Nassarius

Nassarius arcularius, copyright H. Zell.

Belongs within: Nassariidae.

Nassarius is a genus of small to medium-sized dog whelks with an ovate shell that is more or less heavily calloused on the columella and/or parietal wall.

<==Nassarius Duméril 1805 BR17
    |--*N. (Nassarius) arcularius (Linnaeus 1758) P61, WG71 (see below for synonymy)
    |    |--N. a. arcularius BD86
    |    `--N. a. plicatus (Roeding 1798) BD86
    |--N. (Caesia Adams & Adams 1853 non Br. 1810 [ICBN]) C75
    |    `--N. (*C.) perpinguis (Hinds 1844) C75
    |--N. (Cryptonassarius Cernohorsky 1975) C75
    |    `--N. (*C.) ephamillus (Watson 1882) (see below for synonymy) C75
    |--N. (Hima Leach in Gray 1852) ZC10, C75 [incl. Reticunassa Iredale 1936 C75]
    |    |--N. (H.) festivus (Powys 1835) ZC10
    |    |--N. (H.) fraterculus (Dunker 1860) ZC10
    |    |--N. (H.) multigranosus (Dunker 1847) ZC10
    |    `--N. (H.) pauperus (Gould 1850) ZC10 [=*Reticunassa paupera C75]
    `--N. (Varicinassa) variciferus (Adams 1851) ZC10

Nassarius incertae sedis:
  N. albescens (Dunker 1846) BW09
  N. albus (Say 1826) BC01
  N. aoteanus Finlay 1927 P61
  N. arcadioi Rolán & Hernández 2005 F05
  N. babylonicus C60
  N. bicallosus (Smith 1876) MG-H11
  N. burchardi (Dunker in Philippi 1849) HJ08
  N. caboverdensis Rolàn 1984 O96
  N. caelolineatus Nesbitt & Pitt 1986 BC01
  N. cerritensis (Arnold 1903) PP78
  N. circumcinctus (Adams 1851) O96
  N. conoidalis BD12
  N. cooperi B56
  N. coronatus (Bruguière 1789) WG71 [=Buccinum coronatum F27, Arcularia coronata F27]
  N. corpulentus (Adams 1852) [incl. Cancellaria nassiformis Lesson 1842] PH90
  N. crebricostatus (Schepman 1911) MG-H11
  N. crenulatus SP72
  N. delicatus BD12
  N. distortus (Adams 1852) C75
  N. ferrusaci Payraudeau 1826 O96
  N. gaudiosus (Hinds 1844) BW09
  N. gibbosulus (Linné 1758) O96
  N. glans (Linnaeus 1758) BD86
  N. granifer (Kiener 1834) BW09
  N. hepaticus ZLK11
  N. insculptus (Carpenter 1864) PP78
  N. johnsoni C60
  N. karinae Usticke 1971 [=N. nanus Usticke 1959 non (Adams 1852)] BC01
  N. kraussianus (Dunker 1846) HJ08
  N. limacinus (Dall 1917) S66
  N. luteostomus (Broderip & Sowerby 1829) H92
  N. monilis Kiener 1834 C75
  N. nodiferus Powys 1835 [incl. Nassa stolida Adams 1852, Nassarius stolidus] C75
  N. olivaceus (Bruguière 1789) C75
  N. pagodus ZLK11
  N. papillosus (Linnaeus 1758) BW09
  N. particeps Hedley 1915 P61
  N. pullus ZLK11
  N. pyrrhus C75
  N. reticulatus (Linnaeus 1758) D88
  N. sinusigerus BD12
  N. smitsorum Kool 1990 BC01
  N. socialis A27
  N. spiratus (Adams 1852) P61
  N. succinctus (Adams 1851) ZLK11, ZC10 [=N. (Zeuxis) succinctus ZC10]
  N. tiarula (Kiener 1841) H78
  N. trivittatus H64
  N. vibex BD12

Nassarius (*Cryptonassarius) ephamillus (Watson 1882) [=Nassa ephamilla, Alectrion ephamilla, Nassa (Tritia) ephamilla; incl. Arcularia dipsacoides Hedley 1907, Nassarius dipsacoides, Reticunassa dipsacoides, Nassa dissimilis Watson 1886, Alectrion dissimilis, Nassarius dissimilis, Reticunassa flindersi Cotton & Godfrey 1938] C75

*Nassarius (Nassarius) arcularius (Linnaeus 1758) P61, WG71 [=Buccinum arcularia P61, Arcularia arcularia H09, B. (Nassa) arcularia G20]

*Type species of generic name indicated

REFERENCES

[A27] Allan, R. S. 1927. The geology and palaeontology of the Lower Waihao Basin, south Canterbury, New Zealand. Transactions and Proceedings of the New Zealand Institute 57: 265–309.

[BD12] Bandel, K., & D. T. Dockery III. 2012. Protoconch characters of Late Cretaceous Latrogastropoda (Neogastropoda and Neomesogastropoda) as an aid in the reconstruction of the phylogeny of the Neogastropoda. Freiberger Forschungshefte C542 (psf 20): 93–128.

[BD86] Barash, A., & Z. Danin. 1986. Further additions to the knowledge of Indo-Pacific Mollusca in the Mediterranean Sea (Lessepsian migrants). Spixiana 9 (2): 117–141.

[B56] Berry, S. S. 1956. Mollusca dredged by the Orca off the Santa Barbara Islands, California, in 1951. Journal of the Washington Academy of Sciences 46 (5): 150–157.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[BW09] Bryce, C., & C. Whisson. 2009. The macromolluscs of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, Western Australia. Records of the Western Australian Museum Supplement 77: 177–208.

[C75] Cernohorsky, W. O. 1975. The taxonomy of some Indo-Pacific Mollusca. Part 3. With descriptions of new taxa and remarks on an Ecuadorian fossil species of Turridae. Records of the Auckland Institute and Museum 12: 213–234.

[C60] Cox, L. R. 1960. Gastropoda: general characteristics of Gastropoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I84–I169. Geological Society of America, and University of Kansas Press.

[D88] D’Asaro, C. N. 1988. Micromorphology of neogastropod egg capsules. Nautilus 102 (4): 134–148.

