |--P. dostali Baehr 2010 B10
    |--P. fenelli Hudson 2000 B10
    |--P. gilesi Moore 1977 LM87
    |--P. grossi Moore 1977 LM87
    |--P. hypharpagioides Sloane 1895 LM87
    |--P. nigrescens Baehr 1984 B10
    |--P. orientalis G89
    |--P. sumlini Baehr 1999 B10
    |--P. variabilis Moore 1991 B10
    `--P. zietzi Sloane 1895 B10

*Type species of generic name indicated

REFERENCES

[B10] Baehr, M. 2010. Two new species of the genus Pogonus from northern and western Australia (Coleoptera: Carabidae). Records of the Western Australian Museum 25 (4): 441–448.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 5–72.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Tasmanorites

Tasmanorites grossus, from here.

Belongs within: Trechinae.

Tasmanorites is a genus of flightless ground beetles found in damp habitats in Tasmania. They are characterised by small discal pores on the elytra, with the second elytral stria terminating near the posterior discal pore placed well back on the elytral declivity.

Tasmanorites Jeannel 1927 LM87
    |--*T. nitens (Putzeys 1874) [=Trechus nitens] LM87
    |--T. aberrans Moore 1972 LM87
    |--T. austrinus (Sloane 1920) [=Trechus austrinus] LM87
    |--T. blackburni (Sloane 1920) [=Trechus blackburni] LM87
    |--T. brevinotatus (Sloane 1920) [=Trechus brevinotatus] LM87
    |--T. cordicollis Moore 1972 LM87
    |--T. elegans Moore 1972 LM87
    |--T. flavipes (Lea 1910) [=Idacarabus flavipes] LM87
    |--T. glaebarum Moore 1972 LM87
    |--T. grossus Moore 1972 LM87
    |--T. intermedius Moore 1972 LM87
    |--T. laticollis Moore 1983 LM87
    |--T. longinotatus (Sloane 1920) [=Trechus longinotatus] LM87
    |--T. madidus Moore 1972 LM87
    |--T. magnus Moore 1972 LM87
    |--T. pullus Moore 1972 LM87
    |    |--T. p. pullus LM87
    |    `--T. p. minor Moore 1972 LM87
    |--T. riparius Moore 1972 LM87
    `--T. tasmaniae (Blackburn 1901) [=Trechus tasmaniae] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Rhysodinae

Kaveinga abbreviata, copyright David R. Maddison.

Belongs within: Conjunctae.

The Rhysodinae are a group of ground beetles found as both adults and larvae in old, rotted logs (Lawrence & Britton 1991).

Characters (from Lawrence & Britton 1991): Black and shiny, subcylindrical to slightly flattened, head and pronotum with deep grooves that are often filled with dried organic material; antennae stout, moniliform, 11-segmented; mouthparts more or less concealed beneath expanded mentum; head strongly constricted posteriorly to form narrow neck; fore coxae globular, trochantins concealed, fore coxal cavities externally closed (internally open); mid coxal cavities moderately separated, laterally open; metasternum without transverse suture; legs relatively short, with weakly developed antenna cleaner on fore tibia, tarsal formula 5-5-5; abdomen with six ventrites, three connate. Larvae elongate, somewhat flattened and lightly sclerotised with short legs, no urogomphi, and transverse rows of asperities or interrupted ridges on T1 to 6 or 7; head prognathous, protracted; stemmata absent; antenna four-segmented; frontoclypeal suture absent; labrum fused; mandibular mola absent; maxillae and labium reduced, usually clothed with fringed membranes, maxilla with single mala, maxillary palp three-segmented, labial palp one-segmented; hypostomal rods present; legs six-segmented, with one tarsal claws; urogomphi absent; spiracles annular.

