Alysson oppositus, copyright Centre for Biodiversity Genomics.

Belongs within: Bembicidae.

The Alyssontini are a group of small, slender wasps found in Eurasia, Africa and North America that stock their nests with species of Auchenorrhyncha (Hemiptera) (Bohart & Menke 1976).

Characters (from Bohart & Menke 1976, as Alyssonini): Eyes broadly separated, inner orbits essentially parallel or converging slightly below; ocelli well developed. Labrum small, usually concealed; palpal formula 6-4. Scutum with admedian lines close together, sometimes short or faint; notauli absent; no oblique scutal carina but tegula partly covered by lateral scutal flange; scutellum normally convex, not overlapping metanotum. Omaulus present; no acetabular carina nor sternaulus; episternal sulcus undefined above, represented by a pitted remnant between lower end of omaulus and anteroventral pleural margin; scrobal sulcus extending to omaulus, extension horizontal in Alysson but obliquely downward in Didineis. Metapleuron consisting of upper area only, tapering below; propodeum rather long, depressed, rectilinear, areolate, bearing small dorsolateral teeth or angles. Midcoxae approximate, precoxal sulcus complete; midtibia with two apical spurs, one or none (in some male Didineis); foretarsal rake of female weakly developed; plantulae present. Forewing media diverging before or after cu-a; stigma rather large (area greater than that of tegula); three submarginal cells, II petiolate and receiving second or both recurrent veins. Hindwing media diverging before cu-a; jugal lobe small. Gaster sessile; sternum I with a double carina toward base; male with seven visible sterna in addition to protruding pair of spines from sternum VIII; male genitalia with long gonobase, volsella rather simple but elongate, cuspis and digitus not differentiated; female pygidial plate distinct, sharply edged, setose.

<==Alyssontini SN18
    |--Didineis Wesmael 1852 BM76
    |    |--*D. lunicornis (Fabricius 1798) (see below for synonymy) BM76
    |    |--D. aculeata (Cresson 1865) [=Alyson aculeatus] BM76
    |    |--D. bactriana Gussakovskij 1937 BM76
    |    |--D. barbieri Beaumont 1968 BM76
    |    |--D. botscharnikovi Gussakovskij 1937 BM76
    |    |--D. bucharica Gussakovskij 1937 BM76
    |    |--D. clavimana Gussakovskij 1937 BM76
    |    |--D. crassicornis Handlirsch 1888 BM76
    |    |--D. dilata Malloch & Rohwer 1930 BM76
    |    |--D. koshevnikovi Kokujev 1906 BM76
    |    |--D. latimana Malloch & Roher 1930 BM76
    |    |--D. latro (Beaumont 1967) [=Alysson latro] BM76
    |    |--D. nigricans Morice 1911 BM76
    |    |--D. nodosa Fox 1894 [incl. D. clypeata Malloch & Rohwer 1930, D. santacrucae Bradley 1920] BM76
    |    |--D. ogloblini Gussakovskij 1937 BM76
    |    |--D. orientalis Cameron 1897 BM76
    |    |--D. pannonica Handlirsch 1888 BM76
    |    |--D. peculiaris Fox 1895 BM76
    |    |--D. ruthenica Gussakovskij 1937 BM76
    |    |--D. sibirica Gussakovskij 1937 BM76
    |    |    |--D. s. sibirica BM76
    |    |    `--D. s. nipponica Tsuneki 1968 BM76
    |    |--D. stevensi Rohwer 1923 BM76
    |    |--D. texana (Cresson 1872) [=Alyson texanus] BM76
    |    |--D. turanica Gussakovskij 1937 BM76
    |    |--D. vierecki Rohwer 1911 [incl. D. crassicornis Viereck 1906 non Handlirsch 1888] BM76
    |    |--D. wuestneii Handlirsch 1888 BM76
    |    `--D. zimini Gussakovskij 1937 BM76
    `--Alysson Panzer 1806 [=Allyson (l. c.), Alyson Jurine 1807] BM76
         |--*A. spinosus (Panzer 1801) (see below for synonymy) BM76
         |    |--A. s. spinosus BM76
         |    `--A. s. jaroslavensis Kokujev 1906 BM76
         |--A. annulipes Cameron 1897 BM76
         |--A. cameroni Yasumatsu & Masuda 1933 BM76
         |--A. conicus Provancher 1889 BM76
         |--A. costai Beaumont 1953 BM76
         |--A. erythrothorax Cameron 1902 BM76
         |--A. flavomaculatus Cameron 1901 [=Alyson flavomaculatus] BM76
         |--A. formosanus Tsuneki 1968 BM76
         |--A. guichardi Arnold 1951 BM76
         |--A. guignardi Provancher 1887 [incl. A. interstitialis Cameron 1902, A. petiolatus Cameron 1902] BM76
         |--A. guillarmodi Arnold 1944 BM76
         |--A. harbinensis Tsuneki 1967 BM76
         |--A. katkovi Kokujev 1906 BM76
         |--A. madecassus Arnold 1945 BM76
         |--A. maracandensis Radoszkowski 1877 [incl. A. incertus Radoszkowski 1877] BM76
         |--A. melleus Say 1837 BM76
         |--A. ocellatus Beaumont 1967 BM76
         |--A. oppositus Say 1837 BM76
         |--A. pertheesi Gorski 1852 [incl. A. festivus Mocsáry 1879] BM76
         |--A. pictetii Handlirsch 1895 BM76
         |--A. radiatus Fox 1894 BM76
         |--A. ratzeburgi Dahlbom 1843 BM76
         |--A. ruficollis Cameron 1898 BM76
         |--A. seyrigi Arnold 1945 BM76
         |--A. striatus Fox 1894 BM76
         |--A. taiwanus Sonan 1947 BM76
         |--A. testaceitarsis Cameron 1902 BM76
         |--‘Alyson’ tomentosum Macleay 1826 (n. d.) C85
         |--A. triangulifer Provancher 1887 BM76
         |    |--A. t. triangulifer BM76
         |    `--A. t. shawi Bradley 1920 BM76
         |--A. tricolor Lepeletier & Serville 1825 BM76
         `--A. verhoeffi Tsuneki 1967 BM76

*Alysson spinosus (Panzer 1801) [=Pompilus spinosus, *Alyson spinosus; incl. Sphex bimaculatus Panzer 1798 non Fuesslin 1775] BM76

*Didineis lunicornis (Fabricius 1798) [=Pompilus lunicornis, Alyson lunicornis; incl. Didineis kennedii Curtis 1836] BM76

*Type species of generic name indicated


[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[C85] Cardale, J. C. 1985. Vespoidea and Sphecoidea. In: Walton, D. W. (ed.) Zoological Catalogue of Australia vol. 2. Hymenoptera: Formicoidea, Vespoidea and Sphecoidea pp. 150–303. Australian Government Publishing Service: Canberra.

[SN18] Sann, M., O. Niehuis, R. S. Peters, C. Mayer, A. Kozlov, L. Podsiadlowski, S. Bank, K. Meusemann, B. Misof, C. Bleidorn & M. Ohl. 2018. Phylogenomic analysis of Apoidea sheds new light on the sister group of bees. BMC Evolutionary Biology 18: 71.

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