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[H64] Hampson, G. R. 1964. Redescription of a commensal pelecypod, Rochefortia cuneata, with notes on ecology. Nautilus 77 (4): 125–128.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[H92] Herbert, D. G. 1992. Revision of the Umboniinae in southern Africa and Mozambique (Mollusca: Prosobranchia: Trochidae). Annals of the Natal Museum 33 (2): 379–459.

[H78] Houston, R. S. 1978. Notes on the spawning and egg capsules of two prosobranch gastropods: Nassarius tiarula (Kiener, 1841) and Solenosteira macrospira (Berry, 1957). Veliger 20 (4): 367–368.

[HJ08] Huisman, J. M., D. S. Jones, F. E. Wells & T. Burton. 2008. Introduced marine biota in Western Australian waters. Records of the Western Australian Museum 25 (1): 1–44.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[O96] Oliverio, M. 1996. Contrasting developmental strategies and speciation in N.E. Atlantic prosobranchs: a preliminary analysis. In: Taylor, J. D. (ed.) Origin and Evolutionary Radiation of the Mollusca pp. 261–266. Oxford University Press: Oxford.

[PH90] Petit, R. E., & M. G. Harasewych. 1990. Catalogue of the superfamily Cancellarioidea Forbes and Hanley, 1851 (Gastropoda: Prosobranchia). Nautilus 103 (Suppl. 1): 1–69.

[PP78] Poorman, F. L., & L. H. Poorman. 1978. Additional molluscan records from Bahía de Los Angeles, Baja California Norte. Veliger 20 (4): 369–374.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4^th ed. Whitcombe and Tombs Limited: Christchurch.

[S66] Shasky, D. R. 1966. Range and bathymetric extensions for Olivella inconspicua and Nassarius limacinus. Nautilus 80 (1): 35–36.

[SP72] Smythe, K. R., & W. W. A. Phillips. 1972. Some observations on the fauna of the Maldive Islands (Indian Ocean) part VIII. Marine shells. Journal of the Bombay Natural History Society 69 (2): 290–296.

[WG71] Wilson, B. R., & K. Gillett. 1971. Australian Shells: illustrating and describing 600 species of marine gastropods found in Australian waters. A. H. & A. W. Reed: Sydney.

[ZC10] Zhang, S., & Z. Chen. 2010. Species composition and faunal characteristics of Buccinacea (Mollusca, Gastropdoa) of the Yellow Sea and Bohai Sea. In: China-Russia Bilateral Symposium: Proceedings of the China-Russia Bilateral Symposium of "Comparison on Marine Biodiversity in the Northwest Pacific Ocean", 10–11 October 2010, Qingdao (China) pp. 117–118. Institute of Oceanology, Chinese Academy of Sciences; A. V. Zhirmunsky Institute of Marine Biology, Far East Branch of the Russian Academy of Sciences.

[ZLK11] Zou, S., Q. Li & L. Kong. 2011. Additional gene data and increased sampling give new insights into the phylogenetic relationships of Neogastropoda, within the caenogastropod phylogenetic framework. Molecular Phylogenetics and Evolution 61: 425–435.

Zemitrella

Zemitrella chaova, copyright G. & Ph. Poppe.

Belongs within: Columbellidae.

Zemitrella is a New Zealand genus of columbellids with a high-spired, almost straight-sided shell and a narrow, parallel-sided aperture.

<==Zemitrella Finlay 1927 P61
    |--*Z. sulcata (Hutton 1873) P61 (see below for synonymy)
    |--Z. annectens Powell 1937 P61
    |--Z. attenuata Powell 1940 P61
    |--Z. benthicola Dell 1956 P61
    |--Z. chaova (Reeve 1859) P61
    |--Z. constans Powell 1955 P61
    |--Z. curvirostris Powell 1937 P61
    |--Z. daemona (Webster 1906) P61 [=Atilia daemona F27]
    |--Z. fallax Powell 1940 P61
    |--Z. finlayi Powell 1933 P61
    |--Z. inconspicua (Marshall 1918) [=Mitrella inconspicua] F27
    |--Z. laevigata (Suter 1908) P61
    |--Z. laevirostris Powell 1940 P61
    |--Z. lincolnensis M54
    |--Z. menkeana M54
    |--Z. pseudomarginata (Suter 1908) P61 [=Mitrella pseudomarginata F27; incl. Atilia biconica F27]
    |--Z. pulla M54
    |--Z. regis Powell 1940 P61
    |--Z. rosea (Hutton 1873) P61 [=Obeliscus roseus F27, Mitrella rosea F27; incl. Alcira sanguinea F27]
    |--Z. semiconvexa M54
    |--Z. sericea Powell 1937 P61
    |--Z. stephanophora (Suter 1908) P61
    |--Z. turgida Powell 1937 P61
    |--Z. websteri (Suter 1913) P61
    `--Z. whangaroaensis Dell 1956 P61

*Zemitrella sulcata (Hutton 1873) P61 [=Lachesis sulcata P61, Columbella huttoni Suter 1904 F27; incl. C. inconstans Suter 1906 F27, C. varians Sowerby 1832 F27, H09, Pyrene varians H09]

*Type species of generic name indicated

REFERENCES

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[M54] Macpherson, J. H. 1954. Molluscs (sea shells and snails). In: Willis, J. M. (ed.) The Archipelago of the Recherche pp. 55–63. Australian Geographical Society: Melbourne.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4^th ed. Whitcombe and Tombs Limited: Christchurch.

Liratilia

Liratilia conquisita, copyright Te Papa Tongarewa.

Belongs within: Columbellidae.

Liratilia is a New Zealand genus of columbellids with high-spired shells with a prominent sculpture of spiral cords or keels, and a narrow, parallel-sided aperture.

<==Liratilia Finlay 1927 P61
    |--*L. conquisita (Suter 1907) [=Daphnella conquisita] P61
    |    |--L. c. conquisita P61
    |    |--L. c. chathamensis Dell 1956 P61
    |    `--L. c. nestoris Fleming 1948 P61
    |--L. angulata (Suter 1908) P61 [=Alcira angulata F27]
    |--L. compta Powell 1930 P61
    |--L. elegantula Powell 1937 P61
    |--L. eremita (Murdoch & Suter 1906) P61 [=Pleurotoma (Leucosyrinx) eremita F27]
    |--L. gracilis Powell 1940 P61
    |--L. isolata Dell 1956 P61
    |--L. sinuata Powell 1937 P61
    `--L. subnodosa Powell 1934 P61

*Type species of generic name indicated

REFERENCES

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4^th ed. Whitcombe and Tombs Limited: Christchurch.

