Empidoidea

Protempis antennata, from Grimaldi & Engel (2005).

Belongs within: Neoneura.
Contains: Hybotidae, Dolichopodidae, Empididae.

The Empidoidea are a clade of long-legged flies supported by molecular data. Members of this clade are predators as adults and larvae (Grimaldi & Engel 2005). In the past this group has been divided between the families Dolichopodidae and Empididae but recognition of the latter as paraphyletic has led to its division between the Empididae sensu stricto, Atelestidae, Hybotidae and basal Dolichopodidae. The Atelestidae have the costa ending at or near vein M₁₊₂, vein R₅ originating as far or further from the humeral crossvein than the length of said crossvein, the prosternum and proepisternum separated, and symmetrical, unrotated male terminalia (Wahlberg & Johanson 2018).

<==Empidoidea [Oreogetoninae]
    |  i. s.: Homalocnemis WJ18
    |         Gondwanamyia WJ18
    |--Protoreogeton admirabilis GE05
    |--Homalocnemimus abjugatus GE05
    |--Protempis [Protempididae, Protempidinae] GE05
    |    |--P. antennata GE05
    |    `--P. minuta Ran 1998 GC99
    `--+--+--Hybotidae WJ18
       |  `--+--Dolichopodidae WJ18
       |     `--+--Empididae WJ18
       |        `--Ragadidae WJ18
       |             |--Iteaphila [Iteaphilinae] WJ18
       |             |    |--I. macquarti SK81
       |             |    `--+--I. furcata WJ18
       |             |       `--Anthepiscopus SK81
       |             `--Ragadinae WJ18
       |                  |--Dipsomyia WJ18
       |                  |--Hormopeza brevicornis WJ18, SK81
       |                  |--Hydropeza WJ18
       |                  |--Zanclotus WJ18
       |                  `--Ragas WJ18
       |                       |--R. primigenia SK81
       |                       `--R. unica WJ18
       `--Atelestidae WT11
            |  i. s.: Dianafranskia fisheri GE05
            |--Atelestinae WJ18
            |    |--Acarteroptera WJ18
            |    |--Alavesia subiasi WJ18, GE05
            |    |--Meghyperus WJ18
            |    |--Atelestus WT11 [incl. Platycnema K87]
            |    |     `--A. pulicarius WT11
            |    `--Atelestites Grimaldi & Cumming 1999 GC99
            |         `--*A. senectus Grimaldi & Cumming 1999 GC99
            `--+--Burmitempis halteralis Cockerell 1917 GC99
               |--Nemedina [Nemedininae] WJ18
               |    `--N. alamirabilis GC99
               |--Prolatomyia Grimaldi & Cumming 1999 GC99
               |    `--*P. elongata Grimaldi & Cumming 1999 GC99
               |--Cretodromia Grimaldi & Cumming 1999 GC99
               |    `--*C. glaesa Grimaldi & Cumming 1999 GC99
               |--Phaetempis Grimaldi & Cumming 1999 GC99
               |    `--*P. lebanensis Grimaldi & Cumming 1999 GC99
               |--Nemedromia Grimaldi & Cumming 1999 GC99
               |    |--*N. campania Grimaldi & Cumming 1999 GC99
               |    |--N. telescopica Grimaldi & Cumming 1999 GC99
               |    `--N. turonia Grimaldi & Cumming 1999 GC99
               `--Neoturonius Grimaldi & Cumming 1999 GC99
                    |--*N. asymmetrus Grimaldi & Cumming 1999 GC99
                    |--N. cretatus Grimaldi & Cumming 1999 GC99
                    `--N. vetus Grimaldi & Cumming 1999 GC99

*Type species of generic name indicated

REFERENCES

[GC99] Grimaldi, D., & J. Cumming. 1999. Brachyceran Diptera in Cretaceous ambers and Mesozoic diversification of the Eremoneura. Bulletin of the American Museum of Natural History 239: 1–124.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[K87] Kessel, E. L. 1987. Platypezidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 681–688. Research Branch, Agriculture Canada.

[SK81] Steyskal, G. C., & L. V. Knutson. 1981. Empididae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 607–624. Research Branch, Agriculture Canada.

[WJ18] Wahlberg, E., & K. A. Johanson. 2018. Molecular phylogenetics reveals novel relationships within Empidoidea (Diptera). Systematic Entomology 43: 619–636.

[WT11] Wiegmann, B. M., M. D. Trautwein, I. S. Winkler, N. B. Barr, J.-W. Kim, C. Lambkin, M. A. Bertone, B. K. Cassel, K. M. Bayless, A. M. Heimberg, B. M. Wheeler, K. J. Peterson, T. Pape, B. J. Sinclair, J. H. Skevington, V. Blagoderov, J. Caravas, S. N. Kutty, U. Schmidt-Ott, G. E. Kampmeier, F. C. Thompson, D. A. Grimaldi, A. T. Beckenbach, G. W. Courtney, M. Friedrich, R. Meier & D. K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the USA 108 (14): 5690–5695.

Last updated: 18 July 2021.