Bushveld elephant shrew Elephantulus intufi, copyright Yathin.

Belongs within: Atlantogenata.

The Macroscelididae, elephant shrews, are a group of small, cursorial insectivorous mammals known from Africa. Members of this group have long, mobile snouts that provide the inspiration for their vernacular name.

<==Macroscelididae [Dipogales, Macroscelidea, Macroscelidoidea]
    |  i. s.: Chambius kasserinensis Hartenberger 1986 GH99
    |         Miorhynchocyon OB13
    |--Mylomygale Broom 1948 [Mylomygalinae] V67
    |    `--M. spiersi Broom 1948 B78
    |--Myohyracinae [Myohyracidae] M78
    |    |--Protypotheroides Stromer 1922 M78
    |    |    `--P. beetzi Stromer 1922 B78
    |    `--Myohyrax Andrews 1914 M78
    |         |--*M. oswaldi Andrews 1914 M78 [incl. M. doederleini Stromer 1926 B78]
    |         `--M. osborni D07
    |--Rhynchocyon Peters 1847 V67 [incl. Rhinonax Thomas 1918 V67; Rhynchocyoninae B78, Rhynchocyonini]
    |    |  i. s.: R. clarki Butler & Hopwood 1957 B78
    |    |         R. rusingae Butler 1969 B78
    |    |         R. stuhlmanni JP84
    |    |--+--R. chrysopygus FS15
    |    |  `--R. udzungwensis FS15
    |    `--+--R. cirnei FS15
    |       `--R. petersi FS15
    `--Macroscelidinae [Macroscelidini] B78
         |  i. s.: Palaeothentoides Stromer 1932 V67
         |           `--P. africanus Stromer 1932 B78
         |         Metolbodotes Schlosser 1910 [=Metoldobotes] V67
         |           `--M. stromeri Schlosser 1910 B78
         |         Nasilio Thomas & Schwann 1906 V67
         |--+--Macroscelides Smith 1829 FS15, V67 (see below for synonymy)
         |  |    `--M. proboscideus ANG03
         |  |         |--M. p. proboscideus B78
         |  |         `--M. p. vagans Butler & Greenwood 1976 B78
         |  `--+--‘Elephantulus’ rozeti FS15
         |     `--Petrodromus Peters 1846 FS15, V67 [incl. Cercoctenus Hollister 1916 V67, Mesoctenus Thomas 1918 V67]
         |          |--P. sultan GM71
         |          `--P. tetradactylus IT07
         `--Elephantulus Thomas & Schwann 1906 FS15, V67 [incl. Elephantomys Broom 1937]
              |  i. s.: E. antiquus Broom 1948 B78
              |         E. broomi [incl. Elephantomys langi Broom 1937] B78
              |         E. vandami B78
              |--+--E. revoili FS15
              |  `--+--E. fuscipes FS15 [=Nasilio fuscipes B78]
              |     `--E. fuscus FS15 [=Nasilio fuscus B78]
              |          |--E. f. fuscus B78
              |          `--E. f. leakeyi (Butler & Greenwood 1965) B78
              `--+--+--E. myurus FS15
                 |  `--+--E. edwardii FS15
                 |     `--E. pilicaudus FS15
                 `--+--+--*E. rupestris B78, FS15
                    |  `--E. intufi FS15 [=Elephantomys intufi B78]
                    `--+--E. brachyrhynchus FS15 [=Nasilio brachyrhynchus B78]
                       `--E. rufescens FS15

Macroscelides Smith 1829 FS15, V67 [incl. Diposorex Blainville 1838 V67, Eumerus Geoffroy Saint-Hailaire 1829 V67, Macroscelis Fischer 1830 V67, Rhinomys Lichtenstein 1831 V67]

*Type species of generic name indicated


[AM02] Asher, R. J., M. C. McKenna, R. J. Emry, A. R. Tabrum & D. G. Kron. 2002. Morphology and relationships of Apternodus and other extinct, zalambdodont, placental mammals. Bulletin of the American Museum of Natural History 273: 1–117.

[B78] Butler, P. M. 1978. Insectivora and Chiroptera. In: Maglio, V. J., & H. B. S. Cooke (eds) Evolution of African Mammals pp. 56–68. Harvard University Press: Cambridge (Massachusetts).

[D07] Dixon, D. 2007. The Complete Illustrated Encyclopedia of Dinosaurs & Prehistoric Creatures. Hermes House: London.

[FS15] Faurby, S., & J.-C. Svenning. 2015. A species-level phylogeny of all extant and late Quaternary extinct mammals using a novel heuristic-hierarchical Bayesian approach. Molecular Phylogenetics and Evolution 84: 14–26.

[GH99] Gheerbrant, E. & J.-L. Hartenberger. 1999. Nouveau mammifère insectivore (?Lipotyphla, ?Erinaceomorpha) de l’Eocène inférieur de Chambi (Tunisie). Paläontologische Zeitschrift 73 (1–2): 143–156.

[GM71] Goodman, M., & G. W. Moore. 1971. Immunodiffusion systematics of the primates I. The Catarrhini. Systematic Zoology 20 (1): 19–62.

[IT07] Isaac, N. J. B., S. T. Turvey, B. Collen, C. Waterman & J. E. M. Baillie. 2007. Mammals on the EDGE: conservation priorities based on threat and phylogeny. PloS One 2 (3): e296.

[JP84] Jolicoeur, P., P. Pirlot, G. Baron & H. Stephan. 1984. Brain structure and correlation patterns in Insectivora, Chiroptera, and Primates. Systematic Zoology 33 (1): 14–29.

[M78] Meyer, G. E. 1978. Hyracoidea. In: Maglio, V. J., & H. B. S. Cooke (eds) Evolution of African Mammals pp. 284–314. Harvard University Press: Cambridge (Massachusetts).

[OB13] O'Leary, M. A., J. I. Bloch, J. J. Flynn, T. J. Gaudin, A. Giallombardo, N. P. Giannini, S. L. Goldberg, B. P. Kraatz, Z.-X. Luo, J. Meng, X. Ni, M. J. Novacek, F. A. Perini, Z. S. Randall, G. W. Rougier, E. J. Sargis, M. T. Silcox, N. B. Simmons, M. Spaulding, P. M. Velazco, M. Weksler, J. R. Wible & A. L. Cirranello. 2013. The placental mammal ancestor and the post-K–Pg radiation of placentals. Science 339: 662–667.

[V67] Van Valen, L. 1967. New Paleocene insectivores and insectivore classification. Bulletin of the American Museum of Natural History 135 (5): 217–284.

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