(From left to right) Left, right, left interior, ventral and dorsal views of Healdia simplicissima, from Hoare & Merrill (2004).

Belongs within: Metacopina.

The Healdiidae are a group of ostracods known from the Devonian to the Lower Cretaceous. Members of the family are characterised by distinctive marginal areas with a duplicature absent or fused with the outer lamella, and the left valve overreaching the right (Benson et al. 1961).

Characters (from Benson et al. 1961): Carapace convex-backed, with suboblong to subtriangular outlines and nearly straight venters in lateral view, commonly with posterior, and less commonly with anterior sculpturing in ridges and spines; valves hinged posterodorsally; left valve larger than right valve, overlap and overreach (where present) left valve over right valve. Without separated calcified inner lamellae but with hinges and contact margins simply ridged and grooved or shouldered in platycopine fashion; adductor muscle scar circular, consisting of numerous spots generally arranged in concentric rings or rows. Most genera probably exhibit moderate sexual dimorphism.

    |--Hermiella G12
    |--Incisurella Cooper 1941 BB61
    |    `--*I. prima Cooper 1941 BB61
    |--Phreatura Jones & Kirkby 1886 BB61
    |    `--*P. concinna Jones & Kirkby 1886 BB61
    |--Daleiella Bouček 1937 M61, BB61
    |    `--*D. corbuloides (Jones & Holl 1869) [=Cythere corbuloides] BB61
    |--Robsoniella Kusnetsova in Mandelstam et al. 1956 BB61
    |    |--*R. obovata Kuznetzova in Mandelstam et al. 1956 BB61
    |    `--R. falklandensis Dingle 1984 G12
    |--Healdioides Coryell & Rozanski 1942 BB61
    |    |--*H. diversus Coryell & Rozanski 1942 BB61
    |    `--H. bradfieldi Coryell & Rozanski 1942 S82
    |--Hungarella Méhes 1911 BB61
    |    `--*H. problematica (Daday 1911) [=Bairdia problematica] BB61
    |--Healdiacypris Bradfield 1935 M61, BB61
    |    |--*H. perplexa Bradfield 1935 BB61
    |    `--H. acuminata BB61
    |--Ogmoconcha Triebel 1941 BB61
    |    |--*O. contractula [=Hungarella contractula] BB61
    |    `--O. convexa Boomer 1991 WSB93
    |--Waylandella Coryell & Billings 1932 [incl. Harltonella Bradfield 1935] BB61
    |    |--*W. spinosa Coryell & Billings 1932 BB61
    |    |--W. ardmorensis BB61
    |    `--W. obesa BB61
    |--Cyrtocyprus Coryell & Williamson 1936 BB61 [=Cyrtocypris Coryell & Williamson 1936 WSB93, BB61]
    |    |--*C. subovata Coryell & Williamson 1936 BB61
    |    |--C. chvorostanensis BB61
    |    `--C. inornata Copeland 1974 WSB93
    |--Seminolites Coryell 1928 HM04
    |    |--*S. truncatus Coryell 1928 [incl. S. elongatus] BB61
    |    |--S. ovalis Cooper 1941 HM04
    |    |--S. perforatus Harlton 1933 HM04
    |    |--S. porosus Robinson 1978 HM04
    |    `--S. symmetricus Cooper 1941 HM04
    |--Cribroconcha Cooper 1941 HM04
    |    |--*C. costata Cooper 1941 BB61
    |    |--C. alabamensis (Erlich 1964) HM04
    |    |--C. conspicua (Harlton 1929) HM04
    |    |--C. fornicata Cooper 1941 HM04
    |    |--C. perplexa Robinson 1959 HM04
    |    `--C. prolixa Hoare & Merrill 2004 HM04
    |--Microcheilinella Geis 1933 HM04 [=Microcheilus Geis 1932 non Kittl 1894 BB61]
    |    |--*M. distorta (Geis 1932) [=*Microcheilus distortus] BB61
    |    |--M. acuta Shi 1987 ZL87
    |    |--M. cordata Cooper 1941 S82
    |    |--M. monospinosa Zhang in Zhang & Liang 1987 ZL87
    |    |--M. postspinosa Chen 1958 ZL87
    |    |--M. speciosa Chen 1958 CB86
    |    |--M. subreniformis Chen 1958 ZL87
    |    `--M. venusta Chen 1958 ZL87
    `--Healdia Roundy 1926 HM04
         |--*H. simplex Roundy 1926 BB61
         |--H. ampla Roundy 1926 HM04
         |--H. asper Cooper 1946 HM04
         |--H. caneyensis Harlton 1927 HM04
         |--H. cara [incl. H. aspinosa] BB61
         |--H. cornigera (Jones & Kirby 1867) HM04
         |--H. glennensis Harlton 1927 HM04
         |--H. leguminoidea S61
         |--H. marginata Harlton 1928 HM04
         |--H. oblonga Bradfield 1935 HM04
         |--H. oklahomaensis Harlton 1927 HM04
         |--H. simplicissima Harlton 1933 HM04
         `--H. sulcata Cooper 1947 S82

*Type species of generic name indicated


[BB61] Benson, R. H., J. M. Berdan, W. A. van den Bold, T. Hanai, I. Hessland, H. V. Howe, R. V. Kesling, S. A. Levinson, R. A. Reyment, R. C. Moore, H. W. Scott, R. H. Shaver, I. G. Sohn, L. E. Stover, F. M. Swain & P. C. Sylvester-Bradley. 1961. Systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q99–Q421. Geological Society of America and University of Kansas Press.

[CB86] Chen D.-Q. & Bao H. 1986. Lower Permian ostracodes from the Chihsia Formation of Jurong and Longtan, Jiangsu Province. Acta Micropalaeontologica Sinica 3 (2): 107–132.

[G12] Guzel, M. 2012. Jurassic and Early Cretaceous ostracods from Western Australia: what they reveal about evolution of the Indian Ocean. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 849–882. Springer.

[HM04] Hoare, R. D., & G. K. Merrill. 2004. A Pennsylvanian (Morrowan) ostracode fauna from Texas. Journal of Paleontology 78 (1): 185–204.

[M61] Moore, R. C. 1961. Supplement—USSR treatise on Ostracoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q422–Q429. Geological Society of America and University of Kansas Press.

[S61] Scott, H. W. 1961. Shell morphology of Ostracoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q21–Q37. Geological Society of America and University of Kansas Press.

[S82] Shi C.-G. 1982. Some early Carboniferous ostracodes from Nylam, Xizang (Tibet). Acta Palaeontologica Sinica 21 (3): 309–314.

[WSB93] Whatley, R. C., D. J. Siveter & I. D. Boomer. 1993. Arthropoda (Crustacea: Ostracoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 343–356. Chapman & Hall: London.

[ZL87] Zhang X.-J. & Liang X.-Y. 1987. Ostracoda from the Taiyuan Formation of Xingyang and Gong Xian districts, Henan. Acta Micropalaeontologica Sinica 4 (3): 293–312.

Last updated: 11 October 2020.

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