Larva of Philorus sp., from the California Department of Fish and Wildlife.

Belongs within: Neoneura.
Contains: Psychodoidea.

The Psychodomorpha is a clade of flies including the moth flies of the Psychodidae and their close relatives. In the circumscription given below, it is supported by molecular data (Wiegmann et al. 2011).

<==Psychodomorpha WT11
    |--+--Kuperwoodidae GE05
    |  |--Hennigmatidae GE05
    |  |--Grauvogelia [Grauvogeliidae] GE05
    |  |    `--G. arzvilleriana GE05
    |  `--+--Psychodoidea WT11
    |     `--Nadiptera [Nadipteridae] GE05
    |          `--N. kaluginae KK02
    `--Blephariceridae [Blepharicerimorpha] WT11
         |--Edwardsina [Edwardsininae] C09
         |    |--E. ferrugiana CM91
         |    `--E. gigantea WT11
         |--Apistomyiinae [Apistomyiini] CM70
         |    |--Apistomyia tonnoiri CM91
         |    |--Parapistomyia CM91
         |    |--Austrocurupira CM91
         |    `--Neocurupira CM70
         `--Blepharicerinae H81
              |  i. s.: Aposonalco amoyote C09
              |         Kelloggina C09
              |         Limonicola lichanos C09
              |--Paltostoma C09 [Paltostomini H81]
              |    |--P. argyrocincta C09
              |    |--P. exserta C09
              |    `--P. schineri C09
              `--Blepharicerini H81
                   |--Bibiocephala grandis H81
                   |--Blepharicera H81
                   |    |--B. ostensackeni H81
                   |    `--B. tenuipes H81
                   |--Dioptopsis H81
                   |    |--D. aylmeri H81
                   |    `--D. sequioarum H81
                   |--Agathon H81
                   |    |--A. comstocki H81
                   |    |--A. doanei H81
                   |    `--A. elegantulus H81
                   `--Philorus H81
                        |--P. californicus H81
                        |--P. jacinto H81
                        `--P. yosemite H81

*Type species of generic name indicated


[CM70] Colless, D. H., & D. K. McAlpine. 1970. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 656–740. Melbourne University Press.

[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).

[C09] Courtney, G. W. 2009. Blephariceridae (net-winged midges). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 237–243. NRC Research Press: Ottawa.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[H81] Hogue, C. L. 1981. Blephariceridae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 191–197. Research Branch, Agriculture Canada.

[KK02] Krzemiński, W., & E. Krzemińska. 2002. Rhaetaniidae, a new family of the Diptera from the Upper Triassic of Great Britain (Diptera: Nematocera). Annales Zoologici 52 (2): 211–213.

[WT11] Wiegmann, B. M., M. D. Trautwein, I. S. Winkler, N. B. Barr, J.-W. Kim, C. Lambkin, M. A. Bertone, B. K. Cassel, K. M. Bayless, A. M. Heimberg, B. M. Wheeler, K. J. Peterson, T. Pape, B. J. Sinclair, J. H. Skevington, V. Blagoderov, J. Caravas, S. N. Kutty, U. Schmidt-Ott, G. E. Kampmeier, F. C. Thompson, D. A. Grimaldi, A. T. Beckenbach, G. W. Courtney, M. Friedrich, R. Meier & D. K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the USA 108 (14): 5690–5695.

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