Tricommatidae

Spinopilar moria, copyright BioEspeleo Consultoria Ambiental.

Belongs within: Grassatores.

The Tricommatidae are a group of small short-legged harvestmen found in southern Brazil and Paraguay. The group is characterised by an elongate coxa II which widely surpasses coxa III in ventral view (Kury 2002).

Tricommatidae [Olynthoidae, Triacommatinae, Trichommantinae, Trichommatinae, Tricommatinae]
    |--Caramaschia Kury 2002 K02
    |    `--*C. singularis Kury 2002 K02
    |--Pseudophalangodes Roewer 1912 K02 [=Pseudophalangodus (l. c.) K03]
    |    `--*P. unicolor Roewer 1912 K03
    |--Rivetinus Roewer 1919 R23, K03 [=Ruvetinus (l. c.) K03]
    |    `--*R. minutus Roewer 1919 K03
    |--Neoarminda Özdikmen & Kury 2006 [=Arminda Roewer 1949 non Krauss 1892] OK06
    |    `--*N. colatinae (Soares & Soares 1946) [=Phalangodella colatinae, *Arminda colatinae] OK06
    |--Bebedoura Roewer 1949 K03
    |    `--*B. rugosa Roewer 1949 K03
    |--Bissulla Roewer 1929 K03 [=Bissula (l. c.) KA-Z11]
    |    `--*B. paradoxa Roewer 1929 K03
    |--Bunostigma Mello-Leitão 1935 K03
    |    `--*B. singulare Mello-Leitão 1935 K03, KA-Z11, K03 [=B. singularis K03]
    |--Lanesoares Roewer 1949 K03
    |    `--*L. inermis (Soares 1944) [=Phalangodella inermis] K03
    |--Lussanvira Mello-Leitão 1935 K03
    |    `--*L. marmorata Mello-Leitão 1935 K03
    |--Otuquisa Roewer 1927 K03
    |    `--*O. distincta Roewer 1927 K03
    |--Pararezendesius Soares 1972 K03
    |    `--*P. luridus Soares 1972 K03
    |--Poecilosophus Mello-Leitão 1949 K03, KA-Z11 [=Soaresula Roewer 1949 K03]
    |    `--*P. melloleitaoi (Soares 1945) [=Paratricommatus melloleitaoi, *Soaresula melloleitaoi] K03
    |--Rezendezius Soares 1945 K03
    |    `--*R. lanei Soares 1945 K03
    |--Saladonus Roewer 1927 K03
    |    `--*S. singularis Roewer 1927 K03
    |--Segundolus Roewer 1949 K03
    |    `--*S. insignitus Roewer 1949 K03
    |--Taquara Mello-Leitão 1936 [=Taguara (l. c.); incl. Therezopachylus Soares 1972] K03
    |    `--*T. pilosa Mello-Leitão 1936 [incl. *Therezopachylus fuscus Soares 1972] K03
    |--Berlesecaptus Mello-Leitão 1940 KP-R07, K03
    |    `--*B. convexus Mello-Leitão 1940 K03
    |--Heteromeloleptes Mello-Leitão 1931 K02 (see below for synonymy)
    |    |--*H. padbergi Mello-Leitão 1931 (see below for synonymy) K03
    |    `--H. fasciatus (Mello-Leitão 1932) [=Phalangodella fasciata, *Mellops fasciatus] K03
    |--Tricommatus Roewer 1912 K02
    |    |--*T. brasiliensis Roewer 1912 K02
    |    `--T. fulvus Soares 1966 K03
    |--Bresslauius Mello-Leitão 1935 [incl. Friburguesia Soares 1946, Pinocchio Mello-Leitão 1940] K03
    |    |--*B. hirsutus Mello-Leitão 1935 (see below for synonymy) K03
    |    `--B. debilis (Mello-Leitão 1940) [=*Pinocchio debilis] K03
    |--Machairoscelis Kury 2003 K03
    |    |--*M. armatipes (Soares 1972) [=Corcovadesia armatipes] K03
    |    `--M. singularis (Soares & Soares 1949) [=Paramitraceras singularis] K03
    |--Paratricommatus Piza 1943 K03
    |    |--*P. modestus Piza 1943 K03
    |    `--P. mahnerti Soares & Soares 1985 K03
    |--Tachusina Strand 1942 K03 [=Tachus Sørensen 1932 non Jurine 1807 K03, Tuchusina (l. c.) KA-Z11]
    |    |--*T. keyserlingii (Sørensen 1932) [=*Tachus keyserlingii] K03
    |    `--T. fuscomaculata (Soares & Soares 1947) [=Paramitraceras fuscomaculatus, Timoleon fuscomaculatus] K03
    |--Zalanodius Mello-Leitão 1936 K03
    |    |--*Z. bicornutus 1936 K03
    |    `--Z. hirsutus (Soares & Soares 1970) [=Phalangodella hirsuta] K03
    |--Simonoleptes Soares & Soares 1954 K02
    |    |--*S. latifemur Soares & Soares 1954 K03
    |    |--S. insulanus Soares & Soares 1966 K03
    |    `--S. obstectispiracula Spares & Soares 1954 K03
    |--Arruda Mello-Leitão 1940 K03
    |    |--*A. insignis Mello-Leitão 1940 K03
    |    |--A. mutilata Mello-Leitão 1940 K03
    |    `--A. pectinata Mello-Leitão 1940 K03
    |--Corcovadesia Soares & Soares 1954 K03, KA-Z11 (see below for synonymy)
    |    |--*C. inermis Soares & Soares 1954 [=Liops inermis] K03
    |    |--C. hexabunus (Mello-Leitão 1940) KA-Z11 [=*Liops hexabunus K03]
    |    `--C. venefica (Soares 1966) KA-Z11 [=*Buzioleptes veneficus K03, Liops veneficus K03]
    |--Camarana Mello-Leitão 1935 K02
    |    |--*C. minor Mello-Leitão 1935 K03
    |    |--C. bicoloripes Soares 1974 K03
    |    |--C. flavipalpi Soares 1945 K03
    |    |--C. rousseti Soares & Soares 1946 K03
    |    `--C. unica Soares 1944 K03
    |--Spinopilar Mello-Leitão 1940 KP-G08 (see below for synonymy)
    |    |--*S. armatus Mello-Leitão 1940 KP-G08
    |    |--S. anomalis (Sørensen 1932) KP-G08 (see below for synonymy)
    |    |--S. apiacaensis Kury 1992 KP-G08
    |    |--S. friburguensis (Soares 1946) KP-G08 [=Saladonus friburguensis K03]
    |    `--S. moria Kury & Pérez-González 2008 KP-G08
    `--Pseudopachylus Roewer 1912 (see below for synonymy) K03
         |--*P. longipes Roewer 1912 (see below for synonymy) K03
         |--P. alticola (Soares 1945) [=Olynthus alticola] K06
         |--P. cocaiensis (Soares 1972) K06 [=Paramitraceras cocaiensis K03]
         |--P. eximius (Mello-Leitão 1936) K06 (see below for synonymy)
         |--P. martensi Kury 2006 K03
         `--P. nigripes (Mello-Leitão 1932) K06 [=Tricommatus nigripes K03; incl. P. lissonotus Mello-Leitão 1938 K03]

