Mitobatinae

Neoancistrotus gracilis, copyright Osvaldo Villarreal Manzanilla.

Belongs within: Gonyleptidae.

The Mitobatinae are a group of gonyleptid harvestmen with a usually elongate, unarmed femur IV in the male (Bragagnolo & Pinto-da-Rocha 2012).

Characters (from Bragagnolo & Pinto-da-Rocha 2012): Anterior margin of prosoma with frontal hump present and unarmed. Armature of ocularium usually with two spiniform apophyses. Armature of scutal areas variable, area III usually armed with pair of spiniform apophyses. Usually four areas on dorsal scutum; sometimes areas III and IV fused. Posterior margin may be armed with pair of tubercles. Free tergites unarmed or with median tubercles. Chelicera of similar size in both sexes. Femur of pedipalps only with short and thick ventral apical setae, patella and tibia unarmed. Coxa IV dorsally visible in entire extension in both sexes, with dorsoapical apophysis that can be robust and bifid in males (sexual dimorphism) or extremely reduced. Male femur IV straight and usually much elongated, usually more than four times scutal size, rarely about three times, and usually unarmed. Penis with ventral plate well defined, usually pyriform (sometimes rectangular), with setae on distal margin. Ventral process present; dorsal process absent. Stylus elongate, usually straight.

