Carabus dehaanii, copyright Show Ryu.

Belongs within: Geadephaga.
Contains: Clivina, Pamborus.

The Carabinae are a group of ground beetles that, in the sense used herein, include species with open procoxal and mesocoxal cavities (other classifications expand the 'Carabinae' to cover a much larger, probably non-monophyletic subgroup of the Carabidae). In members of the tribe Carabini, the mandibles lack teeth and have a rugose upper surface, and the clypeus possesses a setigerous puncture at each anterior corner. Members of the Cychrini have smooth and toothed mandibles, and lack prominent setae on the clypeus (Lawrence & Britton 1991).

<==Carabinae [Carabitae] LM87
    |  i. s.: Ceroglossus chilensis (Eschscholtz 1829) B14
    |         Damaster blaptoides Kollar 1836 B14
    |         Acoptolabrus B14
    |         Coptolabrus B14
    |         Clivina LB91
    |--Carabini B90
    |    |--Calosoma Weber 1801 B90, LM87 [=Callisoma Agassiz 1846 LM87]
    |    |    |  i. s.: C. australe Hope 1845 LM87
    |    |    |         C. cyaneoventre Mandl 1955 LM87
    |    |    |         C. frigidum B14
    |    |    |         C. inquisitor L02
    |    |    |         C. scrutator B14
    |    |    |--*C. (Calosoma) sycophanta (Linnaeus 1758) [=Carabus sycophanta] LM87
    |    |    `--C. (Australodrepa Lapouge 1929) LM87
    |    |         |--C. (A.) oceanicum Perroud & Montrouzier 1864 (see below for synonymy) LM87
    |    |         `--C. (A.) schayeri Erichson 1842 [incl. C. curtisi Hope 1845, C. grandipenne Castelnau 1867] LM87
    |    `--Carabus Linnaeus 1758 [Carabina] L58
    |         |  i. s.: C. americanus Linnaeus 1758 L58
    |         |         C. auratus L02
    |         |         C. coriaceus Linnaeus 1758 L58
    |         |         C. crepitans Linnaeus 1758 L58
    |         |         C. cruxminor Linnaeus 1758 L58
    |         |         C. ferrugineus Linnaeus 1758 L58
    |         |         C. glabratus L91
    |         |         C. granulatus Linnaeus 1758 L58 [incl. C. granulatus var. solskyanus C01]
    |         |         C. henningi (see below for synonymy) C01
    |         |         C. hortensis Linnaeus 1758 L58
    |         |         C. inquisitor Linnaeus 1758 L58
    |         |         C. intricatus Linnaeus 1761 VPB11
    |         |         C. krueberi C01
    |         |         C. latus Linnaeus 1758 L58
    |         |         C. leucophthalmus Linnaeus 1758 L58
    |         |         C. lividus Linnaeus 1758 L58
    |         |         C. mannerheimi C01
    |         |         C. maurus C01
    |         |         C. melanocephalus Linnaeus 1758 L58
    |         |         C. menetriesi C01
    |         |         C. multipunctatus Linnaeus 1758 L58
    |         |         C. nemoralis E04
    |         |         C. nitens Linnaeus 1758 L58 [incl. C. nitens var. fennicus C01]
    |         |         C. piceus Linnaeus 1758 L58
    |         |         C. regalis [incl. C. regalis var. cyanicollis, C. regalis var. pasianix] C01
    |         |         C. sibiricus C01
    |         |         C. smaragdinus [incl. C. smaragdinus var. mandshuricus] C01
    |         |         C. spinipes Linnaeus 1758 L58
    |         |         C. vaporariorum Linnaeus 1758 L58
    |         |         C. violaceus Linnaeus 1758 L58
    |         `--C. (Ohomopterus) SS06
    |              |--C. (O.) dehaanii SS06
    |              |--C. (O.) iwawakianus SS06
    |              |--C. (O.) uenoi (Ishikawa 1960) SS06, I92
    |              |--C. (O.) yaconinus SS06
    |              `--C. (O.) yamato SS06
    |--Cychrini [Cychrinae, Pamborinae] LB91
    |    |--Maronetus B90
    |    |--Scaphinotus petersi B90, B14
    |    |--Spaheroderus B90
    |    |--Cychropsis B14
    |    |--Pamborus LM87
    |    `--Cychrus B14
    |         |--C. attenuatus B14
    |         |--C. caraboides Linnaeus 1758 B14
    |         |--C. cordicollis B14
    |         `--C. rostratus L02
    `--Notiophilus Duméril 1806 Z93 [Notiophilini B14]
         |--N. aeneus (Herbst 1806) B14
         |--N. aquaticus C01
         |--N. biguttatus (Fabricius 1779) VPB11
         `--N. quadripunctatus (Dejean 1826) VPB11

Calosoma (Australodrepa) oceanicum Perroud & Montrouzier 1864 [incl. C. (*A.) timorense Chaudoir 1869, C. walkeri Waterhouse 1898] LM87

Carabus henningi [incl. C. henningi var. roeschkei Csiki 1901, C. henningi var. sajanicus Csiki 1901] C01

Hololeius ceylanicus (Nietner 1856) [=Chlaenius ceylanicus; incl. C. nitidulus Dejean 1826, *Hololeius nitidulus, Hololius ornatus Tryon 1890] LM87

*Type species of generic name indicated


[B90] Bell, R. T. 1990. Insecta: Coleoptera Carabidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1053–1092. John Wiley & Sones: New York.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[E04] Emberson, R. M. 2004. A European ground beetle, Harpalus tardus (Panzer) (Coleoptera: Carabidae), in Canterbury, and the predisposition of European carabids to establish in exotic temperate locations. New Zealand Entomologist 27: 39–42.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[L91] Lundqvist, L. 1991. Rearing deutonymphs of Iphidosoma fimetarium (J. Müller), a mesostigmatic mite associated with carabid beetles. In: Schuster, R., & P. W. Murphy (eds) The Acari: Reproduction, development and life-history strategies pp. 447–452. Chapman & Hall: London.

[SS06] Sota, T., & M. Sasabe. 2006. Utility of nuclear allele networks for the analysis of closely related species in the genus Carabus, subgenus Ohomopterus. Systematic Biology 55 (2): 329–344.

[VPB11] Varet, M., J. Pétillon & F. Burel. 2011. Comparative responses of spider and carabid beetle assemblages along an urban-rural boundary gradient. Journal of Arachnology 39 (2): 236–243.

[Z93] Zimmerman, E. C. 1993. Australian Weevils (Coleoptera: Curculionoidea) vol. 3. Nanophyidae, Rhynchophoridae, Erirhinidae, Curculionidae: Amycterinae, literature consulted. CSIRO Australia.

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