Copelatus haemorrhoidalis, photographed by Gábor Keresztes.

Belongs within: Dytiscidae.

The Copelatinae is a speciose clade of diving beetles, most species of which are included in the genus Copelatus.

Characters (from Miller 2001): Metacoxal lines closely approximated medially (metacoxal lines lost in Lacconectus); spermatheca strongly reduced; scutellum externally visible; metatarsal claws equal in length in both sexes.

<==Copelatinae [Copelatini]
    |  i. s.: Aglymbus M01
    |--Agaporomorphus M01
    |    |--A. knischi M01
    |    `--A. perairai M01
    `--+--Lacconectus M01
       |    |--L. formosanus M01
       |    `--L. peguensis RB02
       `--Copelatus Erichson 1832 M01
            |  i. s.: C. australiae M03
            |         C. distinctus M01
            |         C. ferrugineus TDC06
            |         C. longicornis R89
            |         C. melanarius TDC06
            |         C. nakamurai Guéorguiev 1970 I92
            |         C. nomax S86
            |         C. ragazzii G89
            |         C. strinatii Guignot 1958 S86
            |         C. subterraneus Gueorguiev 1978 S86
            `--+--C. haemorrhoidalis M01
               `--+--C. glyphicus M01
                  `--C. kalaharii M01

*Type species of generic name indicated


[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880. – IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a 7: 5-72.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[M01] Miller, K. B. 2001. On the phylogeny of the Dytiscidae (Insecta: Coleoptera) with emphasis on the morphology of the female reproductive system. Insect Systematics and Evolution 32: 45-92.

[M03] Miller, K. B. 2003. The phylogeny of diving beetles (Coleoptera: Dytiscidae) and the evolution of sexual conflict. Biological Journal of the Linnaean Society 79: 359-388.

[R89] Régimbart, M. 1889. Enumération des Haliplidae, Dytiscidae et Gyrinidae recueillis par M.r le Prof. L. Balzan dans l'Amérique méridionale et description de quelques autres espèces voisines. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a 7: 256-268.

[RB02] Ribera, I., R. G. Beutel, M. Balke & A. P. Vogler. 2002. Discovery of Aspidytidae, a new family of aquatic Coleoptera. Proceedings of the Royal Society of London Series B – Biological Sciences 269: 2351-2356.

[S86] Spangler, P. J. 1986. Insecta: Coleoptera. In Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) (L. Botosaneanu, ed.) pp. 622-631. E. J. Brill / Dr. W. Backhuys: Leiden.

[TDC06] Timms, B. V., B. Datson & M. Coleman. 2006. The wetlands of the Lake Carey catchment, northeast Goldfields of Western Australia, with special reference to large branchiopods. Journal of the Royal Society of Western Australia 89 (4): 175-183.

No comments:

Post a Comment

Markup Key:
- <b>bold</b> = bold
- <i>italic</i> = italic
- <a href="">FoS</a> = FoS