<==Mitobatinae
    |--Metamitobates Roewer 1913 (see below for synonymy) K03
    |    `--M. squalidus (Perty 1832) DG09 (see below for synonymy)
    |--Despirus Roewer 1929 [incl. Ibarroides Mello-Leitão 1932, Segadasius Mello-Leitão 1949] K03
    |    `--*D. parvulus (Roewer 1927) (see below for synonymy) K03
    |--Dyscocyrtoides Mello-Leitão 1923 KA-Z11 [=Discocyrtoides K03; incl. Mitobatoides Mello-Leitão 1927 K03]
    |    `--D. nigricans (Mello-Leitão 1922) (see below for synonymy) K03
    |--Mitobatula Roewer 1931 K03
    |    `--*M. castanea Roewer 1931 K03
    |--Encheiridium Kury 2003 K03
    |    |--*E. ruschii (Mello-Leitão 1942) [=Despirus ruschii, Discocyrtoides ruschii] K03
    |    `--E. montanum (Mello-Leitão 1941) [=Despirus montanus, Discocyrtoides montanus] K03
    |--Ischnotherus Kury 1991 K03
    |    |--*I. tenebrosus Kury 1991 K03
    |    `--I. pardus (Kury 1989) [=Discocyrtoides pardus] K03
    |--Ruschia Mello-Leitão 1940 K03
    |    |--*R. vellutina Mello-Leitão 1940 [=Discocyrtoides vellutinus] K03
    |    `--R. maculata (Soares 1974) [=Discocyrtoides maculatus] K03
    |--Mitobates Sundevall 1833 K03 (see below for synonymy)
    |    |--*M. triangulus Sundevall 1833 K03 (see below for synonymy)
    |    |--M. elegantulus (Mello-Leitão 1940) [=Neoancistrotus elegantulus, Ancistrotellus elegantulus] K03
    |    |--M. inermis Kury 1989 K03
    |    `--M. pulcher (Mello-Leitão 1932) [=Despiroides pulcher, Discocyrtoides pulcher] K03
    |--Longiperna Roewer 1929 K03
    |    |--*L. concolor (Mello-Leitão 1923) [=Discocyrtoides concolor] K03
    |    |--L. areolata (Soares 1944) [=Discocyrtoides areolatus] K03
    |    |--L. cancellata (Roewer 1913) DG09 [=Neomitobates cancellatus K03, Metamitobates cancellatus K03]
    |    |--L. coxalis (Roewer 1943) [=Mitobatula coxalis] K03
    |    |--L. curitibana Kury 2003 [=L. cancellata Roewer 1943 non Neomitobates cancellatus Roewer 1913] K03
    |    |--L. heliaca (Soares 1942) [=Despirus heliacus, Discocyrtoides heliacus] K03
    |    |--L. insperata (Soares & Soares 1947) [=Ancistrotellus insperatus] K03
    |    |--L. paranensis (Soares & Soares 1947) [=Discocyrtoides paranensis] K03
    |    `--L. zonata (Mello-Leitão 1935) [=Despirus zonatus, Discocyrtoides zonatus] K03
    |--Promitobates Roewer 1913 K03 (see below for synonymy)
    |    |--*P. hexacanthus (Koch 1839) K03, DG09 [=Ancistrotus hexacanthus K03, Goniosoma hexacanthum K03]
    |    |--P. bellus (Soares 1945) [=Ancistrotellus bellus] K03
    |    |--P. difficilis (Mello-Leitão 1931) [=*Batomites difficilis] K03
    |    |--P. granulosissimus Mello-Leitão 1932 [incl. P. decoratus Mello-Leitão 1932] K03
    |    |--P. hatschbachi Soares 1945 K03
    |    |--P. hauseri (Šilhavý 1979) [=Ancistrotellus hauseri] K03
    |    |--P. margaritatus Roewer 1931 K03
    |    |--P. mendax Soares 1945 K03
    |    |--P. ornatus (Mello-Leitão 1922) (see below for synonymy) K03
    |    `--P. viridigranulatus (Soares & Soares 1946) DG09 [=Ancistrotellus viridigranulatus K03]
    `--Neoancistrotus Mello-Leitão 1927 [incl. Neomitobatoides Mello-Leitão 1933] K03
         |--*N. thiacanthus Mello-Leitão 1927 [=Ancistrotellus thiacanthus] K03
         |--N. bifurcatus (Koch 1839) DG09 DG09 (see below for synonymy)
         |--N. bipustulatus (Mello-Leitão 1940) [=Ancistrotellus bipustulatus] K03
         |--N. bristowei (Mello-Leitão 1924) [=Ancistrotus bristowei, Ancistrotellus bristowei] K03
         |--N. dubius (Mello-Leitão 1932) [=Ancistrotellus dubius] K03
         |--N. gracilis (Roewer 1917) [=Ancistrotus gracilis, Ancistrotellus gracilis] K03
         |--N. guapimirim (Soares & Soares 1946) [=Ancistrotellus guapimirim] K03
         |--N. intermedius (Mello-Leitão 1936) [=Ancistrotellus intermedius] K03
         |--N. maculipalpi (Soares & Soares 1948) (see below for synonymy) K03
         |--N. nigroides (Soares & Soares 1947) [=Ancistrotellus nigroides] K03
         |--N. obscurus (Mello-Leitão 1931) (see below for synonymy) K03
         `--N. rosai (Mello-Leitão 1933) [=*Neomitobatoides rosae, Ancistrotellus rosae, Neoancistrotus rosae] K03

*Despirus parvulus (Roewer 1927) [=Discocyrtus parvulus, Discocyrtoides parvulus; incl. Discocyrtoides granulosus Mello-Leitão 1936, *Ibarroides minutus Mello-Leitão 1932, Discocyrtoides minutus, *Segadasius monticola Mello-Leitão 1949] K03

Dyscocyrtoides nigricans (Mello-Leitão 1922) [=Ancistrotus nigricans, *Mitobatoides nigrcans, M. nigrigans (l. c.); incl. Despirus piracicabensis Piza 1938, De. ustus Mello-Leitão 1937, *Discocyrtoides violaceus Mello-Leitão 1923, Di. ypirangae Mello-Leitão 1923] K03

Metamitobates Roewer 1913 [incl. Ancistrotellus Roewer 1923, Ancistrotus Koch 1839 non Audinet-Serville 1832, Ancystrotus (l. c.), Neobourguyia Mello-Leitão 1940, Neomitobates Roewer 1913] K03