<==Rhysodinae [Rhysodidae]
    |--Leoglymmius Bell & Bell 1978 [Leoglymmiini] LM87
    |    `--*L. lignarius (Olliff 1885) (see below for synonymy) LM87
    |--Rhysodini LM87
    |    |--Rhysodes sulcatus LM87
    |    `--Kaveinga Bell & Bell 1978 LM87
    |         |--*K. (Kaveinga) abbreviata (Lea 1904) [=Rhysodes abbreviatus] LM87
    |         `--K. (Angekiva Bell & Bell 1978) LM87
    |              `--K. (*A.) frontalis (Grouvelle 1903) [=Rhysodes frontalis] LM87
    |--Clinidiini LM87
    |    |--Clinidium LM87
    |    `--Rhyzodiastes Fairmaire 1895 LM87
    |         |--*R. parumcostatus (Fairmaire 1868) [=Rhysodes parumcostatus] LM87
    |         |--R. burnsi (Oke 1932) [=Rhysodes burnsi] LM87
    |         `--R. mirabilis (Lea 1904) [=Rhysodes mirabilis] LM87
    `--Ommoglymiini LM87
         |--Ommoglymius Ganglbauer 1892 LM87
         |    |--*O. germari (Ganglebauer 1892) [=Rhysodes germari] LM87
         |    `--O. ichthyocephalus (Lea 1904) [=Rhysodes ichthyocephalus] LM87
         `--Yamatosa Bell & Bell 1979 [=Yamatoa Bell 1977 non Kiriakoff 1967] LM87
              |--*Y. niponensis (Lewis 1888) [=*Yamatoa niponensis] LM87
              `--Y. planata (Lea 1904) [=Rhysodes planatus] LM87

*Leoglymmius lignarius (Olliff 1885) [=Rhysodes lignarius; incl. R. blackburni Grouvelle 1903, R. trichosternus Lea 1904] LM87

*Type species of generic name indicated

REFERENCES

Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Tachyina

Tachys scutellaris, copyright Udo Schmidt.

Belongs within: Bembidiinae.

The Tachyina are a cosmopolitan group of ground beetles characterised by an apicolateral notch on the protibiae, and usually an apical recurrent groove on the elytra.

