Copelatus haemorrhoidalis, photographed by Gábor Keresztes.

Belongs within: Dytiscidae.

The Copelatinae is a speciose clade of diving beetles, most species of which are included in the genus Copelatus. Larvae, where known, are presumed solid feeders with serrated mandibles and a foregut including a crop.

Characters (from Miller 2001): Metacoxal lines closely approximated medially (metacoxal lines lost in Lacconectus); spermatheca strongly reduced; scutellum externally visible; metatarsal claws equal in length in both sexes.

<==Copelatinae [Copelatini]
    |  i. s.: Aglymbus M01
    |--Agaporomorphus M01
    |    |--A. knischi M01
    |    `--A. perairai M01
    `--+--Lacconectus M01
       |    |--L. formosanus M01
       |    `--L. peguensis RB02
       `--Copelatus Erichson 1832 M01
            |  i. s.: *C. posticatus (Fabricius 1801) [=Dystiscus posticatus] LM87
            |         C. ater Sharp 1882 LM87
            |         C. australiae Clark 1863 (see below for synonymy) LM87
            |         C. australis (Clark 1863) [=Celina australis; incl. Co. torcutus Guignot 1954] LM87
            |         C. bakewelli Balfour-Browne 1939 LM87
            |         C. boulevardi Watts 1978 LM87
            |         C. clarki Sharp 1882 LM87
            |         C. daemeli Sharp 1882 LM87
            |         C. distinctus M01
            |         C. divisus Balfour-Browne 1939 [=C. gentilis divisus] LM87
            |         C. elongatulus Macleay 1871 LM87
            |         C. ferrugineus Sharp 1882 [incl. C. erraticus Balfour-Browne 1939, C. lividus Sharp 1882] LM87
            |         C. gapa Watts 1978 LM87
            |         C. glyptus Guignot 1954 LM87
            |         C. gracilis Sharp 1882 LM87
            |         C. irregularis Macleay 1871 LM87
            |         C. longicornis R89
            |         C. marginatus Sharp 1882 LM87
            |         C. melanarius Sharp 1882 LM87
            |         C. nakamurai Guéorguiev 1970 I92
            |         C. nigrolineatus Sharp 1882 LM87
            |         C. nomax S86
            |         C. parvulus S62
            |         C. politus M86
            |         C. punctipennis Lea 1899 LM87
            |         C. ragazzii G89
            |         C. rasilis Lea 1899 LM87
            |         C. simplex Clark 1863 [incl. C. labratus Sharp 1882, C. victoriae Clark 1863] LM87
            |         C. strinatii Guignot 1958 S86
            |         C. subterraneus Gueorguiev 1978 S86
            |         C. tenebrosus Régimbart 1880 LM87
            |--C. haemorrhoidalis M01
            `--+--C. glyphicus M01
               `--C. kalaharii M01

Copelatus australiae Clark 1863 [incl. C. acuductus Clark 1863, C. extensus Sharp 1882, C. nigritulus Sharp 1882, C. tarsalis Balfour-Browne 1939] LM87

*Type species of generic name indicated


[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[M86] Macleay, W. 1886. The insects of the Fly River, New Guinea, "Coleoptera". Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 136–157.

[M01] Miller, K. B. 2001. On the phylogeny of the Dytiscidae (Insecta: Coleoptera) with emphasis on the morphology of the female reproductive system. Insect Systematics and Evolution 32: 45–92.

[R89] Régimbart, M. 1889. Enumération des Haliplidae, Dytiscidae et Gyrinidae recueillis par M.r le Prof. L. Balzan dans l’Amérique méridionale et description de quelques autres espèces voisines. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 256–268.

[RB02] Ribera, I., R. G. Beutel, M. Balke & A. P. Vogler. 2002. Discovery of Aspidytidae, a new family of aquatic Coleoptera. Proceedings of the Royal Society of London Series B—Biological Sciences 269: 2351–2356.

[S62] Spangler, P. J. 1962. Natural history of Plummers Island, Maryland. XIV. Biological notes and description of the larva and pupa of Copelatus glyphicus (Say) (Coleoptera: Dytiscidae). Proceedings of the Biological Society of Washington 75: 19–24.

[S86] Spangler, P. J. 1986. Insecta: Coleoptera. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 622–631. E. J. Brill/Dr W. Backhuys: Leiden.

Last updated: 30 January 2018.

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