Cominella

Quoy's whelk Cominella quoyana, copyright Icolmer.

Belongs within: Buccinoidea.

Cominella is an Australasian genus of whelks whose shell has a high and acuminate spire, a broad notch for the anterior canal, spiral ribs or lirae on the aperture, and a smooth columella (Wilson & Gillett 1971).

<==Cominella Gray 1850 [Cominellini] WG71
    |--C. (Cominella) P61
    |    |--*C. (C.) testudinea (Bruguière 1789) [=Buccinum testudineum] BR17
    |    |--C. (C.) maculosa (Martyn 1784) P61 [=Buccinum maculosum F27a]
    |    `--C. (C.) virgata Adams & Adams 1853 P61
    |         |--C. v. virgata P61
    |         `--C. v. brookesi Powell 1952 P61
    |--C. (Acominia Finlay 1927) P61
    |    |--C. (*A.) adspersa (Bruguière 1789) [=Buccinum adspersum] P61
    |    |    |--C. a. adspersa [incl. C. nimia] P61
    |    |    `--C. a. melo (Lesson 1840) P61
    |    `--C. (A.) hendersoni Marwick 1926 F27a
    |--C. (Cominista Finlay 1927) P61
    |    `--C. (*C.) glandiformis (Reeve 1847) P61 [=Buccinum glandiforme P61; incl. B. luridum F27a]
    |--C. (Cominula Finlay 1927 P61
    |    |--C. (*C.) quoyana Adams 1854 P61
    |    |--C. (C.) euthriaformis Powell 1929 P61
    |    |--C. (C.) exsculpta [=Procominula exsculpta] A27
    |    `--C. (C.) youngi Powell 1940 P61
    `--C. (Eucominia Finlay 1927) P61
         |--C. (*E.) nassoides (Reeve 1846) [=Buccinum nassoides] P61
         |    |--C. n. nassoides P61
         |    `--C. n. fouveauxana Powell 1946 P61
         |--C. (E.) elegantula Finlay 1926 P61
         |--C. (E.) ellisoni P61
         |    |--C. e. ellisoni P61
         |    `--C. e. consobrina Powell 1933 P61
         |--C. (E.) haroldi Powell 1946 P61
         |--C. (E.) iredalei Finlay 1928 P61
         |--C. (E.) marlboroughensis Powell 1946 P61
         |--C. (E.) mirabilis PG98
         |    |--C. m. mirabilis PG98
         |    `--C. m. powelli PG98
         |--C. (E.) nodicincta (von Martens 1878) P61
         |--C. (E.) olsoni Dell 1956 P61
         `--C. (E.) otakauica Powell 1946 P61

Cominella incertae sedis:
  C. carinata H86
  C. compacta Marwick 1926 F27a
  C. costata F27a
  C. eburnea Reeve 1846 WG71
  C. errata Finlay 1924 [=Buccinum carinatum Hutton 1873 (preoc.)] F27b
  C. inflata H86
  C. lineolata Lamarck 1809 WG71 [incl. C. maura Adams 1855 H08]
  C. maculata H86
  C. papyracea C79
  C. robinsoni H86
  C. subnodosa H79
  C. zealandica (Reeve 1846) F27a

*Type species of generic name indicated

REFERENCES

[A27] Allan, R. S. 1927. The geology and palaeontology of the Lower Waihao Basin, south Canterbury, New Zealand. Transactions and Proceedings of the New Zealand Institute 57: 265–309.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[C79] Cook, P. L. 1979. Some problems in interpretation of heteromorphy and colony integration in Bryozoa. In: Larwood, G., & B. R. Rosen (eds) Biology and Systematics of Colonial Organisms pp. 193–210. Academic Press: London.

[F27a] Finlay, H. J. 1927a. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[F27b] Finlay, H. J. 1927b. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488–533.

[H79] Haast, J. von. 1879. Geology of the Provinces of Canterbury and Westland, New Zealand. A report comprising the results of official explorations. "Times" Office: Christchurch.

[H08] Hedley, C. 1908. Studies on Australian Mollusca. Part X. Proceedings of the Linnean Society of New South Wales 33: 456–489, pls 7–10.

[H86] Hutton, F. W. 1886. The Mollusca of the Pareora and Oamaru systems of New Zealand. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 205–237.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4^th ed. Whitcombe and Tombs Limited: Christchurch.

[PG98] Probert, P. K., & S. L. Grove. 1998. Macrobenthic assemblages of the continental shelf and upper slope off the west coast of South Island, New Zealand. Journal of the Royal Society of New Zealand 28: 259–280.

[WG71] Wilson, B. R., & K. Gillett. 1971. Australian Shells: illustrating and describing 600 species of marine gastropods found in Australian waters. A. H. & A. W. Reed: Sydney.

Buccinulum (Euthrena)

Buccinulum robustum, copyright Auckland War Memorial Museum.

Belongs within: Buccinoidea.

The subgenus Euthrena of the genus Buccinulum is a group of whelks bearing a shell with flattened whorls with a sculpture of low broad axials, and a very small protoconch that is weakly axially ribbed at an early stage, the transition from juvenile to adult sculpture being a very well-marked brephic stage of narrow strong axial and spiral cords producing a coarse reticulation with the points of intersection marked as blunt gemmules (Finlay 1927).