<==Tachyina R-JC01
    |--Pericompsus LeConte 1852 R-JC01, LM87
    |    |--*P. (Pericompsus) ephippiatus (Say 1834) [=Bembidium ephippiatum] LM87
    |    `--P. (Upocompsus Erwin 1974) LM87
    |         |--P. (*U.) australis (Schaum 1863) (see below for synonymy) LM87
    |         |--P. (U.) bogani (Darlington 1963) [=Tachys bogani] LM87
    |         |--P. (U.) habitans (Sloane 1896) [=Tachys habitans] LM87
    |         |--P. (U.) olliffi (Sloane 1896) [=Tachys olliffi] LM87
    |         |--P. (U.) pubifrons (Darlington 1963) [=Tachys pubifrons] LM87
    |         |--P. (U.) punctipennis (Macleay 1871) [=Bembidium punctipenne] LM87
    |         |--P. (U.) semistriatus (Blackburn 1888) [=Tachys semistriatus] LM87
    |         |--P. (U.) seticollis (Sloane 1896) [=Tachys seticollis] LM87
    |         `--P. (U.) yarrensis (Blackburn 1892) [=Tachys yarrensis] LM87
    `--Tachys Stephens 1828 (see below for synonymy) LM87
         |--*T. scutellaris Stephens 1828 LM87
         |--‘Elaphropus’ aethiopicus B55
         |--‘Elaphropus’ afer B55
         |--T. amplipennis (Macleay 1871) [=Bembidium amplipenne] LM87
         |--‘Tachyura’ apicalis B55
         |--T. atridermis Sloane 1921 LM87
         |--‘*Paratachys’ austinicus Casey 1918 LM87
         |--T. australicus Sloane 1896 LM87
         |--T. baldiensis Blackburn 1891 LM87
         |--T. banksi Sloane 1921 LM87
         |--T. bembidiiformis Jordan 1894 [incl. T. helmsi Sloane 1898] LM87
         |--‘Eotachys’ bicoloratus B55
         |--T. bioculatus Putz 1875 B89
         |--T. bipustulatus (Macleay 1871) [=Bembidium bipustulatum; incl. T. froggatti Sloane 1896] LM87
         |--‘Elaphrus’ bistriatus Duftschmid 1812 [=*Eotachys bistriatus] LM87
         |--T. blackburni Sloane 1921 LM87
         |--T. bolellus Darlington 1962 LM87
         |--T. brevicornis Chaudoir 1846 [=*Polyderis brevicornis] LM87
         |--T. buprestioides Sloane 1896 LM87
         |--‘Eotachys’ caffer B55
         |--T. captus Blackburn 1888 LM87
         |--‘*Elaphropus’ caraboides Motschulsky 1862 LM87
         |--T. carinulatus Sloane 1921 LM87
         |--‘Tachyura’ conspicua B55
         |--T. convexicollis GWW10
         |--T. convexulus Darlington 1963 LM87
         |--T. convexus (Macleay 1871) (see below for synonymy) LM87
         |--T. cruciger Putzeys 1875 [incl. T. queenslandicus Sloane 1903] LM87
         |--T. curticollis Sloane 1896 LM87
         |--‘Tachylopha’ devroeyiana B55
         |--T. ectromioides Sloane 1896 LM87
         |--‘Tachyura’ emerita B55
         |--T. fasciatus Motschulsky 1851 LM87 (see below for synonymy)
         |--‘Tachyura’ fausta B55
         |--T. flavicornis Sloane 1921 LM87
         |--‘Tachylopha’ gerardiana B55
         |--‘Eotachys’ gilvus B55
         |--‘Tachyura’ grandicollis [=Amaurotachys grandicollis] B55
         |    |--T. g. grandicollis B55
         |    |--T. g. michaelseni B55
         |    `--T. g. nigrolimbatus B55
         |--T. iaspideus Sloane 1896 LM87
         |--T. impressipennis Motschulsky 1860 LM87 (see below for synonymy)
         |--T. infuscatus Blackburn 1888 LM87
         |--T. latissimus (Motschulsky 1851) (see below for synonymy) LM87
         |--T. leai Sloane 1896 LM87
         |--‘Polyderis’ leleupi Basilewsky 1953 B55
         |--‘Eotachys’ leopoldvilleanus B55
         |--T. lindi Blackburn 1888 LM87
         |--T. lividus (Bates 1871) [=Tachyta livida] LM87
         |--T. lucasii Jacq. Duv. 1852 G89
         |--T. macleayi Sloane 1896 LM87
         |--T. mastersi Sloane 1921 [=Bembidium sexstriatum Macleay 1871 non Elaphrus sexstriatus Duftschmid 1812] LM87
         |--T. mitchelli Sloane 1895 LM87
         |--T. mulwalensis Sloane 1900 LM87
         |--T. nervosus Sloane 1903 LM87
         |--T. obliquiceps Sloane 1903 LM87
         |--‘Lopha’ ovata Motschulsky 1851 [=*Tachylopha ovata] LM87
         |--T. ovensensis Blackburn 1891 LM87
         |--T. plagiatus Putzeys 1875 [incl. T. doddi Sloane 1903] LM87
         |--‘Elaphrus’ quadrisignatus Duftschmidt 1812 [=*Tachyura quadrisignata] LM87
         |--T. similis Blackburn 1888 LM87
         |--T. solidus Sloane 1921 LM87
         |--T. spenceri Sloane 1896 LM87
         |--T. striolatus (Macleay 1871) [=Bembidium striolatum; incl. T. brightensis Blackburn 1891] LM87
         |--T. sydneyensis Sloane 1923 LM87
         |--T. transversicollis (Macleay 1871) [=Bembidium transversicolle; incl. T. adelaidae Blackburn 1888] LM87
         |--T. trunci Darlington 1963 LM87
         |--T. uniformis Blackburn 1888 LM87
         |--‘Tachyura’ vagans B55
         |--‘Tachyura’ variabilis B55
         |--T. victoriensis (Blackburn 1891) [=Bembidium victoriense] LM87
         |--‘Tachyura’ viverra B55
         |--T. wilsoni Sloane 1921 LM87
         `--T. zonatus Andrewes 1925 [=T. ephippiatus Putzeys 1875 non Bembidion ephippiatum Say 1834] LM87

Pericompsus (*Upocompsus) australis (Schaum 1863) [=Tachys australis; incl. T. flindersi Blackburn 1888, T. monochrous Schaum 1863, Bembidium rubicundum Macleay 1871, B. tersatum Broun 1893] LM87

Tachys Stephens 1828 [incl. Elaphropus Motschulsky 1839, Eotachys Jeannel 1941, Paratachys Casey 1918, Polyderis Motschulsky 1862, Tachylopha Motschulsky 1862, Tachyura Motschulsky 1862] LM87

Tachys convexus (Macleay 1871) [=Bembidium convexum; incl. B. bistriatum Macleay 1871 non Elaphrus bistriatus Duftschmid 1812] LM87

