Vertigo pusilla, copyright O. Gargominy.

Belongs within: Pupilloidea.

The Vertigininae are a group of snails with reduced tentacles and a penis lacking an appendix and with a simple retractor.

<==Vertigininae [Vertigineae, Vertigini]
    |--Bothriopupa Pilsbry 1898 BP90
    |    `--B. variolosa BP90
    |--Sterkia Pilsbry 1898 BP90
    |    `--S. eyriese BP90
    |         |--S. e. eyriese BP90
    |         `--S. e. rhoadsi BP90
    `--Vertigo Müller 1773 BR17
         |--*V. pusilla Müller 1774 BR17
         |--V. californica BP90
         |--V. coloradensis PF15
         |    |--V. c. coloradensis PF15
         |    `--V. c. arizonensis PF15
         |--V. diversidens E99
         |--V. inserta WK12
         |--V. interferens (Deshayes 1863) TTE93
         |--V. meramecensis VanDevender 1979 NC01
         |--V. milium BP90
         |--V. nouleti E99
         |--V. ovata PF15
         |--V. oviformis Michaud in Deshayes 1863 TTE93
         |--V. parvula BP90
         |--V. pygmaea G59
         |--V. tridentata G59
         `--V. ventricosa G59

*Type species of generic name indicated


[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BP90] Burch, J. B., & T. A. Pearce. 1990. Terrestrial Gastropoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 201–309. John Wiley & Sones: New York.

[E99] Esu, D. 1999. Contribution to the knowledge of Neogene climate changes in western and central Europe by means of non-marine molluscs. In: J. Agustí, L. Rook & P. Andrews (eds) The Evolution of Neogene Terrestrial Ecosystems in Europe pp. 328–354. Cambridge University Press.

[G59] Grimm, W. 1959. Land snails of Carroll County, Maryland. Nautilus 72 (4): 122–127.

[NC01] Nekola, J. C., & B. F. Coles. 2001. Systematics and ecology of Gastrocopta (Gastrocopta) rogersensis (Gastropoda: Pupillidae), a new species of land snail from the Midwest of the United States of America. Nautilus 115 (3): 105–114.

[PF15] Pilsbry, H. A., & J. H. Ferriss. 1915. Mollusca of the southwestern states, VII: the Dragoon, Mule, Santa Rita, Baboquivari, and Tucson Ranges, Arizona. Proceedings of the Academy of Natural Sciences of Philadelphia 67 (2): 363–418, pls 8–15.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

[WK12] Whisson, C. S., & F. Köhler. 2012. Gastrocopta (Mollusca, Gastropoda, Pupillidae) in the Pilbara region of Western Australia. ZooKeys 261: 15–39.

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