<==Buccinulum (Euthrena Iredale 1918) P61
    |--B. (*E.) vittatum (Quoy & Gaimard 1833) P61 [=Fusus vittatus P61, *Euthrena vittata F27]
    |    |--B. v. vittatum P61
    |    `--B. v. costulatum (Suter 1913) P61 [=Euthrena costulata F27]
    |--B. (E.) bicinctum (Hutton 1873) P61 [=Fusus bicinctus F27, Euthrena bicincta F27]
    |--B. (E.) colensoi (Suter 1908) P61
    |--B. (E.) flavescens (Hutton 1884) P61
    |--B. (E.) heteromorphum Powell 1929 P61
    |    |--B. h. heteromorphum P61
    |    `--B. h. bucknilli Powell 1929 P61
    |--B. (E.) kaikouraense Powell 1929 P61
    |--B. (E.) littorinoides (Reeve 1846) P61
    |--B. (E.) maketuense Powell 1929 P61
    |--B. (E.) mariae Powell 1940 P61
    |--B. (E.) martensianum (Hutton 1878) P61 [=Euthrena martensiana F27]
    |--B. (E.) motutaraense Powell 1929 P61
    |--B. (E.) robustum Powell 1929 P61
    |--B. (E.) strebeli (Suter 1908) P61
    |    |--B. s. strebeli P61
    |    |--B. s. exsculptum Powell 1929 P61
    |    `--B. s. mestayerae Powell 1929 P61
    `--B. suteri Powell 1934 P61

*Type species of generic name indicated

REFERENCES

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4^th ed. Whitcombe and Tombs Limited: Christchurch.

Turbinelloidea

Vasum ceramicum, photographed by Frédéric Ducarme.

Belongs within: Neogastropoda.
Contains: Volutomitridae, Columbariidae, Ptychatractidae, Costellariidae.

The Turbinelloidea are a clade of neogastropods supported by molecular phylogenetic analysis. The Turbinellidae, chank and vase shells, are generally large gastropods with a thick, spindle-shaped shell (the largest known gastropod, Syrinx aruanus, belongs to this family). They possess a long, thin proboscis that folds into a non-evaginable proboscis sheath (Harasewych & Petit 1989).

Turbinelloidea BR17
    |--+--Volutomitridae FP15
    |  `--+--Columbariidae FP15
    |     `--+--Ptychatractidae FP15
    |        `--Costellariidae FP15
    `--Turbinellidae [Vasidae] FP15
         |  i. s.: Tudivasum MG-H11
         |           |--T. inermis (Angas 1878) MG-H11
         |           `--T. spinosa (Adams & Adams 1863) MG-H11
         |         Surculina Dall 1908 BK11, P66 [incl. Phenacoptygma Dall 1918 O27]
         |           |--*S. blanda (Dall 1908) [=Daphnella (*Surculina) blanda] P66
         |           `--S. cortezi (Dall 1908) P66 [=Daphnella (Surculina) cortezi P66, *Phenacoptygma cortezi O27]
         |--Tudicula Adams & Adams 1864 ES88 [=Tudicla Röding 1798 BR05; Tudiclidae, Tudiclinae BR05, Tudiculinae]
         |    |--*T. spirillus (Linnaeus 1767) [=Murex spirillus] BR17
         |    |--T. armigera Adams 1855 WG71
         |    |--T. inermis Angas 1878 WG71
         |    |--‘Tudicla’ monheimi (Müller 1851) TTE93
         |    |--T. rasilistoma Abbott 1959 ES88
         |    |--T. spinosa Adams & Adams 1863 WG71
         |    `--T. zanzibarica Abbott 1958 ES88
         |--Vasum Röding 1798 ES88 [=Cynodonta Schumacher 1817 BR05; Cynodontidae, Cynodontinae, Vasinae ES88]
         |    |  i. s.: V. cassiforme (Kiener 1840) ES88
         |    |         V. crosseanum (Souverbie 1875) ES88
         |    |         V. floridanum McGinty 1940 P89
         |    |         V. globulum (Lamarck 1816) [incl. V. globulum antiguensis Usticke 1969] BC01
         |    |         V. humerosum Vaughan 1896 DK10
         |    |         V. lactisfloris Ferrario 1983 BC01
         |    |         V. rhinoceros (Gmelin 1791) ES88
         |    |         V. stephanti Emerson & Sage 1988 ES88
         |    |         V. tubiferum (Anton 1839) ES88
         |    |         V. turbinellus (Linnaeus 1758) ES88 (see below for synonymy)
         |    |--*V. (Vasum) ceramicum (Linnaeus 1758) [=Murex ceramicus, *Cynodonta ceramica] BR17
         |    `--V. (Altivasum Hedley 1914) DK10, WG71
         |         `--V. (A.) flindersi Verco 1914 DK10, WG71
         `--Turbinellinae [Xancidae, Xancinae] HP89
              |--Syrinx Röding 1798 HP89
              |    `--*S. aruanus (Linnaeus 1758) HP89 (see below for synonymy)
              |--Scolymus Swainson 1835 [Scolyminae] BR17
              |    `--‘Turbinella’ angulata (Lightfoot 1786) (see below for synonymy) BR17
              `--Xancus Röding 1798 BR05 [=Turbinella Lamarck 1799 BR05; incl. Buccinella Perry 1811 PH90]
                   |--*X. pyrum (Linnaeus 1767) BR17 (see below for synonymy)
                   |--‘Turbinellus’ acuminatus C64
                   |--‘Cancellaria’ alabamensis Gabb 1860 [=Turbinella alabamensis] PH90
                   |--‘Turbinella’ caestus C64
                   |--‘Turbenella’ ceratus C64
                   |--‘Turbinellus’ infundibulum [=Murex infundibulum; incl. T. filosus, T. gibbulus] G79
                   |--‘Turbinella’ iricolor Hombr. & Jacquinot 1853 H09
                   |--‘Turbinellus’ nassa [incl. T. brasilianus, T. cingulifera, Murex knorri, T. knorri, T. rudis] G79
                   |--‘Turbinella’ scolymoides Dall 1890 P89
                   `--‘Turbinella’ subnassatula Souverbie 1872 H09

*Syrinx aruanus (Linnaeus 1758) HP89 [=Murex aruanus HP89, Busycon aruanus HP89, Fulgur aruanus HP89, Megalatractus aruanus WG71; incl. Fusus proboscidiferus Lamarck 1816 HP89, Me. proboscidiferus HP89]

‘Turbinella’ angulata (Lightfoot 1786) [incl. T. umbilicaris Lamarck 1816, *Scolymus umbilicaris] BR17

Vasum turbinellus (Linnaeus 1758) ES88 [=Voluta turbinella H09; incl. Turbinella cornigera Lamarck 1822 BR17, *Scolymus cornigerus BR17, Vasum turbinellus var. cornigerum SP72]

*Xancus pyrum (Linnaeus 1767) BR17 [=Voluta pyrum BR17, Murex (*Turbinella) pyrum non M. pyrum Linnaeus 1758 G20, BR17; incl. *Buccinella caerulea Perry 1811 PH90]

*Type species of generic name indicated

REFERENCES

[BK11] Bouchet, P., Y. I. Kantor, A. Sysoev & N. Puillandre. 2011. A new operational classification of the Conoidea (Gastropoda). Journal of Molluscan Studies 77: 273–308.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[DK10] Darragh, T. A., & G. W. Kendrick. 2010. Eocene molluscs from the Merlinleigh Sandstone, Carnarvon Basin, Western Australia. Records of the Western Australian Museum 26 (1): 23–41.

