Ozodiceromyia sp., possibly O. notata, copyright Yurika Alexander.

Belongs within: Therevidae.

The Therevinae are a widespread group of stiletto flies characterised primarily by features of the genitalia. Members include the diverse North American genus Ozodiceromyia, characterised by areas of reduced pubescence on the frons, thorax and/or abdomen, with the frons usually being shiny black in females and having a small shiny black area in males (Gaimari & Webb 2009).

Characters (from Winterton et al. 2001): Male genitalia with dorsal apodeme of parameral sheath free from gonocoxal apodeme; ventral apodeme of parameral sheath simple. Female genitalia with acanthophorites well developed; intersegmental membrane between sternite 8 and furca sclerotised; tergite 8 joined to tergite 9+10 by sclerotised bridge; two spermathecae; spermathecal sac present. Pupa with alar spine present. Larva with labial beard present; anterior maxillary palp large.

<==Therevinae GW09
    `--Cyclotelini GW09
         |--Anolinga longiventris GW09
         |--Breviperna placida GW09
         |--Coleiana GW09
         |--Crebraseta GW09
         |--Nesonana GW09
         |--Spiracolis GW09
         |--Cyclotelus GW09
         |    |--C. bellus GW09
         |    |--C. nigrifrons GW09
         |    |--C. pictipennis GW09
         |    |--C. rufiventris GW09
         |    `--C. sumichrasti GW09
         `--Ozodiceromyia GW09
              |--O. argentifera GW09
              |--O. costalis BT04
              |--O. mexicana GW09
              |--O. notata GW09
              `--O. signatipennis GW09

Therevinae incertae sedis:
  Anabarhynchus WY01
    |--A. calceatus CM91
    `--A. tristis WY01
  Irwiniella WY01
  Megathereva WY01
  Platycarenum WY01
  Brachylinga cinerea IL81, GW09
  Thereva Latreille 1802 L02
    |--*T. plebeja [=Bibio plebeja] L02
    |--T. anilis G20
    |--T. bipunctata K01
    |--T. fucata GW09
    |--T. marginula K01
    `--T. nobilitata RD77
  Psilocephala WY01
    |--P. electrella GE05
    |--P. hypogaea IL81
    `--P. scudderi IL81

*Type species of generic name indicated


[BT04] Bybee, S. M., S. D. Taylor, C. R. Nelson & M. F. Whiting. 2004. A phylogeny of robber flies (Diptera: Asilidae) at the subfamilial level: molecular evidence. Molecular Phylogenetics and Evolution 30: 789–797.

[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).

[GW09] Gaimari, S. D., & D. W. Webb. 2009. Therevidae (stiletto flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 633–647. NRC Research Press: Ottawa.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[IL81] Irwin, M. E., & L. Lyneborg. 1981. Therevidae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 513–523. Research Branch, Agriculture Canada.

[K01] Kertész, K. 1901. Legyek [Dipteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 179–201. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms' General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

[WY01] Winterton, S. L., L. Yang, B. M. Wiegmann & D. K. Yeates. 2001. Phylogenetic revision of Agapophytinae subf. n. (Diptera: Therevidae) based on molecular and morphological evidence. Systematic Entomology 26: 173–211.

Last updated: 2 January 2021.

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