Tachys fasciatus Motschulsky 1851 LM87 [=Trechus fasciatus LM87, Eotachys fasciatus B55; incl. Bembidium atriceps Macleay 1871 LM87, Tachys trechiformis Jordan 1894 LM87, B. triangulare Nietner 1858 LM87]

Tachys impressipennis Motschulsky 1860 LM87 [=Eotachys impressipennis B55; incl. T. dohertyi Jordan 1894 LM87, T. sinuaticollis Sloane 1903 LM87]

Tachys latissimus (Motschulsky 1851) [=Elaphropus latissimus; incl. Bembidium bifoveatum Macleay 1871, B. ovatum Macleay 1871] LM87

*Type species of generic name indicated

REFERENCES

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B89] Bates, H. W. 1889. Viaggio di Leonardo Fea in Birmani e regioni vicine. XVI.—On some Carabidae from Burma collected by Mr. L. Fea. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 100–111.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 5–72.

[GWW10] Guthrie, N. A., T. Weir & K. Will. 2010. Localised and regional patterns in ground-dwelling beetle assemblages in a semi-tropical arid zone environment. Records of the Western Australian Museum Supplement 78 (1): 169–184.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.

Laccopterum

Laccopterum deauratum, copyright Dash Huang.

Belongs within: Scaritinae.

Laccopterum is an Australian genus of scaritine ground beetles characterised by ovate and foveate elytra, tridentate anterior tibiae and two supra-orbital punctures.

<==Laccopterum Macleay 1878 [=Laccoscaphus Sloane 1905] LM87
    |--L. macleayi Sloane 1897 [=Carenum spencei Macleay 1878 non Westwood 1841, *Laccopterum spencei] LM87
    |--L. cyaneum (Fabricius 1775) (see below for synonymy) LM87
    |--L. darwiniense (Macleay 1878) [=Carenum darwiniense] LM87
    |--L. deauratum (Macleay 1864) [=Carenum deauratum] LM87
    |--L. doddi (Sloane 1916) [=Laccoscaphus doddi; incl. Laccoscaphus doddi triordinata Sloane 1917] LM87
    |--L. foveigerum (Chaudoir 1868) [=Carenum foveigerum] LM87
    |--L. foveipenne (Macleay 1873) [=Carenum foveipenne] LM87
    |--L. foveolatum (Macleay 1864) [=Carenum foveolatum] LM87
    |--L. gemmatum (Westwood 1842) [=Carenum gemmatum] LM87
    |--L. humerale Sloane 1900 LM87
    |--L. lacunosum Macleay 1887 LM87
    |--L. loculosum (Newman 1842) [=Carenum loculosum; incl. C. variolosum Macleay 1864] LM87
    |--L. multiimpressum (Castelnau 1867) [=Carenum multiimpressum] LM87
    |--L. quadriseriatum (Sloane 1907) [=Laccoscaphus quadriseriatus] LM87
    |--L. salebrosum (Macleay 1871) [=Carenum salebrosum] LM87
    `--L. spencei (Westwood 1841) (see below for synonymy) LM87

Laccopterum cyaneum (Fabricius 1775) [=Scarites cyaneus; incl. Laccoscaphus cyaneus aenescens Sloane 1923, Carenum fabricii Westwood 1842] LM87

Laccopterum spencei (Westwood 1841) [=Carenum spencei; incl. C. digglesi Macleay 1873 non Eutoma digglesi Macleay 1869] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Scaritinae

Big-headed ground beetle Scarites subterraneus, copyright Kaldari.

Belongs within: Carabidae.
Contains: Laccopterum, Scaraphites, Conopterum, Neocarenum, Monocentrum, Euryscaphus, Carenum, Dyschirius, Clivina, Dyschiriodes.

The Scaritinae are a diverse group of ground beetles with body forms commonly modified for digging. The body is pedunculate with a marked constriction between pro- and mesothorax. The fore coxal cavities are closed behind and the mid coxal cavities laterally open, partially closed by the mesepimeron (Lawrence & Britton 1991). The tribe Clivinini includes relatively small forms with a prominent, setiform unguitractor plate on the tarsus whereas members of the Scaritini have a much smaller plate and tend to be much larger in overall size (Ball & Bousquet 2001).

