Ascobolus carbonarius, photographed by Miro Švidroň.
Belongs within: Pezizomycotina.
Contains: Pezizaceae,
Thecotheus, Pyronemataceae, Helvellaceae.
The Pezizales is a clade of fungi including, among others, the morels and truffles.
Characters (from Kirk
et al. 2008): Operculate discomycetes; cupfungi. Stroma absent. Ascomata apothecial or cleistothecial, rarely absent, often large, discoid, cupulate or ± globose, sometimes stalked, often brightly coloured; excipulum usually thick-walled, fleshy or membranous, composed of thin-walled pseudoparenchymatous cells. Interascal tissue of simple or moniliform paraphyses, often pigmented and swollen at the apices, absent in some cleistothecial taxa. Asci elongated, persistent, thin-walled, usually without obvious apical thickening, opening by a circular pore (operculum) or vertical split, the wall sometimes blueing in iodine, the asci cylindrical to ± globose and usually indehiscent in cleistothecial taxa. Ascospores usually ellipsoidal, aseptate, hyaline to strongly pigmented, often ornamented, usually without a sheath. Anamorphs hyphomycetous where known, usually with sympodial proliferation. Saprobes on soil and very rotten wood, some coprophilous, mycorrhizal, some hypogeous (and then mycorrhizal).
Pezizales [Otideaceae, Pezizomycetes]
|--+--Pezizaceae JK06
| `--Ascobolaceae EB03Outline
| |--Clestoiodophanus EB03Outline
| |--Cubonia EB03Outline
| |--Saccobolus EB03Outline
| |--Thecotheus EB03Outline
| `--Ascobolus JK06 [incl. Seliniella Arx & Müller 1955 RS99]
| |--A. carbonarius LK04
| |--A. crenulatus JK06
| |--A. denudatus HWB03
| |--A. immersus [incl. *Seliniella macrospora Arx & Müller 1955] RS99
| `--A. lineolatus ST99
`--+--+--Pyronemataceae JK06
| `--Sarcoscyphaceae EB03Outline
| |--Aurophora EB03Outline
| |--Chorioactis EB03Outline
| |--Desmazierella acicola EB03Outline, E99
| |--Geodina EB03Outline
| |--Kompsoscypha EB03Outline
| |--Nanoscypha EB03Outline
| |--Pseudopithyella EB03Outline
| |--Sarcoscypha coccinea JK06
| |--Thindia EB03Outline
| |--Wynnea EB03Outline
| |--Cookeina EB03Outline [incl. Boedijnopeziza EB03Notes]
| | `--C. instititia [=*Boedijnopeziza instititia] EB03Notes
| `--Pithya LK04
`--+--Caloscypha JK06 [anam. Geniculodendron EB03Notes; Caloscyphaceae]
| `--C. fulgens JK06 [anam. Geniculodendron pyriforme EB03Notes]
`--+--Helvellaceae JK06
`--+--‘Gyromitra’ melaleucoides LK04
`--+--Discinaceae EB03Outline
| |--Gymnohydnotrya EB03Outline
| |--Hydnotrya EB03Outline
| |--Pseudorhizina sphaerospora EB03Outline, RS99
| `--Gyromitra JK06
| |--G. californica JK06
| `--G. esculenta GS02
`--Morchellaceae EB03Outline
|--Disciotis venosa LK04
|--Verpa conica LK04
`--Morchella JK06
|--M. deliciosa J87
|--M. elata GS02
`--M. esculenta JK06
Pezizales incertae sedis:
Loculotuber EB03Outline
Microeurotium EB03Outline
Orcadia Sutherland 1915 EB03Outline, RS99
`--*O. ascophylli Sutherland 1915 [incl. O. pelvetiana Sutherland 1915] RS99
Inermisia aggregata JKW03
Carbomyces [Carbomycetaceae] EB03Outline
Glaziella [Glaziellaceae] EB03Outline
Karstenella [Karstenellaceae] EB03Outline
Rhizina [Rhizinaceae] EB03Outline
Ascodesmidaceae EB03Outline
|--Ascodesmis EB03Outline
`--Eleutherascus EB03Outline
Terfeziaceae H03
|--Delastria rosea H03
`--Terfezia H03
|--T. claveryi H03
`--T. leptoderma H03
Tuberaceae EB03Outline
|--Choiromyces EB03Outline
|--Dingleya EB03Outline
|--Labyrinthomyces EB03Outline
|--Reddelomyces EB03Outline
`--Tuber EB03Outline
|--T. brumale PHK96
|--T. magnatum SL02
|--T. melanosporum LS01
`--T. puberulum RS99
Sarcosomataceae EB03Outline
|--Donadinia EB03Outline
|--Galiella EB03Outline
|--Korfiella EB03Outline
|--Neournula EB03Outline
|--Plectania nigrella EB03Outline, GS02
|--Pseudoplectania EB03Outline
|--Sarcosoma EB03Outline
|--Selenaspora EB03Outline
|--Strobiloscypha EB03Outline
|--Urnula hiemalis EB03Outline, JKW03
`--Wolfina EB03Outline
*Type species of generic name indicated
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