Helicininae

Pleuropoma extincta, copyright Bronwen Scott.

Belongs within: Helicinoidea.

The Helicininae are a group of operculate terrestrial snails in which the radula possesses five lateral teeth, including a comb-lateral and an accessory plate (Knight et al. 1960).

<==Helicininae BR05
    |--Bourciera Pfeiffer 1852 BR05 [incl. Pseudhelicina Sykes 1907 KC60; Bourcierinae]
    |    `--*B. helicinaeformis (Pfeiffer 1852) [=Cyclostoma helicinaeforme] BR17
    |--Ceratopoma Moellendorff 1893 KC60, S88 (see below for synonymy)
    |    `--*C. caroli (Kobelt 1886) [=Helicina caroli] KC60
    |--Hemipoma Wagner 1905 KC60
    |    `--*H. hakodadiensis (Hartmann 1890) [=Helicina hakodadiensis] KC60
    |--Orobophana Wagner 1905 KC60
    |    `--*O. uberta (Gould 1847) [=Helicina uberta] KC60
    |--Sturanya Wagner 1905 [incl. Sturanyella Pilsbry & Cooke 1934, Sturyanella] KC60
    |    `--*S. plicatilis (Mousson 1865) [=Helicina plicatilis] KC60
    |--Palaeohelicina Wagner 1905 (Palaeohelicina) KC60
    |    |--*P. (P.) fischeriana (Montrouzier 1863) [=Helicina fischeriana] KC60
    |    `--P. (P.) mayri Clench 1958 BC01
    |--Olygyra Say 1818 BR05 (see below for synonymy)
    |    |--*O. orbiculata Say 1818 [=Helicina (*Olygyra) orbiculata] KC60
    |    `--O. sprucei [=Helicina (Oligyra) sprucei] A66
    |--Sulfurina Möllendorff 1893 [incl. Hypostrongyla Tomlin 1930, Pestomena Iredale 1941 (n. n.)] KC60
    |    |--*S. (Sulfurina) citrina (Grateloup 1840) [=Helicina citrina] KC60
    |    `--S. (Kosmetopoma Wagner 1905) KC60
    |         `--S. (*K.) amaliae (Kobelt 1886) [=Helicina amaliae] KC60
    |--Schasicheila Shuttleworth 1852 [=Atoyac Baker 1928, Schasichila Fischer 1885] KC60
    |    |--*S. (Schasicheila) alata (Pfeiffer 1848) [=Helicina alata, *Atoyac alata] KC60
    |    |--S. (Misantla Baker 1928) KC60
    |    |    `--S. (*M.) misantlensis Fischer & crosse 1893 KC60
    |    `--S. (Necaxa Baker 1928) KC60
    |         `--S. (*N.) minuscula (Pfeiffer 1859) [=Helicina minuscula] KC60
    |--Alcadia Gray 1840 (see below for synonymy) KC60
    |    |  i. s.: A. delmonteana Pilsbry 1932 BC01
    |    |         A. pompholyx (Pilsbry 1933) [=Helicina pompholyx] BC01
    |    |--*A. (Alcadia) major (Gray 1824) [=Helicina major] KC60
    |    |--A. (Analcadia Wagner 1907) KC60
    |    |    `--A. (*A.) dysoni (Pfeiffer 1849) [=Helicina dysoni] KC60
    |    |--A. (Emoda Adams & Adams 1856) [incl. Glyptemoda Clench & Aguayo 1950) KC60
    |    |    `--A. (*E.) silacea (Morelet 1849) [=Helicina silacea] KC60
    |    |--A. (Idesa Adams & Adams 1856) (see below for synonymy) KC60
    |    |    `--A. (*I.) rotunda (d’Orbigny in Sagra 1842) [=Helicina rotunda] KC60
    |    |--A. (Penisoltia Baker 1954) KC60
    |    |    `--A. (*P.) hispida (Pfeiffer 1839) [=Helicina hispida] KC60
    |    `--A. (Striatemoda Baker 1940) [incl. Diaphana Guppy 1895 non Brown 1827] KC60
    |         `--A. (S.) subfusca (Menke 1828) (see below for synonymy) KC60
    |--Helicina Lamarck 1799 BR05 (see below for synonymy)
    |    |  i. s.: H. chionea Pilsbry 1949 BC01
    |    |         H. clappi BP90
    |    |         H. hollandi Adams 1849 BC01
    |    |         H. laciniosa Mighels 1845 [=Pleuropoma laciniosa; incl. Orobophana berniceia] J49
    |    |         H. rotelloidea Mighels 1845 [incl. Pleuropoma hawaiiensis] J49
    |    |--*H. (Helicina) neritella Lamarck 1801 (see below for synonymy) KC60
    |    |--H. (Oxyrhombus Fischer & Cross 1893) [incl. Angulata Wagner 1905, Tamsiana Wagner 1905] KC60
    |    |    `--H. (*O.) amoena Pfeiffer 1845 KC60
    |    |--H. (Pseudoligyra Baker 1954) KC60
    |    |    `--H. (*P.) tenuis Pfeiffer 1849 KC60
    |    `--H. (Tristramia Crosse 1863) [incl. Caloplisma Fischer & Crosse 1893, Retorquata Wagner 1905] KC60
    |         `--H. (*T.) salvini Tristram 1861 KC60
    `--Pleuropoma Moellendorff 1893 KC60, KKW12 (see below for synonymy)
         |--*P. dichroa (Möllendorff 1890) [=Helicina dichroa] SS10
         |--P. brecanrio Stanisic in Stanisic, Shea et al. 2010 SS10
         |--'Helicina (Pachystoma)’ concentrica A66
         |--P. draytonensis (Pfeiffer 1857) [incl. Helicina diversicolor Cox 1866] SS10
         |--P. extincta (Odhner 1917) SS10
         |--P. gladstonensis (Cox 1864) SS10
         |--P. gouldiana (Forbes 1851) (see below for synonymy) SS10
         |--P. jana (Cox 1872) SS10
         |--P. louisiadensis (Forbes 1851) S88
         |--P. macleayi (Brazier 1876) SS10
         |--P. queenslandica Stanisic in Stanisic, Shea et al. 2010 SS10
         |--P. reflexilabiata Stanisic in Stanisic, Shea et al. 2010 SS10
         |--P. richardson Stanisic in Stanisic, Shea et al. 2010 SS10
         |--P. solomonensis (Smith 1885) S88
         |--P. spatei Stanisic in Stanisic, Shea et al. 2010 SS10
         |--P. talusata Stanisic in Stanisic, Shea et al. 2010 SS10
         |--P. verecunda (Gould 1859) [=Helicina verecunda, P. (*Aphanoconia) verecunda, *Sphaeroconia verecunda] KC60
         `--P. walkeri (Smith 1894) KKW12 (see below for synonymy)