Scaritinae [Scarititae, Scrobifera] B14
    |--Migadopini [Migadopinae] R-JC01
    |    |--Migadops latus R-JC01
    |    `--Stichonotus Sloane 1910 LB91, LM87
    |         |--*S. leai Sloane 1910 LM87
    |         |--S. limbatus Sloane 1915 LM87
    |         `--S. piceus Sloane 1915 LM87
    |--Scaritini B90
    |    |--Euryscaphus LB91
    |    |--Carenum LB91
    |    |--Dyschirius B90
    |    |--Pasimachus Bonelli 1813 BB01
    |    |    |  i. s.: P. elongatus LeConte 1846 BB01
    |    |    |         P. subangulatus (Chaudoir 1862) B14
    |    |    `--P. (Emydopterus Lacordaire 1854) BB01
    |    `--Scarites Fabricius 1775 BB01
    |         |--S. eurytus G89
    |         |--S. gigas C74
    |         |--S. guineensis [incl. S. guineensis var. asphaltinus] G89
    |         |--S. pyracmon B35
    |         |--S. rugosus B35
    |         |--S. senegalensis B55
    |         `--S. (Scarites) subterraneus Fabricius 1775 BB01
    `--Clivinini BB01
         |--Clivina BB01
         |--Ardistomis Putzeys 1846 BB01
         |--Dyschiriodes BB01
         |--Oxydrepanus Putzeys 1866 BB01
         |    `--O. rufus (Putzeys 1846) BB01
         |--Halocoryza Alluaud 1919 BB01
         |    `--H. arenaria (Darlington 1939) BB01
         |--Aspidoglossa Putzeys 1846 BB01
         |    `--A. subangulata (Chaudoir 1843) BB01
         |--Semiardistomis Kult 1950 BB01
         |    |--S. puncticollis (Dejean 1831) BB01
         |    `--S. viridis (Say 1823) BB01
         |--Akephorus LeConte 1852 BB01
         |    |--A. marinus LeConte 1852 BB01
         |    `--A. obesus LeConte 1866 B14
         `--Schizogenius Putzeys 1846 BB01
              |  i. s.: S. freyi Baehr 1983 BB01
              `--S. (Genioschizus Whitehead 1972) BB01
                   `--S. (G.) crenulatus LeConte 1852 BB01

Scaritinae incertae sedis:
  Ochyropus gigas B55
  Taeniolobus B55
    |--T. laevipennis B55
    |--T. lucidus B55
    |    |--T. l. lucidus B55
    |    `--T. l. strigiceps B55
    `--T. vilhenai Basilewsky 1955 B55
  Trilophus congoanus B55
  Syleter Andrewes 1941 [=Psilus Putzeys 1877 nec Panzer 1806 nec Fischer de Waldheim 1813] LM87
    |--*S. paradoxus (Putzeys 1868) [=Ardistomis paradoxa] LM87
    |--S. andrewesi (Basilewsky 1931) [=Dyschirius andrewesi, Psilus andrewesi] B55
    |--S. congoensis B55
    `--S. papua Darlington 1962 LM87
  Pseudoclivina Kult 1947 B55
    `--P. grandis B55
  Rhysocara Sloane 1916 LM87
    `--*R. crassa Sloane 1916 LM87
  Platysphyrus Sloane 1905 LM87
    `--*P. tibialis Sloane 1905 LM87
  Clivinarchus Sloane 1896 LM87
    `--*C. perlongus Sloane 1896 LM87
  Laccopterum LM87
  Philoscaphus Macleay 1871 LM87
    |--*P. tuberculatus (Macleay 1863) [=Carenum tuberculatum] LM87
    |--P. barnardi Macleay 1888 LM87
    |--P. bicostatus Sloane 1905 LM87
    |--P. carinatus (Macleay 1864) [=Carenum carinatum] LM87
    |--P. costalis Macleay 1873 (see below for synonymy) LM87
    `--P. mastersii Macleay 1871 [incl. P. duboulayi Blackburn 1890] LM87
  Scaraphites LM87
  Conopterum LM87
  Neocarenum LM87
  Trichocarenum Blackburn 1892 LM87
    |--*T. elderi Blackburn 1892 LM87
    |--T. castelnaui Sloane 1905 LM87
    `--T. cylindricum Sloane 1897 LM87
  Neoscaphus Sloane 1888 LM87
    `--*N. simplex Sloane 1888 LM87
  Epilectus Blackburn 1888 [=Carenarchus Sloane 1905, Eurygnathus Blackburn 1888 non Wollaston 1854] LM87
    |--*E. fortis (Blackburn 1888) [=*Eurygnathus fortis, *Carenarchus fortis] LM87
    `--E. mastersi (Macleay 1869) [=Neocarenum mastersi] LM87
  Monocentrum LM87
  Steganomma Macleay 1887 LM87
    `--*S. porcatum Macleay 1887 LM87
  Geoscaptus Chaudoir 1855 LM87
    |--*G. laevissimus Chaudoir 1855 (see below for synonymy) LM87
    |--G. cacus (Macleay 1863) (see below for synonymy) LM87
    |--G. crassus Sloane 1895 LM87
    |--G. macleayi Chaudoir 1879 LM87
    `--G. plicatulus (Castelnau 1867) [=Scarites plicatulus] LM87
  Omoglymmius B14
    |--O. americanus (Laporte 1836) B14
    `--O. hamatus B14
  Solenogenys funkei Adis 1981 B14

