Tutu Coriaria arborea, copyright Krzysztof Ziarnek, Kenraiz.

Belongs within: Cucurbitales.

Coriaria, tutu, is a genus of shrubs and trees with a disjunct distribution in southern Europe, eastern Asia, New Zealand and Central and South America (Allan 1961).

Characters (from Allan 1961): Subshrubs, shrubs or small trees with quadrangular branchlets and simple opposite or whorled exstipulate leaves. Flowers small, pentamerous, usually in racemes; sepals imbricate, persistent; petals keeled on inside. Stamens 10, free, or the petal-opposed ones adnate; anthers large. Carpels free, 5–10, one-celled; receptacles conical; styles free, elongate. Fruits of 5–10 indehiscent cocci surrounded by the enlarged succulent petals; ovules solitary in each carpel, pendulous. Seeds compressed, with thin endosperm and straight embryos.

<==Coriaria Linnaeus 1753 [Coriariaceae, Coriariales, Coriarieae, Coriariineae] A61
    |--C. angustissima Hook. f. 1864 A61
    |--C. arborea Lindsay 1868 A61
    |    |--C. a. var. arborea A61
    |    `--C. a. var. kermadecensis Oliver 1942 A61
    |--C. kingiana Colenso 1844 (see below for synonymy) A61
    |--C. lurida Kirk 1899 A61
    |--C. nepalensis WM09
    |--C. plumosa Oliver 1942 [incl. C. lurida var. parvifolia Cockayne & Allan 1927] A61
    |--C. pottsiana Oliver 1942 A61
    |--C. pteridoides Oliver 1942 [incl. C. lurida var. acuminata Cockayne & Allan 1927] A61
    |--C. ruscifolia A61
    |--C. × sarlurida Cockayne & Allan 1927 [C. lurida × C. sarmentosa] CA27
    |--C. sarmentosa Forst. f. 1786 [incl. C. tutu Lindsay 1868] A61
    `--C. thymifolia Humb. & Bonpl. ex Willd. 1806 A61

Coriaria kingiana Colenso 1844 [incl. C. thymifolia var. undulata Petrie 1921, C. lurida var. undulata (Petrie) Allan 1931] A61

*Type species of generic name indicated


[A61] Allan, H. H. 1961. Flora of New Zealand vol. 1. Indigenous Tracheophyta: Psilopsida, Lycopsida, Filicopsida, Gymnospermae, Dicotyledones. R. E. Owen, Government Printer: Wellington (New Zealand).

[CA27] Cockayne, L., & H. H. Allan. 1927. Notes on New Zealand floristic botany, including descriptions of new species, &c. (No. 5). Transactions and Proceedings of the New Zealand Institute 57: 48–72.

[WM09] Wang, H., M. J. Moore, P. S. Soltis, C. D. Bell, S. F. Brockington, R. Alexandre, C. C. Davis, M. Latvis, S. R. Manchester & D. E. Soltis. 2009. Rosid radiation and the rapid rise of angiosperm-dominated forests. Proceedings of the National Academy of Sciences of the USA 106 (10): 3853–3858.

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