Suboval ambersnail Catinella vermeta, copyright Ron Matsumoto.

Belongs within: Heterobranchia.
Contains: Succineinae.

The Succineoidei are a group of terrestrial molluscs whose members have optic tentacles that retract that do not invert, and elasmognathous jaws with lateral winged processes and a greatly enlarged attachment plate. The cosmopolitan Succineidae are thin-shelled, conical snails with a greatly enlarged aperture and strongly impressed sutures. Apical whorls have a distinctly nipple-like appearance. Many species are found along the margins of water bodies, feeding on algal films growing on damp surfaces, and the Miocene Papyrotheca may have been entirely aquatic (Gray 1988). The Australasian Athoracophoridae are brightly coloured slugs with the mantle confined to a small triangular area in the centre of the back. The internal shell has been reduced to scattered fragments or granules of calcium (Stanisic et al. 2010).

<==Succineoidei [Elasmognatha, Heterurethra, Succineida, Succineiformes, Succineoinei]
    |--Athoracophoridae [Athoracophoracea, Athoracophoroidea, Tracheopulmonata] SS10
    |    |--Aneitella virgata T84
    |    |--Triboniophorus Humbert 1863 SS10
    |    |    `--*T. graeffei Humbert 1863 (see below for synonymy) SS10
    |    |--Vomanus Iredale 1915 P61
    |    |    `--*V. marmoreus (Hutton 1879) [=Konophora marmoreus] P61
    |    |--Amphiconophora Suter 1897 P61
    |    |    |--*A. gigantea (Suter 1909) P61
    |    |    `--A. verrucosa (Simroth 1889) P61
    |    |--Aneitea Gray 1860 [Aneiteadae, Aneiteidae, Aneiteinae] BR05
    |    |    |--*A. macdonaldii Gray 1860 BR17
    |    |    `--A. simrothi T84
    |    |--Athoracophorus Gould 1852 (see below for synonymy) BR05
    |    |    `--*A. bitentaculatus (Quoy & Gaimard 1832) P61 [=Limax bitentaculatus P61, *Janella bitentaculata BR17]
    |    |         |--A. b. bitentaculatus P61
    |    |         `--A. b. rufovenosus Suter 1909 P61
    |    `--Pseudaneitea Cockerell 1891 P61
    |         |--*P. papillata (Hutton 1879) [=Athoracophorus papillatus] P61
    |         |--P. dendyi (Suter 1897) P61
    |         |--P. huttoni (Suter 1909) P61
    |         |--P. martensi (Suter 1909) P61
    |         |--P. schauinslandi (Plate 1897) P61
    |         |--P. simrothi (Suter 1896) P61
    |         `--P. sorenseni Powell 1955 P61
    `--Succineidae [Diplotrema, Succineacea, Succineoidea] SS10
         |  i. s.: Quickia FGN07
         |         Spirancinea Iredale 1945 SS10
         |           `--*S. norfolkensis (Sykes 1900) (see below for synonymy) SS10
         |         Papyrotheca G88
         |         Omalonyx T84
         |           |--O. felina Guppy 1872 G79
         |           |--O. matheroni T84
         |           `--O. unguis G79
         |--Catinella Pease 1870 [Catinellinae] BR05
         |    |--*C. rubida Pease 1870 BR17
         |    |--C. avara BP90
         |    |--C. gelida (Baker 1927) [=Succinea grosvenori gelida] H63
         |    |--C. oklahomarum (Webb 1953) G68
         |    |    |--G. o. oklahomarum G68
         |    |    `--G. o. loculosa (Webb 1954) G68
         |    |--C. pinicola Grimm 1960 G68
         |    |--C. rehderi H63
         |    |--C. stretchiana (Bland 1865) [=Succinea stretchiana] H63
         |    |--C. vermeta H68
         |    `--C. wandae H63
         `--Succineinae BR05

Athoracophorus Gould 1852 [=Janella Gray 1850 non Grateloup 1838; Athoracophorinae, Janellidae, Janellinae] BR05

*Spirancinea norfolkensis (Sykes 1900) [=Succinea norfolkensis; incl. Su. humerosa Preston 1913, Su. nepeanensis Preston 1913] SS10

*Triboniophorus graeffei Humbert 1863 [incl. T. brisbanensis Pfeiffer 1900, T. krefftii Keferstein 1865, Aneitea graeffei rosea Hedley 1893, T. schuttei Keferstein 1865] SS10

*Type species of generic name indicated


[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BP90] Burch, J. B., & T. A. Pearce. 1990. Terrestrial Gastropoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 201–309. John Wiley & Sones: New York.

[FGN07] Fontaine, B., O. Gargominy & E. Neubert. 2007. Land snail diversity of the savanna/forest mosaic in Lopé National Park, Gabon. Malacologia 49 (2): 313–338.

[G79] Gibbons, J. S. 1879. Comparison of Omalonyx unguis, D'Orb., with O. felina, Guppy. Journal of Conchology 2: 99–101.

[G88] Gray, J. 1988. Evolution of the freshwater ecosystem: the fossil record. Palaeogeography, Palaeoclimatology, Palaeoecology 62: 1–214.

[G68] Grimm, F. W. 1968. A note on Catinella oklahomarum. Nautilus 81 (3): 84–85.

[H63] Hubricht, L. 1963. Some Succineidae, with a new species. Nautilus 76 (4): 135–138.

[H68] Hubricht, L. 1968. The land snails of Mammoth Cave National Park, Kentucky. Nautilus 82 (1): 24–28.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4th ed. Whitcombe and Tombs Limited: Christchurch.

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.

[T84] Tillier, S. 1984. Patterns of digestive tract morphology in the limacisation of helicarionid, succineid and athoracophorid snails and slugs (Mollusca: Pulmonata). Malacologia 25 (1): 173–192.

Last updated: 19 February 2022.

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