Geoscaptus cacus (Macleay 1863) [=Scarites cacus; incl. S. approximatus Macleay 1865, S. bipunctatus Castelnau 1867, S. mitchellii Castelnau 1867, S. ruficornis Castelnau 1867, S. subporcatulus Macleay 1865, S. substriatus Castelnau 1867, S. waterhousei Macleay 1865] LM87

*Geoscaptus laevissimus Chaudoir 1855 [incl. Scarites bostocki Castelnau 1867, S. damastes Macleay 1863, S. geryon Macleay 1863, S. jacksoniensis Macleay 1865, S. planiusculus Macleay 1865] LM87

Philoscaphus costalis Macleay 1873 [incl. P. crassus Blackburn 1888, P. lateralis Macleay 1873, P. tepperi Blackburn 1888] LM87

*Type species of generic name indicated

REFERENCES

[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B90] Bell, R. T. 1990. Insecta: Coleoptera Carabidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1053–1092. John Wiley & Sones: New York.

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d'Urville. Faune entomologique de l'océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[C74] Coineau, Y. 1974. Éléments pour une monographie morphologique, écologique et biologique des Caeculidae (Acariens). Mémoires du Muséum National d’Histoire Naturelle, nouvelle série, Série A, Zoologie 81: 1–299, 24 pls.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso “Esploratore” dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 5–72.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2^nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.

Last updated: 19 December 2020.

Adelotopus

Adelotopus haemorrhoidalis, copyright Udo Schmidt.

Belongs within: Pseudomorphini.

Adelotopus is a genus of pseudomorphin ground beetles found in Australia and south-east Asia, characterised by a reduced chaetotaxy and foliaceous female stylomeres (Baehr 2008).

