Amber snail Succinea putris, copyright Charles J. Sharp.

Belongs within: Heterobranchia.

The Succineoidei are a group of terrestrial molluscs whose members have optic tentacles that retract that do not invert, and elasmognathous jaws with lateral winged processes and a greatly enlarged attachment plate (Stanisic et al. 2010). The cosmopolitan Succineidae are thin-shelled, conical snails whereas the Australasian Athoracophoridae are brightly coloured slugs.

<==Succineoidei [Elasmognatha, Heterurethra, Succineida, Succineiformes, Succineoinei]
    |--Athoracophoridae [Athoracophoracea, Athoracophoroidea, Tracheopulmonata] SS10
    |    |--Aneitella virgata T84
    |    |--Triboniophorus Humbert 1863 SS10
    |    |    `--*T. graeffei Humbert 1863 (see below for synonymy) SS10
    |    |--Vomanus Iredale 1915 P61
    |    |    `--*V. marmoreus (Hutton 1879) [=Konophora marmoreus] P61
    |    |--Amphiconophora Suter 1897 P61
    |    |    |--*A. gigantea (Suter 1909) P61
    |    |    `--A. verrucosa (Simroth 1889) P61
    |    |--Aneitea Gray 1860 [Aneiteadae, Aneiteidae, Aneiteinae] BR05
    |    |    |--*A. macdonaldii Gray 1860 BR17
    |    |    `--A. simrothi T84
    |    |--Athoracophorus Gould 1852 (see below for synonymy) BR05
    |    |    `--*A. bitentaculatus (Quoy & Gaimard 1832) P61 [=Limax bitentaculatus P61, *Janella bitentaculata BR17]
    |    |         |--A. b. bitentaculatus P61
    |    |         `--A. b. rufovenosus Suter 1909 P61
    |    `--Pseudaneitea Cockerell 1891 P61
    |         |--*P. papillata (Hutton 1879) [=Athoracophorus papillatus] P61
    |         |--P. dendyi (Suter 1897) P61
    |         |--P. huttoni (Suter 1909) P61
    |         |--P. martensi (Suter 1909) P61
    |         |--P. schauinslandi (Plate 1897) P61
    |         |--P. simrothi (Suter 1896) P61
    |         `--P. sorenseni Powell 1955 P61
    `--Succineidae [Diplotrema, Succineacea, Succineoidea] BR05
         |  i. s.: Quickia FGN07
         |         Spirancinea Iredale 1945 SS10
         |           `--*S. norfolkensis (Sykes 1900) (see below for synonymy) SS10
         |         Papyrotheca G88
         |         Omalonyx matheroni T84
         |--Catinella Pease 1870 [Catinellinae] BR05
         |    |--*C. rubida Pease 1870 BR17
         |    `--C. avara BP90
         `--Succineinae BR05
              |--Hyalimax Adams & Adams 1855 [Hyalimacidae, Hyalimacinae] BR05
              |    `--*H. perlucidus (Quoy & Gaimard 1832) [=Limax perlucidus] BR17
              |--Austrosuccinea Iredale 1937 [incl. Arborcinea Iredale 1937] S88
              |    |--*A. australis (Férussac 1861) [=Succinea australis] SS10
              |    |--A. macgillivrayi (Cox 1864) [incl. Succinea nortoni Cox 1864] SS10
              |    `--A. strigillata (Adams & Angas 1864) W08
              |--Oxyloma Westerlund 1885 [Oxylomatinae, Oxylominae] BR05
              |    |--*O. dunkeri (Pfeiffer 1865) [=Succinea dunkeri] BR17
              |    |--O. elegans (Risso 1826) CP06
              |    |--O. pfeifferi (Rossmässler 1835) V09, CP06
              |    `--O. retusa BP90
              `--Succinea Draparnaud 1801 SS10 [incl. Papusuccinea Iredale 1941 S88; Succineae]
                   |--*S. putris (Linnaeus 1758) SS10 [=Helix putris SS10, Bulimus putris PB27]
                   |--S. aperta Lea 1838 F27
                   |--‘Papusuccinea’ archeyi (Powell 1933) P61
                   |--S. avara B59
                   |--S. baumanni Sturany 1894 V07
                   |--S. boissyi (Deshayes 1863) TTE93
                   |--S. caducca Mighels 1845 J49
                   |--S. chudeaui [incl. S. chudeaui var. leyneni Dupuis & Putzeys 1923] PB27
                   |--S. congoensis PB27
                   |--S. coxi Finlay 1927 [=S. aperta Cox 1868 non Lea 1838] F27
                   |--S. grosvenori B59
                   |--S. ovalis BP90
                   |--S. propinqua T84
                   |--S. rotundata Gould 1845 [incl. S. patula Mighels 1845 non Lamarck in de Blainville 1827] J49
                   |--S. strubelli [=*Papusuccinea strubelli] P61
                   `--S. wilsoni Lea 1864 BF89

