Pea urchin Echinocyamus pusillus, copyright Marine Ecological Surveys Ltd.

Belongs within: Clypeasteroida.

The Fibulariidae are a group of echinoids known in temperate and tropical regions from the Upper Cretaceous to the present (Durham 1966).

Characters (from Durham 1966): Shape variable; petals variable, indistinct or simple, open; pore pairs not conjugate, pores rounded; food grooves absent or indistinct; primordial plates simple; internal supports absent or radial partitions only.

<==Fibulariidae [Fibularina, Fistularina] D66
    |--Cyamidia Lambert & Thiéry 1914 D66
    |    `--*C. nummulitica (Duncan & Sladen 1884) [=Echinocyamus nummulitica] M89
    |--Fibulaster Lambert & Thiéry 1914 [=Crustulina Pomel 1883 non Menge 1868] D66
    |    `--*F. michelini (Cotteau 1861) M89 [=Sismondia michelini M89, *Crustulina michelini D66]
    |--Lenicyamidia Brunnschweiler 1962 D66
    |    `--*L. compta Brunnschweiler 1962 M89
    |--Mortonia Gray 1852 D66
    |    `--*M. australis (Desmoulins 1837) [=Fibularia australis] M89
    |--Thagastea Pomel 1888 [=Thegaster (l. c.)] D66
    |    `--*T. wetterlei Pomel 1888 M89
    |--Fibularia Lamarck 1816 M89
    |    |--*F. ovulum Lamarck 1816 M89
    |    `--F. volva Agassiz 1846 BM09
    |--Fibulariella Mortensen 1948 D66
    |    |--*F. acuta (Yoshiwara 1898) [=Fibularia acuta] M89
    |    `--F. oblonga M89
    |--Tarphypygus Arnold & Clark 1927 D66
    |    |--*T. ellipticus Arnold & Clark 1927 M89
    |    `--T. clarki M89
    `--Echinocyamus van Phelsum 1774 MG-H11, M89 (see below for synonymy)
         |--*E. pusillus Müller 1776 M89 [=Spatagus pusillus D66]
         |--E. elongatus D66
         |--E. platytatus Clark 1914 MG-H11
         `--*Anaster’ studeri Sismonda 1841 D66

Echinocyamus van Phelsum 1774 MG-H11, M89 [incl. Echinocentrotus (l. c.) D66; incl. Anaster Sismonda 1841 D66, Fibularia Lambert 1891 (nom. rej.) non Lamarck 1816 D66]

*Type species of generic name indicated


[BM09] Bryce, C., & L. Marsh. 2009. Echinodermata (Asteroidea, Echinoidea and Holothuroidea) of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, Western Australia. Records of the Western Australian Museum Supplement 77: 209–220.

[D66] Durham, J. D. 1966. Clypeasteroids. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt U. Echinodermata 3 vol. 2 pp. U450–U491. The Geological Society of America, Inc., and The University of Kansas Press.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[M89] Mooi, R. 1989. Living and fossil genera of the Clypeasteroida (Echinoidea: Echinodermata): an illustrated key and annotated checklist. Smithsonian Contributions to Zoology 488: 1–51.

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