Metamitobates squalidus (Perty 1832) DG09 [=Goniosoma squalidum K03, *Ancistrotellus squalidus K03, *Ancistrotus squalidus K03; incl. *Metamitobates genusulphureus Roewer 1913 K03, Metamit. genusulfureas (l. c.) K03, Metamit. genusulfureus (l. c.) K03, *Neomitobates gracilipes Roewer 1913 K03, Metamit. gracilipes K03, *Neobourguyia rosai Mello-Leitão 1940 K03, Despiroides rosai K03, Discocyrtoides rosai K03, Ancistrotus urceolaris Bertkau 1880 K03, Neomitobates venustus Mello-Leitão 1933 K03, Metamibotates (l. c.) venustus K03]

Mitobates Sundevall 1833 K03 [incl. Despiroides Mello-Leitão 1932 K03, Dolichoscelis Hope 1836 K03, KA-Z11, Metaroeweria Mello-Leitão 1936 K03]

*Mitobates triangulus Sundevall 1833 K03 [incl. Goniosoma conspersum Perty 1833 K03, Ancistrotus conspersus K03, Mitobates conspersus DG09, *Dolichoscelis haworthii Hope 1837 K03, *Metaroeweria insignis Mello-Leitão 1936 K03, Neoancistrotus nigromaculatus Mello-Leitão 1931 K03, Ancistrotellus nigromaculatus K03, Mitobates stygnoides Mello-Leitão 1923 K03, *Despiroides xanthophthalmus Mello-Leitão 1932 K03, Discocyrtoides xanthophthalmus K03]

Neoancistrotus bifurcatus (Koch 1839) DG09 [=Ancistrotus bifurcatus K03, Ancistrotellus bifurcatus K03, Dolichoscelis bifurcatus K03, Goniosoma bifurcatum K03; incl. Ancistrotellus anomalus Mello-Leitão 1935 K03, Neoancistrotus anomalus K03, Prosancistrotus communis Mello-Leitão in Soares 1945 (n. n.) K03, Ancistrotellus perlatus Mello-Leitão 1932 K03, N. perlatus K03, Ancistrotellus tijucae Soares 1944 K03, N. tijucae K03]

Neoancistrotus maculipalpi (Soares & Soares 1948) [=Ancistrotellus maculipalpi; incl. A. gracilioides Mello-Leitão in Soares 1945 (n. n.), A. luteimanu Mello-Leitão in Soares 1945 (n. n.)] K03

Neoancistrotus obscurus (Mello-Leitão 1931) [=Ancistrotellus obscurus; incl. A. melacanthus Mello-Leitão 1932, Neoancistrotus melacanthus] K03

Promitobates Roewer 1913 K03 [incl. Batomites Mello-Leitão 1931 (nom. inv.) K03, KA-Z11, Leonardosia Mello-Leitão 1935 K03, Promitobatoides Mello-Leitão 1927 K03]

Promitobates ornatus (Mello-Leitão 1922) [=Neomitobates ornatus, *Promitobatoides ornatus; incl. Batomites heteracanthus Mello-Leitão 1935, Promitobates heteracanthus, B. intermedius Mello-Leitão 1935, Neoancistrotus nigripes Mello-Leitão 1935, Ancistrotellus nigripes, *Leonardosia nitida Mello-Leitão 1935, Promitobates nitidus, Batomites nitidus Mello-Leitão 1940 non Leonardosia nitida Mello-Leitão 1935, B. spitzi Mello-Leitão 1931] K03

*Type species of generic name indicated

REFERENCES

Bragagnolo, C., & R. Pinto-da-Rocha. 2012. Systematic review of Promitobates Roewer, 1913 and cladistic analysis of Mitobatinae Simon, 1879 (Arachnida: Opiliones: Gonyleptidae). Zootaxa 3308: 1–48.

[DG09] DaSilva, M. B., & P. Gnaspini. 2009. A systematic revision of Goniosomatinae (Arachnida: Opiliones: Gonyleptidae), with a cladistic analysis and biogeographical notes. Invertebrate Systematics 23: 530–624.

[K03] Kury, A. B. 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Ibérica de Aracnología, volumen especial monográfico 1: 1–337.

[KA-Z11] Kury, A. B., & M. A. Alonso-Zarazaga. 2011. Addenda and corrigenda to the "Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones)". Zootaxa 3034: 47–68.