Cibicides lobatulus, copyright Marek Zajaczkowski & Patrycja Jernas.

Belongs within: Rotaliida.

The Cibicididae are a group of spiral foraminifera known from the Cretaceous to the present. In members of the subfamily Planulininae, the test is free and has a single aperture; in Cibicidinae, the test is attached by its spiral side and some forms have multiple apertures (Loeblich & Tappan 1964b).

Characters (from Loeblich & Tappan 1964b): Test free or attached, trochospiral to nearly planispiral, or later spreading, irregular or cyclical; wall coarsely perforate, radial in structure, septa double (bilamellar); aperture interiomarginal, may extend onto spiral side, and peripheral supplementary apertures may occur.

    |--Planulininae LT64b
    |    |--Cibicidina Bandy 1949 LT64b
    |    |    `--*C. walli Bandy 1949 LT64b
    |    |--Hyalinea Hofker 1951 [=Hofkerinella Bermúdez 1952] LT64b
    |    |    `--*H. balthica (Schröter 1783) [=Nautilus balthicus, *Hofkerinella balthica] LT64b
    |    `--Planulina d’Orbigny 1826 C40
    |         |--*P. ariminensis d’Orbigny 1826 C40
    |         |--P. ornata C40
    |         |--P. renzi H90
    |         `--P. wuellerstorfi B49
    `--Cibicidinae LT64a
         |  i. s.: Stichocibicides Cushman & Bermúdez 1936 C40
         |           `--*S. cubensis Cushman & Bermúdez 1936 C40
         |         Caribeanella Bermúdez 1952 [incl. Oinomikadoina Matsunaga 1954, Pseudocibicidoides Ujiié 1956] LT64b
         |           |--*C. polystoma Bermúdez 1952 LT64b
         |           |--C. katasensis (Ujiié 1956) [=*Pseudocibicidoides katasensis] LT64b
         |           `--C. ogiensis (Matsunaga 1954) [=*Oinomikadoina ogiensis] LT64b
         |         Cycloloculina Heron-Allen & Earland 1908 [=Cycloloceilina (l. c.)] LT64b
         |           |--*C. annulata Heron-Allen & Earland 1908 LT64b
         |           `--C. miocenica C40
         |         Planorbulinoides Cushman 1928 LT64b
         |           `--*P. retinaculata (Parker & Jones in Carpenter, Parker & Jones 1862) (see below for synonymy) LT64b
         |--Dyocibicides Cushman & Valentine 1930 C40 [incl. Rectocibicidella McLean 1956 LT64b]
         |    |--*D. biserialis Cushman & Valentine 1930 C40
         |    |--D. robertsi (McLean 1956) [=*Rectocibicidella robertsi] LT64b
         |    `--D. variabilis LJ98
         |--+--Annulocibicides Cushman & Ponton 1932 C40
         |  |    `--*A. projectus Cushman & Ponton 1932 C40
         |  `--Rectocibicides Cushman & Ponton 1932 C40
         |       `--*R. miocenicus Cushman & Ponton 1932 C40
         |--+--Cibicidella Cushman 1927 C40
         |  |    `--*C. variabilis (d’Orbigny 1826) [=Truncatulina variabilis] LT64b
         |  `--Cyclocibicides Cushman 1927 C40
         |       `--*C. vermiculata (d’Orbigny 1826) [=Planorbulina vermiculata] LT64b
         `--Cibicides Monfort 1808 C40, G86 (see below for synonymy)
              |--*C. refulgens de Montfort 1808 [=*Truncatulina refulgens] LT64b
              |--C. advenum M62
              |--*Craterella’ albescens Dons 1942 [=*Crateriola albescens] LT64b
              |--‘Truncatulina’ boueana F71
              |--C. collinsi B49
              |--*Polyxenes’ cribratus de Montfort 1808 LT64b
              |--*Soldanina’ exagona Costa 1856 LT64b
              |--C. kullenbergi H90
              |--C. lobatulus C40 [=Truncatulina lobulata BK77, Lobatula lobatula M08]
              |--C. mediocris H90
              |--C. neoperforatus H90
              |--C. notocenicus H90
              |--C. novozelandicus H90
              |--C. perforatus H90
              |--C. pseudolobatus S05
              |--*Storilus’ radiatus de Montfort 1808 LT64b
              |--C. strelkovi Mikhalevich 1976 G86
              |--C. temperata H90
              |--‘Truncatulina’ ungeriana H04
              |--C. variabilis LT64a
              `--*Lobatula’ vulgaris Fleming 1828 LT64b

Nomen nudum: Cibicides molestus Battey 1949 B49

Cibicides Monfort 1808 C40, G86 [=Cymbicides (l. c.) LT64b, Truncatulina d’Orbigny 1826 LT64b; incl. Aristeropora Ehrenberg 1858 C40, Craterella Dons 1942 nec Schrammen 1901 nec Kofoid & Campbell 1929 LT64b, Crateriola Strand 1943 LT64b, Lobatula Fleming 1828 C40, Polyxenes de Montfort 1808 LT64b, Soldanina Costa 1856 LT64b, Storilus de Montfort 1808 LT64b]

*Planorbulinoides retinaculata (Parker & Jones in Carpenter, Parker & Jones 1862) [=Planorbulina retinaculata] LT64b

*Type species of generic name indicated


[BK77] Barel, C. D. N., & P. G. N. Kramers. 1977. A survey of the echinoderm associates of the north-east Atlantic area. Zoologische Verhandelingen 156: 1–159.

[B49] Battey, M. H. 1949. The geology of the Tuakau-Mercer area, Auckland. Transactions and Proceedings of the Royal Society of New Zealand 77 (3): 429–455.

[C40] Cushman, J. A. 1940. Foraminifera: Their classification and economic use 3rd ed. Harvard University Press: Cambridge (Massachusetts).

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[G86] Golemansky, V. G. 1986. Rhizopoda: Foraminiferida. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 17–20. E. J. Brill/Dr. W. Backhuys: Leiden.

[H04] Haeckel, E. 1899–1904. Kunstformen der Natur. Bibliographisches Institut: Leipzig und Wien.

[H90] Hayward, B. W. 1990. Use of foraminiferal data in analysis of Taranaki Basin, New Zealand. Journal of Foraminiferal Research 20: 71–83.

[LJ98] Li, Q., N. P. James, B. McGowran, Y. Bone & J. Cann. 1998. Synergetic influence of water masses and Kangaroo Island barrier on foraminiferal distribution, Lincoln and Lacepede shelves, South Australia: a synthesis. Alcheringa 22 (2): 153–176.

[LT64a] Loeblich, A. R., Jr & H. Tappan. 1964a. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

[LT64b] Loeblich, A. R., Jr & H. Tappan. 1964b. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 2. The Geological Society of America and The University of Kansas Press.

[M08] McMillan, I. K. 2008. Aragonitic-walled benthic foraminifera (Epistomina) in the Cretaceous 'mudbelt' off southern Africa, and postmortem cross-shelf transport of tests. African Natural History 4: 17–24.

[M62] Monniot, F. 1962. Recherches sur les graviers a Amphioxus de la région de Banyuls-sur-Mer. Vie et Milieu 13: 231–322.

[S05] Semeniuk, T. A. 2005. Fossil foraminiferal assemblages from Pleistocene seagrass-bank deposits of the southern Perth Basin, Western Australia, and their palaeotemperature implications. Journal of the Royal Society of Western Australia 88 (4): 177–190.

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