<==Adelotopus Hope 1834 B08
    |--*A. gyrinoides Hope 1834 B08
    |--A. adustus Baehr 1997 B08
    |--A. affinis Castelnau 1867 LM87
    |--A. analis Macleay 1871 LM87
    |--A. aphodioides Westwood 1853 LM87
    |--A. apicalis Macleay 1864 LM87
    |--A. basalis Baehr 2008 B08
    |--A. bimaculatus Macleay 1864 [incl. Silphomorpha rufoguttata Blackburn 1893] LM87
    |--A. bolitus OM08
    |--A. brevipennis Macleay 1888 B08
    |--A. brunneus Castelnau 1867 LM87
    |--A. celeripes Lea 1910 LM87
    |--A. coriaceus Baehr 1997 B08
    |--A. cornutus Castelnau 1867 LM87
    |--A. dysticides Newman 1842 [incl. A. fortnumi Hope 1845] LM87
    |--A. elongatulus Macleay 1888 LM87
    |--A. fasciatus Castelnau 1867 LM87
    |--A. filiformis Castelnau 1867 LM87
    |--A. flavus B08
    |--A. haemorrhoidalis Erichson 1842 [incl. A. inquinatus Newman 1842] LM87
    |--A. houstoni Baehr 1997 B08
    |--A. hydrobioides Westwood 1853 LM87
    |--A. insignis Sloane 1910 LM87
    |--A. laevis Macleay 1888 LM87
    |--A. linearis Macleay 1888 LM87
    |--A. longipennis Macleay 1888 LM87
    |--A. maculipennis Macleay 1871 LM87
    |--A. mastersii Macleay 1871 LM87
    |--A. micans Blackburn 1901 LM87
    |--A. minutus Baehr 2008 B08
    |--A. nemosomoides Westwood 1853 LM87
    |--A. niger Notman 1925 LM87
    |--A. occidentalis Castelnau 1867 LM87
    |--A. paroensis Castelnau 1867 B08
    |--A. piceus B08
    |--A. pilbarae Baehr 2008 B08
    |--A. politus Castelnau 1867 LM87
    |--A. punctatus Castelnau 1867 LM87
    |--A. puncticollis Notman 1925 LM87
    |    |--A. p. puncticollis B08
    |    `--A. p. angustemaculatus Baehr 1997 B08
    |--A. rubiginosus Newman 1856 B08 [incl. A. castaneus Castelnau 1867 B08, LM87]
    |--A. rufescens B08
    |--A. scolytides Newman 1842 LM87
    |--A. seriepunctatus Notman 1925 LM87
    |--A. tasmani Blackburn 1901 LM87
    |--A. variolosus Lea 1910 LM87
    |--A. vicinus Castelnau 1867 LM87
    `--A. zonatus Castelnau 1867 LM87

*Type species of generic name indicated

REFERENCES

[B08] Baehr, M. 2008. New and rare pseudomorphine species from Western Australia (Insecta, Coleoptera, Carabidae, Pseudomorphinae). Records of the Western Australian Museum 24 (2): 151–180.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

Pseudomorphini

Graphipterus serrator, copyright Hectonichus.

Belongs within: Harpalinae.
Contains: Adelotopus, Sphallomorpha.

The Pseudomorphini are a group of often flattened ground beetles found living under bark. They are often found in association with ants and larvae of some species are physogastric inquilines in ant nests (Lawrence & Britton 1991). The majority of genera in this group are ovoviviparous (Baehr 2008).

Characters (from Lawrence & Britton 1991): Antennal scape not visible from above; antennae sometimes short and clavate; head with deep antennal sulcus between eye and mandible; legs more or less concealed beneath body, with may be flattened and laterally explanate.

<==Pseudomorphini [Pseudomorphinae]
    |  i. s.: Cryptocephalomorpha B08
    |--+--Pseudomorpha Kirby 1825 OM08, LM87
    |  |    |--*P. excrucians Kirby 1825 LM87
    |  |    |--P. angustata OM08
    |  |    `--P. confusa Notman 1925 LM87
    |  `--Graphipterus Latreille 1802 OM08, L02 [Graphipterini]
    |       |--G. circumdatus Raffr. 1885 G89
    |       |--G. cordiger OM08
    |       |--G. limbatus OM08
    |       |--G. minutus B14
    |       |--G. serrator (Forskål 1775) [=Carabus serrator] G89
    |       |--G. trilineatus [=Carabus trilineatus] L02
    |       `--G. variegatus [=Carabus variegatus] L02
    `--+--+--Adelotopus OM08
       |  |--Paussotropus Waterhouse 1877 B08
       |  |    `--P. cylindricus (Chaudoir 1862) (see below for synonymy) B08
       |  `--Cainogenion Notman 1925 B08
       |       |--*C. ipsoides (Newman 1837) [=Adelotopus ipsoides] B08
       |       |    |--C. i. ipsoides B08
       |       |    `--C. i. occidentale Baehr 1997 B08
       |       |--C. creberrimum (Blackburn 1901) [=Adelotopus creberrimus] LM87
       |       |--C. ephippiatum (Newman 1856) [=Adelotopus ephippiatus] LM87
       |       `--C. obscurum (Castelnau 1867) [=Adelotopus obscurus; incl. A. subopacus Macleay 1871] LM87
       `--+--Orthogonius [Orthogoniinae, Orthogoniini] OM08
          |    |--O. brevilabris B55
          |    |--O. clarkei B55
          |    |--O. davidis B89
          |    |--O. duplicatus [incl. O. alternans] B89
          |    `--O. profundestriatus B89
          `--Sphallomorpha OM08

Paussotropus cylindricus (Chaudoir 1862) [=Adelotopus cylindricus, Cainogenion cylindricum; incl. *Paussotropus parellelus Waterhouse 1877] B08

*Type species of generic name indicated

REFERENCES

[B08] Baehr, M. 2008. New and rare pseudomorphine species from Western Australia (Insecta, Coleoptera, Carabidae, Pseudomorphinae). Records of the Western Australian Museum 24 (2): 151–180.