[ES88] Emerson, W. K., & W. E. Sage III. 1988. A new species of Vasum (Gastropoda: Turbinellidae) from off Somalia. Nautilus 102 (1): 36–39.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[G79] Guppy, R. J. L. 1879. First sketch of a marine invertebrate fauna of the Gulf of Paria and its neighbourhood. Part I.—Mollusca. Journal of Conchology 2: 151–172.

[HP89] Harasewych, M. G., & R. E. Petit. 1989. The nomenclatural status and phylogenetic affinities of Syrinx aruanus Linné, 1758 (Prosobranchia: Turbinellidae). Nautilus 103 (2): 83–84.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. 2 pt 1. Stanford University Press: Stanford University (California).

[PH90] Petit, R. E., & M. G. Harasewych. 1990. Catalogue of the superfamily Cancellarioidea Forbes and Hanley, 1851 (Gastropoda: Prosobranchia). Nautilus 103 (Suppl. 1): 1–69.

[P89] Petuch, E. J. 1989. New species of Malea (Gastropoda: Tonnidae) from the Pleistocene of southern Florida. Nautilus 103 (3): 92–95.

[P66] Powell, A. W. B. 1966. The molluscan families Speightiidae and Turridae: An evaluation of the valid taxa, both Recent and fossil, with lists of characteristic species. Bulletin of the Auckland Institute and Museum 5: 1–184, pls 1–23.

[SP72] Smythe, K. R., & W. W. A. Phillips. 1972. Some observations on the fauna of the Maldive Islands (Indian Ocean) part VIII. Marine shells. Journal of the Bombay Natural History Society 69 (2): 290–296.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

[WG71] Wilson, B. R., & K. Gillett. 1971. Australian Shells: illustrating and describing 600 species of marine gastropods found in Australian waters. A. H. & A. W. Reed: Sydney.

Buccinoidea

Pareuthria fuscata, copyright Manuel Caballer.

Belongs within: Neogastropoda.
Contains: Pisania, Cantharus, Colubrariidae, Melongena, Nassariidae, Columbellidae, Fasciolariidae, Engina, Busyconinae, Beringius, Siphonaliinae, Aeneator, Penion, Buccinulum (Euthrena), Buccinulum (Evarnula), Cominella, Colini, Buccinum, Volutopsius.

The Buccinoidea are a diverse clade of neogastropods characterised by the absence of accessory salivary glands. Members are ecologically diverse; the majority are scavengers or predators but some are parasites or herbivores. The Buccinidae, whelks, are a carnivorous family, including both predators and scavengers, found predominantly in temperate or boreal waters of the Northern Hemisphere. They are morphologically diverse, varying from small to large, commonly with a wide aperture and long proboscis. Eggs may be laid singly or in masses with development often being direct. The Melongenidae, giant whelks, are a mostly tropical group of medium-sized to very large scavengers with an elongate aperture and long head. The Pisaniidae are a predominantly tropical group of mostly medium-sized species with a high aperture, typically with a thickened outer lip, and usually with an axial sculpture of strong, closely-spaced ribs. Belomitra is a North Atlantic genus with moderately large (16–22 mm), thin shells with a tall spire and narrow whorls that are distinctly angulate above the middle (Powell 1966).