Athoracophorus Gould 1852 [=Janella Gray 1850 non Grateloup 1838; Athoracophorinae, Janellidae, Janellinae] BR05

*Succinea norfolkensis (Sykes 1900) [=Succinea norfolkensis; incl. Su. humerosa Preston 1913, Su. nepeanensis Preston 1913] SS10

*Triboniophorus graeffei Humbert 1863 [incl. T. brisbanensis Pfeiffer 1900, T. krefftii Keferstein 1865, Aneitea graeffei rosea Hedley 1893, T. schuttei Keferstein 1865] SS10

*Type species of generic name indicated


[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[B59] Branson, B. A. 1959. An ecologic observation on Succinea avara. Nautilus 72 (4): 145–146.

[BP90] Burch, J. B., & T. A. Pearce. 1990. Terrestrial Gastropoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 201–309. John Wiley & Sones: New York.

[BF89] Burnham, B. H., & P. E. Fell. 1989. Distribution of Melampus bidentatus (Say) and Succinea wilsoni (Lea) within a tidal marsh in eastern Connecticut. Nautilus 103 (3): 109–112.

[CP06] Cameron, R. A. D., B. M. Pokryszko, A. Riedel & A. Wiktor. 2006. Checklists, systematics and the CLECOM initiative: an alternative view from Europe. Malacologia 49 (1): 225–230.

[F27] Finlay, H. J. 1927. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488–533.

[FGN07] Fontaine, B., O. Gargominy & E. Neubert. 2007. Land snail diversity of the savanna/forest mosaic in Lopé National Park, Gabon. Malacologia 49 (2): 313–338.

[G88] Gray, J. 1988. Evolution of the freshwater ecosystem: the fossil record. Palaeogeography, Palaeoclimatology, Palaeoecology 62: 1–214.

[J49] Johnson, R. I. 1949. Jesse Wedgwood Mighels with a bibliography and a catalogue of his species. Occasional Papers on Mollusks 1 (14): 213–231.

[PB27] Pilsbry, H. A., & J. Bequaert. 1927. The aquatic mollusks of the Belgian Congo, with a geographical and ecological account of Congo malacology. Bulletin of the American Museum of Natural History 53 (2): 69–602, pls 10–77.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4th ed. Whitcombe and Tombs Limited: Christchurch.

[S88] Solem, A. 1988. Non-camaenid land snails of the Kimberley and Northern Territory, Australia. I. Systematics, affinities and ranges. Invertebrate Taxonomy 4: 455–604.

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.

[T84] Tillier, S. 1984. Patterns of digestive tract morphology in the limacisation of helicarionid, succineid and athoracophorid snails and slugs (Mollusca: Pulmonata). Malacologia 25 (1): 173–192.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

[V07] Verdcourt, B. 2007. Miscellaneous notes on tropical African non-marine Mollusca. Annales Historico-Naturales Musei Nationalis Hungarici 99: 189–197.

[V09] Verdcourt, B. (ed.) 2009. Additions to the wild fauna and flora of the Royal Botanic Gardens, Kew. XXVI. Miscellaneous records. Kew Bulletin 64 (1): 183–194.

[W08] Wilson, B. 2008. Terrestrial gastropods of Faure Island, Shark Bay, Western Australia. Records of the Western Australian Museum Supplement 75: 21–24.

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