Cyprois marginata, copyright Zoologische Staatssammlung Muenchen.

Belongs within: Ostracoda.
Contains: Suchonellina, Cyprididae, Candonidae, Ilyocyprididae.

The Cypridocopina are a group of ostracods in which three pairs of thoracic legs are present, with the first pair modified to function as a mouthpart and the third pair is bent dorsally and used as a cleaning organ, and the body bears a terminal furca (Benson et al. 1961).

Characters (from Benson et al. 1961, as Cypridacea): Carapace variable in size, shape, and surface ornamentation; calcareous or corneous; mid-ventral margin curved inward. Hinge of ridge-and-groove type or rabbeted; muscle scars a median group of spots, typically not radially or linearly arranged, with additional more anteroventral spots in most genera; inner lamellae present; line of concrescence and inner margin typically separated.

    |--Pontocyprididae (see below for synonymy) MD01
    |    |--Liasina lanceolata (Apostolecsu 1959) WSB93
    |    |--Argilloecia Sars 1866 [=Argillaecia Brady 1870] BB61
    |    |    `--*A. cylindrica Sars 1866 BB61
    |    |--Bakunella Schneider in Mandelstam et al. 1958 BB61
    |    |    `--*B. dorsoarcuata (Zalanyi 1929) [=Pontocypris dorsoarcuata] BB61
    |    |--Clinocypris Mandelstam 1956 BB61
    |    |    `--*C. scolia Mandelstam 1956 BB61
    |    |--Pontocypria Müller 1894 BB61
    |    |    `--*P. spinosa Müller 1894 BB61
    |    |--Propontocypris Sylvester-Bradley 1947 BB61
    |    |    `--*P. trigonella (Sars 1866) [=Pontocypris trigonella] BB61
    |    |--Subulacypris Schneider in Mandelstam et al. 1957 BB61
    |    |    `--*S. subtilis Schneider in Mandelstam et al. 1957 BB61
    |    |--Bosquetia Brady, Crosskey & Robertson 1874 M61, BB61
    |    |    `--*B. robusta Brady, Crosskey & Robertson 1874 BB61
    |    `--Pontocypris Sars 1866 [incl. Cytheropsis McCoy 1849 (n. d.), Erythrocypris Müller 1894] BB61
    |         |--P. mytiloides (Norman 1862) (see below for synonymy) BB61
    |         |--*Cytheropsis’ aldensis M’Coy 1851 BB61
    |         `--P. mawii Jones 1887 [incl. P. mawii var. proxima Jones 1889] C01
    `--Cypridoidea [Cypracea, Cypridacea] M03
         |  i. s.: Camdenidea Swain 1953 BB61
         |           `--*C. camdenensis Swain 1953 BB61
         |         Carbonitidae WSB93
         |           |--Carbonita Strand 1928 [=Carbonia Jones 1870 non Robineau-Desvoidy 1863] BB61
         |           |    |--*C. agnes (Jones 1870) [=*Carbonia agnes] BB61
         |           |    `--C. fabulina G88
         |           `--Gutschickia Scott 1944 WSB93, BB61
         |                |--*G. ninevehensis (Holland 1934) [=Whipplella ninevehensis] BB61
         |                `--G. convexa Tasch 1963 WSB93
         |         Haworthina Kellett 1935 BB61
         |           `--*H. bulleta (Harris & Lalicker 1932) [=Bairdia bulleta] BB61
         |         Hilboldtina Scott & Summerson 1943 BB61
         |           `--*H. multiplicata Scott & Summerson 1943 BB61
         |         Palaeocypris Brongniart 1876 BB61
         |           `--*P. edwardsii Brongniart 1876 BB61
         |         Pruvostina Scott & Summerson 1943 BB61
         |           `--*P. wanlessi Scott & Summerson 1943 BB61
         |         Ranapeltis Bassler 1941 BB61
         |           `--*R. typicalis Bassler 1941 BB61
         |         Scabriculocypris Anderson 1940 BB61
         |           `--*S. trapezoides Anderson 1940 BB61
         |         Suchonella Spizharsky 1937 BB61
         |           |--*S. typica Spizharsky 1937 BB61
         |           `--S. blomi TT05
         |         Suchonellina BB61
         |         Whipplella Holland 1934 BB61
         |           `--*W. cuneiformis Holland 1934 BB61
         |--Cyprididae M03
         |--Candonidae F08
         |--Ilyocyprididae F08
         |--Macrocyprididae [Macrocypridae, Macrocypridoidea, Macrocyprinae] WSB93
         |    |--Macrosarisa exquisita (Kaye 1964) WSB93
         |    |--Macrocypris Brady 1867 CB86
         |    |    |--*M. minna (Baird 1850) [=Cythere minna] BB61
         |    |    `--M. siliquoides Jones 1887 C01
         |    `--Macrocypria Sars 1923 BB61
         |         `--*M. sarsi (Müller 1912) (see below for synonymy) BB61
         `--Notodromadidae [Notodromadinae, Notodromides, Notodrominae, Notodromini] MD01
              |--Notodromas Liljeborg 1853 BB61
              |    `--*N. monachus (Müller 1776) BB61 [=Cypris monacha BB61, Monoculus monachus L02]
              |--Newnhamia King 1855 BB61
              |    `--*N. fenestrata King 1855 BB61
              `--Cyprois Zenker 1854 BB61
                   |--*C. marginata (Strauss 1821) [=Cypris marginata] BB61
                   |--C. ephraimensis Swain 1964 WSB93
                   `--C. tantouensis NG13

