Endomychidae

Dapsa trimaculata, copyright Bobgaia.


Belongs within: Coccinelloidea.

The Endomychidae are a group of often brightly coloured fungus-feeding beetles.

Characters (from Lawrence & Britton 1991): Broadly ovate to elongate and globose to moderately flattened, glabrous or clothed with decumbent and erect hairs, often brightly patterned with red or yellow and black. Frontoclypeal suture present; antennae moderately long, usually with relatively loose club; maxillary palps fusiform, labial palps may be securiform; fore coxae globular, trochantins concealed, fore coxal cavities externally open or closed, internally closed; mid coxal cavities moderately to widely separated, laterally open or closed; tarsi trimerous or tetramerous, second tarsal segment commonly lobed, extending beneath reduced third segment; tarsal claws simple; abdomen with five or six ventrites, none connate, femoral lines on first ventrite usually absent. Larva usually oblong to ovate (rarely elongate), slightly to strongly flattened, usually with lateral tergal processes on all thoracic and most abdominal segments and only rarely with short urogomphi; antennae often long (with segment 2 elongate and 1 and 3 reduced), their sockets well separated from mandibular articulations; head often broad at base, strongly narrowed towards mouth-parts; mala obtuse.

Endomychidae B14
    |--Trochoideus desjardinsi LB91
    |--Displotera LB91
    |--Mycetaea subterranea LB91
    |--Stenotarsus pisoniae LB91
    |--Epipocus [Epipocinae] MW15
    |--Endomychus [Endomychinae] MW15
    |--Eumorphus [Lycoperdininae] B14
    |    `--E. quadriguttatus (Illiger 1800) B14
    |--Encymon LB91
    |    |--E. clavicornis C70
    |    `--E. immaculatus LB91
    `--Dapsa C01
         |--D. barbara Lucas 1846 E12
         |--D. (Phylira) horvathi Csiki 1901 C01
         |--D. limbata C01
         |--D. nigripennis C01
         `--D. trimaculata C01

*Type species of generic name indicated

REFERENCES

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[C70] CSIRO. 1970. The Insects of Australia: A textbook for students and research workers. Melbourne University Press.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[MW15] McKenna, D. D., A. L. Wild, K. Kanda, C. L. Bellamy, R. G. Beutel, M. S. Caterino, C. W. Farnum, D. C. Hawks, M. A. Ivie, M. L. Jameson, R. A. B. Leschen, A. E. Marvaldi, J. V. McHugh, A. F. Newton, J. A. Robertson, M. K. Thayer, M. F. Whiting, J. F. Lawrence, A. Ślipiński, D. R. Maddison & B. D. Farrell. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40 (4): 835–880.

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