Alcadia Gray 1840 [incl. Eualcadia Wagner 1907, Incrustata Wagner 1907, Intusplicata Wagner 1907, Isoltia Guppy 1859, Palliata Wagner 1907, Sericea Wagner 1907] KC60

Alcadia (Idesa Adams & Adams 1856) [incl. Hjalmarsona Baker 1940, Leialcadia Wagner 1907, Mammilla Wagner 1907 non Schumacher 1817, Schrammia Guppy 1895, Weinlandella Baker 1954] KC60

Alcadia (Striatemoda) subfusca (Menke 1828) [=Helicina subfusca; incl. H. striata Lamarck 1822 non Defrance 1821, Alcadia (*Striatemoda) striata] KC60

Ceratopoma Moellendorff 1893 KC60, S88 [incl. Diversicolor Wagner 1905 KC60, Negopenia Iredale 1941 (n. n.) KC60]

Helicina Lamarck 1799 BR05 [=Euneritella Wagner 1905 KC60, Pitonnellus deMontfort 1810 KC60, Pitonnillus deMontfort 1810 KC60; incl. Ampliata Wagner 1907 KC60, Colyma Rafinesque 1815, Festiva Wagner 1910 KC60, Pachytoma Swainson 1840 KC60]

*Helicina (Helicina) neritella Lamarck 1801 [=Helix neritella, *Euneritella neritella, *Pitonnellus neritella] KC60

Olygyra Say 1818 BR05 [=Oligyra BR05; incl. Subglobulosa Wagner 1905 KC60, Succincta Wagner 1905 KC60; Oligyradae, Olygyridae]

Pleuropoma Moellendorff 1893 KC60, KKW12 [incl. Albocincta Wagner 1909 SS10, Aphanoconia Wagner 1905 SS10, Dichroa Wagner 1905 SS10, Pachystoma Wagner 1905 non Guilding 1828 KC60, Reticulata Wagner 1905 SS10, Sculpta Wagner 1905 SS10, Sphaeroconia Wagner 1909 SS10]

Pleuropoma gouldiana (Forbes 1851) [incl. Helicina fumigata Sowerby 1866, H. lizardensis Cox 1868, H. reticulata Pfeiffer 1863, Aphanoconia (Sphaeroconia) subreticulata Wagner 1909] SS10

Pleuropoma walkeri (Smith 1894) KKW12 [=Helicina walkeri KKW12, Orobophana walkeri S88, Aphanoconia (Sphaeroconia) baudinensis Wagner 1909 KKW12]

*Type species of generic name indicated

REFERENCES

[A66] Adams, H. 1866. List of land and freshwater shells collected by Mr. E. Bartlett on the Upper Amazons, and on the River Ucayali, eastern Peru, with descriptions of new species. Proceedings of the Zoological Society of London 1866: 440–445.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[BP90] Burch, J. B., & T. A. Pearce. 1990. Terrestrial Gastropoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 201–309. John Wiley & Sones: New York.

[J49] Johnson, R. I. 1949. Jesse Wedgwood Mighels with a bibliography and a catalogue of his species. Occasional Papers on Mollusks 1 (14): 213–231.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I169–I331. Geological Society of America, and University of Kansas Press.

[KKW12] Köhler, F., V. Kessner & C. Whisson. 2012. New records of non-marine, non-camaenid gastropods (Mollusca: Gastropoda) from islands off the Kimberley coast, Western Australia. Records of the Western Australian Museum 27 (1): 21–39.

[S88] Solem, A. 1988. Non-camaenid land snails of the Kimberley and Northern Territory, Australia. I. Systematics, affinities and ranges. Invertebrate Taxonomy 4: 455–604.

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.