*Macrocypria sarsi (Müller 1912) [=Macrocypris sarsi, Bairdia angusta Sars 1866 non Cythere (Bairdia) angusta Jones 1850] BB61

Pontocyprididae [Argilloeciinae, Clinocyprinae, Pontocypridae, Pontocypridinae, Pontocypridoidea, Pontocyprinae] MD01

Pontocypris mytiloides (Norman 1862) [=Cythere (Bairdia) mytiloides; incl. Cypris serrulata Sars 1863, *Pontocypris serrulata] BB61

*Type species of generic name indicated


[BB61] Benson, R. H., J. M. Berdan, W. A. van den Bold, T. Hanai, I. Hessland, H. V. Howe, R. V. Kesling, S. A. Levinson, R. A. Reyment, R. C. Moore, H. W. Scott, R. H. Shaver, I. G. Sohn, L. E. Stover, F. M. Swain & P. C. Sylvester-Bradley. 1961. Systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q99–Q421. Geological Society of America and University of Kansas Press.

[C01] Chapman, F. 1901. On some fossils of Wenlock age from Mulde, near Klinteberg, Gotland; with notes by Prof. T. Rupert Jones and Dr. F. A. Bather. Annals and Magazine of Natural History, series 7, 7: 141–160, pl. 3.

[CB86] Chen D.-Q. & Bao H. 1986. Lower Permian ostracodes from the Chihsia Formation of Jurong and Longtan, Jiangsu Province. Acta Micropalaeontologica Sinica 3 (2): 107–132.

[F08] Fuhrmann, R. 2008. Die Ostrakoden- und Molluskenfauna des Auelehmprofils Zeitz (Landkreis Burgenland) und ihre Aussage zum Klima sowie zur Landnutzung im jüngeren Holozän Mitteldeutschlands. Mauritiana 20 (2): 253–281.

[G88] Gray, J. 1988. Evolution of the freshwater ecosystem: the fossil record. Palaeogeography, Palaeoclimatology, Palaeoecology 62: 1–214.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 4. Familles naturelles des genres. F. Dufart: Paris.

[M03] Martens, K. 2003. On a remarkable South African giant ostracod (Crustacea, Ostracoda, Cyprididae) from temporary pools, with additional appendages. Hydrobiologia 500: 115–130.

[MD01] Martin, J. W., & G. E. Davis. 2001. An updated classification of the Recent Crustacea. Natural History Museum Los Angeles County, Science Series 39: 1–124.

[M61] Moore, R. C. 1961. Supplement—USSR treatise on Ostracoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q422–Q429. Geological Society of America and University of Kansas Press.

[NG13] Ni, X., D. L. Gebo, M. Dagosto, J. Meng, P. Tafforeau, J. J. Flynn & K. C. Beard. 2013. The oldest known primate skeleton and early haplorhine evolution. Nature 498: 60–64.

[TT05] Tverdokhlebov, V. P., G. I. Tverdokhlebova, A. V. Minikh, M. V. Surkov & M. J. Benton. 2005. Upper Permian vertebrates and their sedimentological context in the South Urals, Russia. Earth-Science Reviews 69: 27–77.

[WSB93] Whatley, R. C., D. J. Siveter & I. D. Boomer. 1993. Arthropoda (Crustacea: Ostracoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 343–356. Chapman & Hall: London.

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