Buccinoidea [Beringioidea] BR17
    |--Belomitra Fischer 1883 BR17 [incl. Pleurobela Monterosato in Locard 1897 P66; Belomitridae FP15]
    |    |--*B. paradoxa Fischer 1883 BR17
    |    |--B. bouteti FP15
    |    |--B. delicatulina (Locard 1897) [=Clionella delicatulina] P66
    |    |--B. fischeri Locard 1897 P66
    |    |--B. lyrata Locard 1897 [=Pleurobela lyrata] P66
    |    `--B. spelta (Locard 1897) [=Pleurotoma spelta, *Pleurobela spelta] P66
    `--+--Melongenidae [Cassidulina, Cassidulinae, Melongeninae] FP15
       |    |--Melongena BR05
       |    |--Tantunia clathrata TTE93
       |    |--Volegalea Iredale 1938 WG71
       |    |    `--V. wardiana Iredale 1938 WG71
       |    |--Rhombopsis Gardner 1916 [=Neptunella Meek 1864 non Gray 1854] P66
       |    |    `--*R. newberryi (Meek & Hayden 1857) [=Fusus newberryi, *Neptunella newberryi] P66
       |    |--Bruclarkia P66
       |    |    `--‘Fusus’ corpulentus Conrad 1849 [=Priscofusus corpulentus] P66
       |    |--Heligmotoma Mayer-Eymar 1896 [Heligmotomidae] BR05
       |    |    `--*H. nilotica (Mayer-Eymar 1896) [=Melongena nilotica] BR17
       |    |--Volema Röding 1798 [Volemidae] BR05
       |    |    |--*V. paradisiaca Röding 1798 BR17
       |    |    `--V. cornuta [=Galeodes (Volema) cornuta] H62
       |    |--Hemifusus ZLK11
       |    |    |--H. colosseus ZLK11
       |    |    |--H. ternatanus ZLK11
       |    |    `--H. tuba ZLK11
       |    `--‘Galeodes’ Röding 1798 non Olivier 1791 BR05
       |         |--*G. melongena (Linnaeus 1758) [=Murex melongena] BR17
       |         |--G. cochlidium (Linnaeus 1758) [=Murex cochlidium] H09
       |         `--G. pricei (Smith 1887) [=Fusus pricei] H09
       `--+--Nassariidae FP15
          |--+--Columbellidae ZLK11
          |  `--Fasciolariidae FP15
          `--Buccinidae [Ceryciidae, Cominellidae, Cominellina, Cominellinae] FP15
               |  i. s.: Manaria kuroharai Azuma 1960 AV03, MG-H11
               |         Eosipho AV03
               |         Anna Risso 1826 P66
               |           `--*A. massena Risso 1826 P66
               |         Savatieria Rochebrune & Mabille 1885 P66
               |           `--*S. frigida Rochebrune & Mabille 1885 P66
               |         Troschelia berniciensis (King 1846) OT04
               |         Neocola Finlay 1926 F26
               |           |--*N. beta [=Austrofusus beta] F26
               |           `--N. cliftonensis (Marwick 1926) [=Aethocola cliftonensis] F26
               |         Zelandiella Finlay 1926 DK10
               |           |--*Z. subnodosa [=Neptunea subnodosa] F26
               |           `--Z. fatua Finlay 1926 DK10
               |         Nassicola Finlay 1926 F26
               |           `--*N. costata F26 [=Neptunea costata F26, N. (Sipho) costatus H79]
               |         Zeapollia Finlay 1926 F26
               |           |--*Z. acuticingulata (Suter 1917) [=Tritonidea acuticingulata] F26
               |           |--Z. colensoi [=Tritonidea colensoi] F26
               |           `--Z. purpuroides (Johnston 1879) [=Ricinula purpuroides] F26
               |         Tritonidea F27
               |           |--T. australis H08
               |           `--T. fusiformis Martin 1883 F27
               |         Tasmeuthria Iredale 1925 F27
               |           |--*T. clarkei [=Siphonalia clarkei] F27
               |           `--T. glirina (Blainv. 1832) (see below for synonymy) F27
               |         Josepha tasmanica M54
               |         Engina BC01
               |         Nassaria MG-H11
               |           |--N. acuminata (Reeve 1844) MG-H11
               |           |--N. bitubercularis (Adams 1855) [=Hindsia bitubercularis] H09
               |           |--N. problematica (Iredale 1936) MG-H11
               |           `--N. solida Kuroda & Habe 1961 MG-H11
               |         Dolicholatirus Bellardi 1883 MG-H11, F27
               |           `--*D. bronni [=Turbinella bronni] F27
               |         Granulifusus kiranus Shuto 1958 MG-H11
               |         Pseudolatirus pallidus Kuroda & Habe 1961 MG-H11
               |         Bailya Smith 1944 BC01
               |           |  i. s.: B. milleri Usticke 1959 BC01
               |           `--B. (Parabailya Watters & Finlay 1989) BC01
               |                `--B. (*P.) weberi (Watters 1983) [=Caducifer (Monostiolium) weberi] BC01
               |         Caducifer decapitata (Reeve 1844) [incl. Pisania englerti Hertlein 1960] BC01
               |         Monostiolum BC01
               |           |--M. biliratum (Reeve 1846) [=Caducifer (Monostiolum) biliratus] PP78
               |           `--M. rosewateri Watters & Finlay 1989 BC01
               |         ‘Paradoxa’ Fernandes & Rolán 1990 nec Marshall 1896 nec Mattir. 1935 (ICBN) BC01
               |           |--*P. confirmata Fernandes & Rolán 1990 BC01
               |           `--P. thomensis Fernandes & Rolán 1990 BC01
               |         Antimitra Iredale 1917 BK11, P66 [incl. Mitromorpha Adams 1865 non Carpenter 1865 P66]
               |           |--*A. aegrota (Reeve 1845) [=Pleurotoma aegrota] P66
               |           |--A. crenulata (Pease 1867) [=Daphnella crenulata] P66
               |           |--‘Mitromorpha’ filosa Carpenter 1865 nec Columbella filosa Dujardin 1837 (not preoc. as in homonymous gen.) nec Daphnella filosa Carpenter 1864 (do.) O27
               |           |--‘Mitromorpha’ gracilior Hemphill 1884 O27
               |           |    |--M. g. gracilior O27
               |           |    `--M. g. intermedia Arnold 1903 O27
               |           |--‘*Mitromorpha’ gracilis O27
               |           `--A. lirata (Adams 1865) [=Mitromorpha lirata] P66
               |         Bathyclionella Kobelt 1905 BK11, P66 [incl. Cryptomitra Dall 1924 P66]
               |           |--*B. quadruplex (Watson 1881) BW80 [=Pleurotoma (Clionella) quadruplex P66]
               |           |--B. brachytoma (Schepman 1913) [=Surcula brachytoma] P66
               |           |--B. climacella (Dall 1895) [=Pleurotoma climacella, Pleurotomella climacella] P66
               |           `--B. paschalis (Thiele 1925) [=Mangelia paschalis] P66
               |         Jumala BK11
               |         Trachypollia Woodring 1928 BD12
               |           `--*T. sclera Woodring 1928 RD75
               |         Fax Iredale 1925 P61
               |           |--*F. tabidus [=Phos tabidus] P61
               |           |--F. mirabilis (Powell 1929) P61
               |           |    |--F. m. mirabilis P61
               |           |    |--F. m. canturiensis Dell 1951 P61
               |           |    |--F. m. nuptialis Dell 1953 P61
               |           |    `--F. m. powelli Fleming 1948 P61
               |           |--F. otagoensis (Finlay 1927) P61
               |           `--F. tenuicostatus F26
               |         Austrobuccinum Stilwell & Zinsmeister 1992 C96
               |         Pseudofax JB12
               |         Bulliopsis Conrad 1862 P12
               |           `--B. bowlerensis Allmon 1990 P12
               |         Hesperisternia petuchi Vermeij 2006 P12
               |         Celatoconus P12
               |         Ptychosalpinx P12
               |         Amycla C64
               |           |--A. californiana Gask. 1851 C64
               |           |--A. chrysalloidea Carpenter 1864 C64
               |           |--A. corniculum PP64
               |           |--A. gouldiana [=Columbella gouldiana, C. gouldii Agass. 1858 (n. n.)] C64
               |           |--A. minor C64
               |           |--A. tuberosa Carpenter 1864 C64
               |           `--A. undata Carpenter 1864 C64
               |         Pseudoneptunea Kobelt 1882 C75
               |           |--*P. varicosa (Roeding 1798) (see below for synonymy) C75
               |           `--P. inflata (Oostingh 1941) [=Siphonalia (Pseudoneptunea) inflata] C75
               |--Busyconinae BR05
               |--Beringius BR05
               |--Siphonaliinae BR05
               |--Donovaniinae BR17
               |    |--Syntagma Iredale 1918 P66 [incl. Lachesis Risso 1826 non Daudin 1803 BR05; Lachesinae]
               |    |    |--*S. minimum Montagu 1803 (see below for synonymy) P66
               |    |    `--‘*Lachesis’ mamillata Risso 1826 BR17
               |    `--Chauvetia Monterosato 1884 P66, BR05 (see below for synonymy)
               |         |--*C. mamillata (Risso 1826) [=Nesaea mamillata, *Donovania mamillata] BR17
               |         |--C. candidissima [=Buccinum candidissima, Nesaea candidissima] P66
               |         |--‘Buccinum’ folineaea [=*Folineaea folineaea] P66
               |         |--‘Buccinum’ lefebvrei Maravigna 1840 P66
               |         `--C. minima PP64
               |--Buccinulinae [Buccinulidae, Buccinulini] BR05
               |    |--Aeneator P61
               |    |--Penion P61
               |    |--Antarctoneptunea BR17
               |    |--Kelletia Bayle 1884 BR17, O27
               |    |    `--*K. kelletii (Forbes 1850) [=Fusus kellettii] O27
               |    |--Ellicea Finlay 1928 P61
               |    |    |--*E. orbita [=Siphonalia orbita] P61
               |    |    `--E. recens Dell 1951 P61
               |    |--Euthrenopsis Powell 1929 P61
               |    |    |--*E. otagoensis Powell 1929 P61
               |    |    |--E. bountyensis Powell 1929 P61
               |    |    `--E. venusta Powell 1929 P61
               |    |--Euthria Gray 1850 F27a
               |    |    |--E. amorimi Fraussen 2004 F05
               |    |    |--E. interrupta Sowerby 1900 S00
               |    |    |--E. linea F27a
               |    |    `--E. plumbea [incl. E. ferrea, E. viridula] S00
               |    `--Buccinulum Deshayes 1830 BR05 [incl. Evarne F27a]
               |         |--B. (Buccinulum) P61
               |         |    |--*B. (B.) lineatum (Gmelin 1791) [=Murex lineatus] BR17
               |         |    |--B. (B.) lineum (Martyn 1784) P61
               |         |    |--B. (B.) pallidum Finlay 1928 P61
               |         |    |--B. (B.) tenuistriatum Powell 1929 P61
               |         |    `--B. (B.) waitangiensis Powell 1933 P61
               |         |--B. (Euthrena) P61
               |         `--B. (Evarnula) P61
               `--Buccininae [Buccinidia] BR05
                    |  i. s.: Cominella BR05
                    |         Parancistrolepidini (see below for synonymy) BR05
                    |           |--Parancistrolepis Azuma 1965 BR05
                    |           |    `--*P. kinoshitai (Kuroda 1931) [=Japelion kinoshitai] BR17
                    |           `--Brevisiphonia Lus 1973 [Brevisiphoniinae, Brevisiphoninae] BR05
                    |                `--*B. circumreta Lus 1973 BR17
                    |         Prosipho Thiele 1912 [Prosiphiinae, Prosiphoninae, Prosiphonini] BR05
                    |           |--*P. gaussianus Thiele 1912 BR17
                    |           |--P. cassandra F26 [=Mitromorpha cassandra H22]
                    |           |--P. pallidula F26 [=Mitromorpha pallida H22]
                    |           |--P. solidus [=Mitromorpha solida] H22
                    |           `--P. tomlini Powell 1957 [incl. P. madigani Tomlin 1948 non Hedley 1916] P61
                    |         Ancistrolepis Dall 1895 [Ancistrolepidinae, Ancistrolepidini, Ancistrolepisinae] BR05
                    |           |--*A. eucosmius (Dall 1891) [=Chrysodomus eucosmius] BR17
                    |           |    |--A. e. eucosmius O27
                    |           |    `--A. e. bicinctus Dall 1919 O27
                    |           |--A. beringianus Dall 1919 O27
                    |           |--A. californicus Dall 1919 O27
                    |           `--A. magnus Dall 1895 O27
                    |         Liomesus Stimpson 1865 (see below for synonymy) BR05
                    |           |--*L. dalei (Sowerby 1825) O27, BR17 (see below for synonymy)
                    |           |--L. nassula Dall 1901 O27
                    |           |--L. nux Dall 1877 O27
                    |           `--L. ooides Middendorff 1848 O27
                    |                |--L. o. ooides O27
                    |                `--L. o. canaliculatus (Dall 1874) [=Buccinopsis canaliculatus] O27
                    |--Colini ZLK11
                    `--+--+--Buccinini BR17
                       |  |    |--Buccinum ZLK11
                       |  |    `--Volutopsius BR17
                       |  `--Volutharpa Fischer 1856 ZLK11, BC01 [Volutharpinae]
                       |       |--*V. deshayesiana Fischer 1856 BR17
                       |       |--V. ampullacea Middendorff 1848 O27 [=Bullia ampullacea C64]
                       |       `--V. perryi (Jay 1856) FP15, BC01 [=Bullia perryi BC01, V. ampullacea perryi BC01]
                       `--+--Japelion latus (Dall 1918) ZLK11, ZC10
                          `--Pareuthria Strebel 1905 ZLK11, P61
                               |--*P. fuscata P61 [=Buccinum fuscatum P61, Cominella fuscata F26]
                               |--P. campbelli (Filhol 1880) P61 [=Cominella campbelli F26]
                               |--P. magellanica [=Cominella magellanica] F26
                               |--P. plumbea ZLK11 [=Cominella plumbea F26]
                               |--P. rosea [=Cominella rosea] F26
                               `--P. valdiviae Thiele 1925 [incl. Probuccinum tenerum Tomlin 1948 non Smith 1907] P61

