Female Chaerilus celebensis, photographed by A. Tietz.

Belongs within: Scorpiones.
Contains: Buthoidea, Scorpionoidea.

The Orthosterni is a clade including the living scorpions and some fossil relatives, characterised by the presence of trichobothria on the pedipalp (Soleglad & Fet 2001).

<==Orthosterni [Orthosternina]
    |  i. s.: Corniops mapesii SF01
    |--Branchioscorpionina SF01
    `--Neoscorpionina [Chaeriloidea, Dionychopodes, Neoscorpii] SF01
         |  i. s.: Chactoidea L03
         |           |--Chactas Gervais 1844 B-TN08
         |           `--Palaeoeuscorpius Lourenço 2003 [Palaeoeuscorpiidae] L03
         |                `--*P. gallicus Lourenço 2003 L03
         |         Superstitionidae SF01
         |           |--Typhlochactas mitchelli SF01
         |           `--Superstitionia donensis SF01
         |         Palaeoburmesebuthus Lourenço 2002 S-BF04
         |           `--P. grimaldii Lourenço 2002 S-BF04
         |--Palaeopisthacanthidae SF01
         |    |--Compsoscorpius elegans SF01
         |    |--Cryptoscorpius americanus SF01
         |    `--Palaeopisthacanthus SF01
         |         |--P. schucherti SF01
         |         `--P. vogelandurdeni SF01
         `--+--Archaeobuthus Lourenço 2001 SF01, S-BF04 [Archaeobuthidae]
            |    `--*A. estephani Lourenço 2001 S-BF04
            `--+--+--Buthoidea SF01
               |  `--Pseudochactas Gromov 1998 [Pseudochactidae] SF01
               |       `--P. ovchinnikovi Gromov 1998 SF01
               `--+--Scorpionoidea SF01
                  `--Chaerilus Sim. 1877 T94 [Chaerilidae SF01, Chaerilini]
                       `--C. celebensis Pocock 1893 T94

*Type species of generic name indicated


[B-TN08] Botero-Trujillo, R., & J. A. Noriega. 2008. First record of the scorpion genus Microtityus from Colombia, with the description of a new species (Scorpiones, Buthidae). Journal of Arachnology 36 (2): 259-266.

[L03] Lourenço, W. R. 2003. The first scorpion fossil from the Cretaceous amber of France. New implications for the phylogeny of Chactoidea. Comptes Rendus Palevol 2: 213-219.

[S-BF04] Santiago-Blay, J. A., V. Fet, M. E. Soleglad & P. R. Craig. 2004. The second Cretaceous scorpion specimen from Burmese amber (Arachnida: Scorpiones). Journal of Systematic Palaeontology 2 (2): 147-152.

[SF01] Soleglad, M. E., & V. Fet. 2001. Evolution of scorpion orthobothriotaxy: A cladistic approach. Euscorpius 1: 1-38.

[T94] Thorell, T. 1894. Förteckning öfver Arachnider från Java och närgränsande öar, insamlade af docenten D:r Carl Aurivillius; jemte beskrifingar å några sydasiatiska och sydamerikanska Spindlar. Bihang till K. Svenska Vetenskaps-Akademiens Handlingar 20 pt. 4 (4): 1-63.

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