Belongs within: Oodini.
Chlaenius is a cosmopolitan genus of often metallic ground beetles with pubescent elytra that have the epipleura interrupted by a twisted fold near the apex. Members of the genus have been divided between over fifty subgenera and monophyly of the genus perhaps warrants investigation.
<==Chlaenius Bonelli 1810 [Chlaeniina] B12
|--C. (Chlaenius) (see below for synonymy) BB01
| |--*C. (C.) festivus (Fabricius 1810) [=Carabus festivus] LM87
| |--C. (C.) aestivus Say 1823 BB01
| |--C. (C.) augustus Newman 1838 BB01
| |--C. (C.) cordicollis Kirby 1837 BB1
| |--C. (C.) cursor BB01
| |--C. (C.) laticollis BB01
| |--C. (C.) platyderus Chaudoir 1856 BB01
| |--C. (C.) ruficauda BB01
| |--C. (C.) sericeus (Forster 1771) BB01
| |--C. (C.) solitarius BB01
| `--C. (C.) viduus Horn 1871 BB01
|--C. (Agostenus Motschulsky 1850) (see below for synonymy) BB01
| |--‘Agostenus’ glaucus BB01
| |--C. (A.) impunctifrons Say 1823 BB01
| |--‘Agostenus’ lithophilus BB01
| |--‘Agostenus’ nemoralis BB01
| |--‘Agostenus’ pennsylvanicus BB01
| |--C. (A.) purpuricollis Randall 1838 BB01
| |--C. (A.) tricolor Dejean 1826 BB01
| |--‘Agostenus’ vafer BB01
| `--‘Agostenus’ variabilipes BB01
|--C. (Anomoglossus Chaudoir 1856) BB01
|--C. (Diapheropsophus) B66
| |--C. (D.) mellyi B66
| `--C. (D.) swinhoei Bates 1866 B66
|--C. (Epomis) B14
| |--C. (E.) circumscriptus (Duftschmid 1812) B14
| `--C. (E.) dejeanii B14
|--C. (Eurydactylus LaFerté Sénectère 1851) [incl. Glyptoderus LaFerté Sénectère 1851] BB01
| |--C. (E.) pimalicus Casey 1914 BB01
| `--C. (E.) tomentosus (Say 1823) BB01
|--C. (Lissauchenius) rufifemoralis OM08
| |--C. r. rufifemoralis OM08
| `--C. r. bimaculatus Dejean 1826 OM08, LM87
|--C. (Pelasmomimus Grundmann 1955) LM87
| `--C. (*P.) greyanus White 1841 [=C. grayianus] LM87
`--C. (Pseudanomoglossus Bell 1960) BB01
`--C. (P.) maxillosus Horn 1876 BB01
Chlaenius incertae sedis:
C. australis Dejean 1831 LM87
C. darlingensis Castelnau 1867 LM87
C. flaviguttatus Macleay 1825 (see below for synonymy) LM87
C. hamifer Chaudoir 1856 [incl. C. bihamatus Chaudoir 1856, C. queenslandicus Sloane 1910] LM87
C. maculiger Castelnau 1867 LM87
C. nigricornis [incl. C. nigricornis var. melanocornis] C01
C. nigripes Macleay 1886 M86
C. ophonoides Fairmaire 1843 (see below for synonymy) LM87
C. planicornis B66
C. rudicollis Chaudoir 1876 [incl. C. bimaculatus Macleay 1864 non Dejean 1826] LM87
C. spoliatus C01
C. subcostatus Macleay 1864 LM87
C. tenellus G89
C. virgulifer C01
Chlaenius flaviguttatus Macleay 1825 [incl. Lissauchenius biguttatus Montrouzier 1860, C. binotatus Dejean 1826, C. guttatus Eschscholtz 1833, C. flaviguttatus immaculipennis Jedlicka 1951, C. maculifer Castelnau 1867, C. punctatus Chaudoir 1856, C. puncticeps Gemminger & Harold 1868] LM87
Chlaenius ophonoides Fairmaire 1843 [incl. C. peregrinus Chaudoir 1856, C. viridescens Gemminger & Harold 1868, C. viridis Montrouzier 1860 non Ménétries 1832] LM87
Chlaenius (Agostenus Motschulsky 1850) [inc. Brachylobus Chaudoir 1876, Chlaeniellus Reitter 1908, Pelasmus Motschulsky 1850] BB01
Chlaenius (Chlaenius) [incl. Callistometus Grundmann 1956, Hemichlaenius Lutschnik 1933 (preoc.), Lithochlaenius Kryzhanovskij 1976] BB01
*Type species of generic name indicated
REFERENCES
[B12] Baehr, M. 2012. A new species of the genus Pheropsophus (Coleoptera: Carabidae: Brachininae) from northern Australia. Records of the Western Australian Museum 27 (1): 62–67.
[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.
[B66] Bates, H. W. 1866. On a collection of Coleoptera from Formosa sent home by R. Swinhoe, Esq., H. B. M. Consul, Formosa. Proceedings of the Zoological Society of London 1866: 339–355.
[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).
[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.
[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.
[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.
[M86] Macleay, W. 1886. The insects of the Fly River, New Guinea, "Coleoptera". Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 136–157.
[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.
Last updated: 3 November 2018.
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