Eteoanisoptera

Giant dragonfly Petalura gigantea, copyright Leonard Müller.


Belongs within: Anisoptera.
Contains: Hemicorduliidae, Gomphoidea, Aeshnoidea, Cavilabiata.

The clade Eteoanisoptera includes the majority of living dragonflies (excluding only the relictual genus Epiophlebia). Members of this group have relatively broad wings that are generally held flat at rest. Within the Eteoanisoptera, the Petaluroidea are a group of large dragonflies united by a number of forewing synapomorphies including a very narrow postnodal space, with many cells distal from the pterostigma, and a wide subdiscoidal triangle containing a number of cross-veins. The petaluroids are regarded as the sister clade of the remaining dragonflies in the Euanisoptera, but this latter clade is less strongly supported (Nel et al. 2001).

Characters (from Grimaldi & Engel 2005): Wings with pterostigmal brace vein (supporting vein under inner edge of pterostigma); discoidal cell divided into two triangular cells; distinctive anal loop; small, frequently darkened area of veinless membrane present at base of wings near anal region; secondary 'CuP' vein associated with expanded anal region in hind wing. Ovipositor vestigial. Naiad able to propel itself using water pressure from rectal chamber.

<==Eteoanisoptera
    |--Petaluroidea [Petalurida] GE05
    |    |--Protolindeniidae RP02
    |    |--Aktassiinae [Aktassiidae] RP02
    |    |    |--Aktassia Pritykina 1968 NB01
    |    |    `--Aeschnogomphus Handlirsch 1906 NB01
    |    `--Petaluridae TH06
    |         |--Uropetala carovei (White 1846) HL02
    |         |--Phenes raptor Rambur 1842 FN02
    |         `--Petalura TH06
    |              |--P. gigantea Leach 1815 TH06
    |              |--P. hesperia Watson 1958 TH06
    |              |--P. ingentissima Tillyard 1908 TH06
    |              |--P. litorea Theischinger 1999 TH06
    |              `--P. pulcherrima Tillyard 1913 TH06
    `--Euanisoptera [Exophytica] RP02
         |--Gomphoidea GE05
         `--+--Aeshnoidea GE05
            `--Cavilabiata GE05

Eteoanisoptera incertae sedis:
  Hemicorduliidae HL02
  Palaeaeschna S02
  Juragomphus Nel, Bechly et al. 2001 [Juragomphidae] NB01
    `--*J. karatauensis Nel, Bechly et al. 2001 NB01
  Gomphomacromiidae FB03
    |--Gomphomacromia Brauer 1864 FB03
    `--Archaeophya TH06
         |--A. adamsi Fraser 1959 TH06
         `--A. magnifica Theischinger & Watson 1978 TH06
  Libellulium longialata GE05

*Type species of generic name indicated

REFERENCES

[FB03] Fleck, G., G. Bechly, X. Martínez-Delclòs, E. Jarzembowski, R. Coram & A. Nel. 2003. Phylogeny and classification of the Stenophlebioptera (Odonata: Epiproctophora). Annales de la Societe Entomologique de France 39 (1): 55-93.

[FN02] Fleck, G., & A. Nel. 2002. The first isophlebioid dragonfly (Odonata: Isophlebioptera: Campterophlebiidae) from the Mesozoic of China. Palaeontology 45 (6): 1123-1136.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[HL02] Haacks, M., & A. Lehmann. 2002. Some observations on dragonflies (Insecta: Odonata) throughout New Zealand. Weta 24: 13-17.

[NB01] Nel, A., G. Bechly, X. Martínez-Delclòs & G. Fleck. 2001. A new family of Anisoptera from the Upper Jurassic of Karatau in Kazakhstan (Insecta: Odonata: Juragomphidae n. fam.) Stuttgarter Beiträge zur Naturkunde Serie B (Geologie und Paläontologie) 314: 1-9.

[RP02] Rasnitsyn, A. P., & L. N. Pritykina. 2002. Superorder Libellulidea Laicharting, 1781. Order Odonata Fabricius, 1792. The dragonflies. In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 97-104. Kluwer Academic Publishers: Dordrecht.

[S02] Sinitshenkova, N. D. 2002. Ecological history of the aquatic insects. In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 388-426. Kluwer Academic Publishers: Dordrecht.

[TH06] Theischinger, G. & J. Hawking. 2006. The Complete Field Guide to Dragonflies of Australia. CSIRO Publishing: Collingwood (Victoria).

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