Chlaenius festivus, copyright Siga.

Belongs within: Oodini.

Chlaenius is a cosmopolitan genus of often metallic ground beetles with pubescent elytra that have the epipleura interrupted by a twisted fold near the apex.

<==Chlaenius Bonelli 1810 B12
    |--*C. (Chlaenius) festivus (Fabricius 1810) [=Carabus festivus] LM87
    |--C. (Diapheropsophus) B66
    |    |--C. (D.) mellyi B66
    |    `--C. (D.) swinhoei Bates 1866 B66
    |--C. (Epomis) B14
    |    |--C. (E.) circumscriptus (Duftschmid 1812) B14
    |    `--C. (E.) dejeanii B14
    |--C. (Lissauchenius) rufifemoralis OM08
    |    |--C. r. rufifemoralis OM08
    |    `--C. r. bimaculatus Dejean 1826 OM08, LM87
    `--C. (Pelasmomimus Grundmann 1955) LM87
         `--C. (*P.) greyanus White 1841 [=C. grayianus] LM87

Chlaenius incertae sedis:
  C. australis Dejean 1831 LM87
  C. darlingensis Castelnau 1867 LM87
  C. flaviguttatus Macleay 1825 (see below for synonymy) LM87
  C. hamifer Chaudoir 1856 [incl. C. bihamatus Chaudoir 1856, C. queenslandicus Sloane 1910] LM87
  C. maculiger Castelnau 1867 LM87
  C. nigricornis [incl. C. nigricornis var. melanocornis] C01
  C. nigripes Macleay 1886 M86
  C. ophonoides Fairmaire 1843 (see below for synonymy) LM87
  C. planicornis B66
  C. rudicollis Chaudoir 1876 [incl. C. bimaculatus Macleay 1864 non Dejean 1826] LM87
  C. ruficauda OM08
  C. spoliatus C01
  C. subcostatus Macleay 1864 LM87
  C. tenellus G89
  C. tomentosus BB68
  C. virgulifer C01

Chlaenius flaviguttatus Macleay 1825 [incl. Lissauchenius biguttatus Montrouzier 1860, C. binotatus Dejean 1826, C. guttatus Eschscholtz 1833, C. flaviguttatus immaculipennis Jedlicka 1951, C. maculifer Castelnau 1867, C. punctatus Chaudoir 1856, C. puncticeps Gemminger & Harold 1868] LM87

Chlaenius ophonoides Fairmaire 1843 [incl. C. peregrinus Chaudoir 1856, C. viridescens Gemminger & Harold 1868, C. viridis Montrouzier 1860 non Ménétries 1832] LM87

*Type species of generic name indicated


[B12] Baehr, M. 2012. A new species of the genus Pheropsophus (Coleoptera: Carabidae: Brachininae) from northern Australia. Records of the Western Australian Museum 27 (1): 62–67.

[BB68] Basford, N. L., J. E. Butler, C. A. Leone & F. J. Rohlf. 1968. Immunologic comparisons of selected Coleoptera with analyses of relationships using numerical taxonomic methods. Systematic Zoology 17 (4): 388–406.

[B66] Bates, H. W. 1866. On a collection of Coleoptera from Formosa sent home by R. Swinhoe, Esq., H. B. M. Consul, Formosa. Proceedings of the Zoological Society of London 1866: 339–355.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[M86] Macleay, W. 1886. The insects of the Fly River, New Guinea, "Coleoptera". Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 136–157.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

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