Buccinoidea incertae sedis:
  Echinofulguridae BR17
    |--Echinofulgur Olsson & Harbison 1953 [Echinofulgurinae] BR05
    |    `--*E. echinatus (Dall 1890) BR17 [=Fulgur echinatus BR17, Busycon echinatum C60]
    |--Levifusus Conrad 1865 [Levifusinae] BR17
    |    `--*L. trabeatus (Conrad 1833) [=Fusus trabeatus] BR17
    `--Protobusycon Wade 1917 BD12 [Protobusyconinae BR17]
         `--*P. cretaceum (Wade 1917) [=Busycon cretaceum] BR17
  Pisaniidae [Pisaniana, Pisaniinae] ZLK11
    |--Notoficula AV03
    |--Falsimohnia AV03
    |--Pisania BR17
    |--Cantharus ZLK11
    |--Solenosteira Dall 1890 AV03
    |    `--S. macrospira (Berry 1957) H78
    |--Pusiostoma Swainson 1840 [Pusiostomatidae] BR05
    |    `--*P. mendicaria (Linnaeus 1758) BR17 [=Voluta mendicaria BR17, Engina mendicaria H09]
    |--Tromina Dall 1918 AV03, RD75
    |    `--*T. unicarinata (Philippi 1868) [=Fusus unicarinatus] RD75
    |--Procominula Finlay 1926 F26
    |    `--*P. pulchra [=Cominella pulchra] F26
    |--Burnupena Iredale 1918 F26
    |    `--*B. delalandii [=Cominella delalandii; incl. Buccinum catarracta] F26
    |--Ratifusus Iredlae 1919 B88
    |    `--R. bednalli M54
    |--Zephos Finlay 1926 F26
    |    |--*Z. cingulata [=Nassa cingulata] F26
    |    `--Z. otagoensis Finlay 1926 F26
    |--Pusio Gray in Griffith & Pidgeon 1834 [incl. Calicantharus Clark 1938] AV03
    |    |--*P. elegans Gray in Griffith & Pidgeon 1834 AV03 (see below for synonymy)
    |    |--P. articulatus (see below for synonymy) G79
    |    `--‘Pisania’ fortis Carpenter 1866 [=*Calicantharus fortis] AV03
    |--Iredalula Finlay 1927 B88
    |    |--*I. striata (Hutton 1873) P61 [=Bela striata P61, Daphnella striata H86]
    |    |--I. alticincta (Murdoch & Suter 1906) P61
    |    `--I. venusta Powell 1934 P61
    `--Lirabuccinum Vermeij 1991 AV03
         |--*L. dirum (Reeve 1846) AV03 (see below for synonymy)
         |--L. branneri (Clark & Arnold 1923) (see below for synonymy) AV03
         |--L. dalli (Clark 1918) AV03
         |--L. fuscolabiatum (Smith 1875) (see below for synonymy) AV03
         |--L. japonicum (Yokoyama 1926) [=Searlesia japonica] AV03
         |--L. miocenicum (Etherington 1931) AV03
         `--L. portlaense (Arnold 1908) AV03
  Neobuccinum eatoni (Smith 1875) ZLK11, S11
  Colubrariidae ZLK11

