Various external views of holotype (1–5) and paratype (6–9) of Quasiumbella carnarvonensis, from Edgell (2003).

Belongs within: Streptophyta.

The Umbellinaceae are a group of fossil protists (currently thought to be early streptophyte algae) known from the Devonian and Lower Carboniferous (Edgell 2003).

Characters (from Edgell 2003): Subconical to spherical gyrogonites, with simple or slightly rimmed aperture and basal pore; wall two-layered with thin dark internal layer and thick outer layer of radial fibres; mostly with smooth outer surface, occasionally with vertical ridges or spiny surface ornamentation.

<==Umbellinaceae [Umbellinae, Umbellinaphyceae, Umbellininae]
    |--Costatumbella E03
    |--Spinumbella Platonov 1974 E03
    |    `--S. spinifera Platonov 1974 E03
    |--Eolagena Lipina 1959 LT64
    |    `--*E. minuta Lipina 1959 LT64
    |--Eovolutina Antropov 1950 LT64
    |    `--*E. elementa Antropov 1940 LT64
    |--Umbellina Loeblich & Tappan 1961 [=Umbella Maslov in Bykova 1955 nec d’Orbigny 1841 nec Scudder 1882] E03
    |    |--*U. bella (Maslov) Loeblich & Tappan 1974 (see below for synonymy) E03
    |    `--‘Umbella’ minima E03
    `--Quasiumbella Poyarkov 1965 [incl. Protoumbella Mamet 1970] E03
         |--*Q. rotunda (Bykova) Poyarkov 1965 [=Umbella rotunda Bykova 1955; incl. U. radiata Konoplina 1959] E03
         |--Q. atipica E03
         |--Q. bulla Edgell 2003 E03
         |--Q. carnarvonensis Edgell 2003 E03
         |--Q. gneudnaensis Edgell 2003 E03
         |--Q. langoora Edgell 2003 E03
         |--Q. pararotunda E03
         |--Q. polonica [=Umbellina polonica Ozonkowa 1962] E03
         `--Q. saccamminiformis (see below for synonymy) E03

Quasiumbella saccamminiformis [=Umbella saccamminiformis Bykova 1955, *Protoumbella saccamminiformis (Bykova) Mamet 1970] E03

*Umbellina bella (Maslov) Loeblich & Tappan 1974 [=*Umbella bella Maslov in Bykova 1955; incl. Umbellina santacrucensis] E03

*Type species of generic name indicated


[E03] Edgell, H. S. 2003. Upper Devonian Charophyta of Western Australia. Micropaleontology 49 (4): 359–374.

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

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