Longitudinal section of Pachyphloia ovata under plane-polarised light, from Groves et al. (2004).

Belongs within: Lagenida.

The Geinitzinidae are a group of uniserial foraminifera known from the Late Carboniferous and Permian, with an ancestrally atelo-monolamellar (i.e. lacking secondary lamellarity), hyaline-radial wall structure. In the derived genus Pachyphloia, the wall becomes plesio-monolamellar, with secondary lamellae enveloping some but not all previous chambers (Groves et al. 2003).

    |--Vervilleina Groves in Groves & Boardman 1999 GAR03
    |    `--V. bradyi (Spandel 1901) GAR03
    |--Nodosinelloides Mamet & Pinard 1992 GAR03
    |    |--N. netschajewi (Cherdyntsev 1914) GAR03
    |    |--N. pinardae Groves & Wahlmann 1997 GRA04
    |    `--N. ronda (Lipina 1949) GAR03
    `--+--Geinitzina Spandel 1901 GAR03 [=Geinitzella Spandel 1898 non Waagen & Wentzel 1866 LT64]
       |    |--‘Textularia’ jonesi Brady 1876 (see below for synonymy) LT64
       |    |--G. ciscoensis C40
       |    |--G. postcarbonica Spandel 1901 GAR03 [incl. G. triangularis F71]
       |    `--G. tcherdynzevi Miklukho-Maclay 1954 W76
       `--+--Howchinella Palmieri in Foster et al. 1985 GAR03
          |    |--*H. woodwardi (Howchin 1895) [=Frondicularia woodwardi] GAR03
          |    `--‘Geinitzina’ caseyi Crespin 1958 GAR03
          `--Pachyphloia Lange 1925 GAR03 (see below for synonymy)
               |--*P. ovata Lange 1925 LT64
               |--P. asymmetrica (Miklukho-Maklay 1954) [=*Parapachyphloia asymmetrica] LT64
               |--P. cukurkoyi Sellier de Civrieux & Dessauvagie 1965 GRA04
               |--P. depressa (Miklukho-Maklay 1954) [=*Parageinitzina depressa] LT64
               |--P. gefoensis (Miklukho-Maklay 1953) [=*Parapermodiscus gefoensis] LT64
               |--P. magna (Miklukho-Maklay 1954) [=*Pseudogeinitzina magna] LT64
               `--P. pigmobesa Wang 1976 W76

Pachyphloia Lange 1925 GAR03 [incl. Parageinitzina Miklukho-Maklay 1954 LT64, Parapachyphloia Miklukho-Maklay 1954 LT64, Parapermodiscus Miklukho-Maklay 1953 LT64, Pseudogeinitzina Miklukho-Maklay 1954 LT64; Pachyphloiidae]

‘Textularia’ jonesi Brady 1876 [=Lunucammina jonesi; incl. Textularia cuneiformis Jones in King 1850 non d’Orbigny 1876, *Geinitzella cuneiformis, *Geinitzina cuneiformis] LT64

*Type species of generic name indicated


[C40] Cushman, J. A. 1940. Foraminifera: Their classification and economic use (3rd ed.) Harvard University Press: Cambridge (Massachusetts).

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1-167.

[GAR03] Groves, J. R., D. Altiner & R. Rettori. 2003. Origin and early evolutionary radiation of the order Lagenida (Foraminifera). Journal of Paleontology 77 (5): 831-843.

[GRA04] Groves, J. R., R. Rettori & D. Altiner. 2004. Wall structures in selected Paleozoic lagenide Foraminifera. Journal of Paleontology 78: 245-256.

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt. C. Protista 2, vol. 1. The Geological Society of America and The University of Kansas Press.

[W76] Wang K. 1976. The foraminifera from the Changhsing Formation in western Guizhou. Acta Palaeontologica Sinica 15 (2): 187-195.

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