Part and counterpart of Aeschnidium heishankowense, from here.

Belongs within: Euepiproctophora.
Contains: Eteoanisoptera.

The Anisoptera as recognised here includes the modern dragonflies (Eteoanisoptera) and a number of close fossil relatives, united by the possession of non-petiolate wings. Excluding the basal Henrotayia marci, whose affinities are somewhat uncertain, Anisoptera are also supported by fore wings with the nodal Cr nearly perpendicular to RA and ScP, and with a pseudo-IR1 vein; the presence of an 'anisopteroid vein' (AA + CuA)b in the hind wing; and a transverse anal triangle in the male. The Aeschnidiidae are united with modern dragonflies by the long hypertriangle in the fore wing in contrast to the short hypertriangle in Liassogomphidae (Fleck et al. 2003).

<==Anisoptera [Pananisoptera]
    |  i. s.: Sona R02 [Sonidae FB03]
    |           `--S. nectes Pritykina 1986 R02
    |         Petaliaeschna fletcheri Fraser 1927 FB03
    |--Henrotayia Fleck, Bechly et al. 2003 [Henrotayiidae] FB03
    |    `--*H. marci Fleck, Bechly et al. 2003 FB03
    `--+--Liassogomphidae [Gomphitidae] FB03
       |    |--Liassogomphus brodiei (Buckman 1843) FB03
       |    |--Elattogomphus latus Bode 1953 NB01
       |    |--Proinogomphus bodei (Handlirsch 1939) NB01
       |    `--Phthitogomphus angulatus (Handlirsch 1939) NB01
       `--Neoanisoptera NB01
            |--Eteoanisoptera FB03
            `--Aeschnidiidae FB03
                 |--Aeschnidium heishankowense ZBH03
                 |--Huizhougenia Lin 1980 KS02
                 |--Nothomacromia [Nothomacromiidae] S02
                 |--Aeschnidopsis flindersiensis RJ93
                 `--Hebeiaeschnidia fengningensis RJ93

*Type species of generic name indicated


[FB03] Fleck, G., G. Bechly, X. Martínez-Delclòs, E. Jarzembowski, R. Coram & A. Nel. 2003. Phylogeny and classification of the Stenophlebioptera (Odonata: Epiproctophora). Annales de la Societe Entomologique de France 39 (1): 55–93.

[KS02] Kluge, N. Yu., & N. D. Sinitshenkova. 2002. Order Ephemerida Latreille, 1810. The true mayflies (=Ephemeroptera Hyatt et Arms, 1891 (s. l.); =Euephemeroptera Kluge, 2000. In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 89–97. Kluwer Academic Publishers: Dordrecht.

[NB01] Nel, A., G. Bechly, X. Martínez-Delclòs & G. Fleck. 2001. A new family of Anisoptera from the Upper Jurassic of Karatau in Kazakhstan (Insecta: Odonata: Juragomphidae n. fam.) Stuttgarter Beiträge zur Naturkunde Serie B (Geologie und Paläontologie) 314: 1–9.

[R02] Rasnitsyn, A. P. 2002. Special features of the study of fossil insects. In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 8–12. Kluwer Academic Publishers: Dordrecht.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.

[S02] Sinitshenkova, N. D. 2002. Ecological history of the aquatic insects. In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 388–426. Kluwer Academic Publishers: Dordrecht.

[ZBH03] Zhou, Z., P. M. Barrett & J. Hilton. 2003. An exceptionally preserved Lower Cretaceous ecosystem. Nature 421: 807–814.

Last updated: 29 May 2022.

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