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B89] Bates, H. W. 1889. Viaggio di Leonardo Fea in Birmani e regioni vicine. XVI.—On some Carabidae from Burma collected by Mr. L. Fea. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 100–111.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 5–72.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

Neocarenum

Neocarenum angustatum, copyright Museums Victoria.

Belongs within: Scaritinae.

Neocarenum is an Australian genus of scaritine ground beetles characterised by moniliform antennae, sub-securiform labial palpi and bidentate anterior tibiae.

<==Neocarenum Castelnau 1867 LM87
    |--N. singulare Castelnau 1867 [=N. singularis] LM87
    |--N. angustatum Sloane 1894 LM87
    |--N. blackburni Sloane 1895 LM87
    |--N. cylindripenne Bates 1874 LM87
    |--N. dingo Sloane 1916 LM87
    |--N. elongatum (Macleay 1864) [=Carenum elongatum; incl. N. kreusleri Castelnau 1867] LM87
    |--N. parviceps Sloane 1894 LM87
    |--N. retusum Bates 1874 LM87
    |--N. rugulosum Macleay 1869 LM87
    `--N. spenceri Sloane 1897 LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Conopterum

Conopterum sp., copyright Reiner Richter.

Belongs within: Scaritinae.

Conopterum is an Australian genus of scaritine ground beetles, characterised by slender, filiform antennae and bidentate anterior tibiae.

<==Conopterum Chaudoir 1868 [incl. Carenidium Chaudoir 1868] LM87
    |--*C. insigne Chaudoir 1868 LM87
    |--C. aberrans (Sloane 1897) [=Eutoma aberrans] LM87
    |--C. atrum (Sloane 1916) [=Carenidium atrum] LM87
    |--C. bicornutum Macleay 1887 LM87
    |--C. bifurcum (Sloane 1916) [=Carenidium bifurcum] LM87
    |--C. chaudoirii (Macleay 1887) [=Carenidium chaudoirii] LM87
    |--C. damelii (Macleay 1869) [=Carenidium damelii] LM87
    |--C. darlingense (Macleay 1887) [=Carenidium darlingense] LM87
    |--C. frenchi (Sloane 1916) [=Carenidium frenchi] LM87
    |--C. gagatinum (Macleay 1864) [=Carenum gagatinum, *Carenidium gagatinum] LM87
    |--C. leai (Sloane 1897) [=Carenidium leai] LM87
    |--C. longipenne (Sloane 1907) [=Carenidium longipenne] LM87
    |--C. modestum Sloane 1888 LM87
    |--C. mucronatum (Macleay 1866) [=Carenum mucronatum] LM87
    |--C. pertenue (Sloane 1916) [=Carenidium pertenue] LM87
    |--C. purpuratum (Sloane 1905) [=Carenidium purpuratum] LM87
    |--C. pyripenne Sloane 1900 LM87
    |--C. riverinae (Macleay 1865) (see below for synonymy) LM87
    |--C. sapphirinum (Bates 1874) [=Carenidium sapphirinum] LM87
    |--C. septentrionale (Macleay 1887) [=Carenidium septentrionale] LM87
    |--C. spaldingii (Macleay 1878) [=Carenidium spaldingii] LM87
    |--C. superbum (Castelnau 1867) (see below for synonymy) LM87
    `--C. tropicale (Macleay 1887) [=Carenidium tropicale] LM87

Conopterum riverinae (Macleay 1865) [=Carenum riverinae; incl. Ca. amabile Castelnau 1867, Conopterum barnardi Macleay 1887, Co. incornutum Macleay 1887, Monocentrum insigne Chaudoir 1868, Co. littorale Macleay 1887, Co. violaceum Macley 1887] LM87

Conopterum superbum (Castelnau 1867) [=Carenum superbum; incl. Carenidium kreuslerae Macleay 1869, Carenidium lacustre Macleay 1873] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.