*Bresslauius hirsutus Mello-Leitão 1935 [=Pucrolia hirsuta; incl. *Friburguesia hirsuta Soares 1946 non Bresslauius hirsutus Mello-Leitão 1935] K03

Corcovadesia Soares & Soares 1954 K03, KA-Z11 [incl. Buzioleptes Soares 1966 K03, Liops Mello-Leitão 1940 nec Fieber 1870 nec Gidley 1906 KA-Z11]

Heteromeloleptes Mello-Leitão 1931 K02 [incl. Derzius Roewer 1949 K03, Melloceras Roewer 1949 K03, Mellops Roewer 1949 K03]

*Heteromeloleptes padbergi Mello-Leitão 1931 [incl. Pseudopachylus anomalus Mello-Leitão 1935, *Derzius anomalus, Pseudophalangodes bimaculatus Mello-Leitão 1932, *Melloceras bimaculatus, Phalangodella jandirae Soares & Soares 1970] K03

Pseudopachylus Roewer 1912 [=Pseudophchylus (l. c.); incl. Caporiacoius Mello-Leitão 1936, Monticola Soares 1944 nec Boie 1822 nec Nalivkin 1930, Monticolina Soares 1944] K03

Pseudopachylus eximius (Mello-Leitão 1936) K06 [=*Caporiacoius eximius K03; incl. C. fallax Soares 1944 K03, Pseudopachylus fallax K03]

*Pseudopachylus longipes Roewer 1912 [incl. *Monticola acutinasua Soares 1944, *Monticolina acutinasua, Pseudopachylus acutinasua] K03

Spinopilar Mello-Leitão 1940 KP-G08 [incl. Olynthus Sørensen 1932 nec Hübner 1819 nec Haeckel 1869 K03, Soerensenibunus Strand 1942 K03, Soerensenolynthus Soares & Soares 1947 K03]

Spinopilar anomalis (Sørensen 1932) KP-G08 [=*Olynthus anomalis K03, O. anomalus K03, *Soerensenibunus anomalis K03, *Soerensenolynthus anomalis K03]

*Type species of generic name indicated

REFERENCES

[K02] Kury, A. B. 2002. A new genus of Tricommatinae from eastern Brazil (Opiliones Laniatores Gonyleptidae). Tropical Zoology 15: 209–218.

[K03] Kury, A. B. 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Ibérica de Aracnología, volumen especial monográfico 1: 1–337.

[K06] Kury, A. B. 2006. A new Tricommatinae from the montane savanna of São Paulo (Opiliones: Laniatores: Gonyleptidae). Zootaxa 1325: 211–217.

[KA-Z11] Kury, A. B., & M. A. Alonso-Zarazaga. 2011. Addenda and corrigenda to the "Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones)". Zootaxa 3034: 47–68.

[KP-G08] Kury, A. B., & A. Pérez-González. 2008. The first cave-dwelling Spinopilar Mello-Leitão 1940 (Opiliones Gonyleptidae Tricommatinae), described from a Brazilian cave. Tropical Zoology 21: 259–267.

[KP-R07] Kury, A. B., & R. Pinto-da-Rocha. 2007. Gonyleptidae Sundevall, 1833. In: Pinto-da-Rocha, R., G. Machado & G. Giribet (eds) Harvestmen: The Biology of Opiliones pp. 196–203. Harvard University Press: Cambridge (Massachusetts).

[OK06] Özdikmen, H., & A. B. Kury. 2006. Three homonymous generic names in Araneae and Opiliones. Journal of Arachnology 34 (1): 279–280.

[R23] Roewer, C.-F. 1923. Die Weberknechte der Erde: Systematische Bearbeitung der bisher bekannnten Opiliones. Gustav Fischer: Jena.