Chauvetia Monterosato 1884 P66, BR05 [=Nesaea Risso 1826 non Leach 1814 P66, Donovania Bucquoy, Dautzenberg & Dollfus 1883 non Leach 1814 BR17; incl. Folineaea Monterosato 1884 P66; Chauvetiinae, Donovaniini]

Liomesus Stimpson 1865 [=Buccinopsis Jeffreys 1867 non Conrad 1857; Buccinopsidae, Liomesidae, Liomesinae, Liomesini] BR05

*Liomesus dalei (Sowerby 1825) O27, BR17 [=Buccinum dalei O27, *Buccinopsis dalei BR17; incl. B. dalei var. eburnea J64]

Lirabuccinum branneri (Clark & Arnold 1923) [=Searlesia branneri; incl. S. kavranensis Sinelnikova in Gladenkov, Sinelnikova & Titova 1984, Lirabuccinum kavranense, S. kurodai Makiyama 1936, S. kurodai kinseiensis Makiyama 1936] AV03

*Lirabuccinum dirum (Reeve 1846) AV03 [=Buccinum dirum AV03, Chrysodomus dirus C64, Searlesia dira O27; incl. Fusus (Neptunaea) incisus Gld. 1849 C64, Tritonium (Fusus) sitchense Middendorff 1849 C64, Chrysodomus sitchensis C64]

Lirabuccinum fuscolabiatum (Smith 1875) [=Euthria fuscolabiata, Searlesia fuscolabiata; incl. S. constricta Dall 1918, Fusus coreanicus Smith 1879, S. coreanica, Euthria hokkaidonis Pilsbry 1901, Fusus modestus Gould 1860 non Philippi 1844, Searlesia modesta, S. shimosensis Oinomikado 1935] AV03

Parancistrolepidini [Parancistrolepididae, Parancistrolepidinae, Parancistrolepisinae] BR05

*Pseudoneptunea varicosa (Roeding 1798) [=Neptunea varicosa, Murex varicosus, Siphonalia (Pseudoneptunea) varicosa; incl. S. bantamensis Martin 1895, S. (P.) bantamensis] C75

Pusio articulatus [=Fusus articulatus; incl. Murex accinctus, Purpura accincta, Buccinum pennatum, B. plumatum] G79

*Pusio elegans Gray in Griffith & Pidgeon 1834 AV03 [=Triton (Pusio) elegans C64; incl. Pisania insignis Rve. 1846 C64]

*Syntagma minimum Montagu 1803 [=Buccinum minimum, Donovania minima; incl. B. brunneum Donovan 1804] P66

Tasmeuthria glirina (Blainv. 1832) [=Purpura glirina; incl. Pisania bednalli, Buccinum discolor Kiener 1834, Tritonidea fusiformis Verco 1896 non Martin 1883, T. fusiformis adcocki Verco 1896, T. subrubiginosa Smith 1879] F27

*Type species of generic name indicated

REFERENCES

[AV03] Amano, K., & G. J. Vermeij. 2003. Evolutionary adaptation and geographic spread of the Cenozoic buccinid genus Lirabuccinum in the North Pacific. Journal of Paleontology 77 (5): 863–872.

[BD12] Bandel, K., & D. T. Dockery III. 2012. Protoconch characters of Late Cretaceous Latrogastropoda (Neogastropoda and Neomesogastropoda) as an aid in the reconstruction of the phylogeny of the Neogastropoda. Freiberger Forschungshefte C542 (psf 20): 93–128.

[B88] Bouchet, P. 1988. Two new species of Metula (Gastropoda: Buccinidae) with a description of the radula of the genus. Nautilus 102 (4): 149–153.

[BK11] Bouchet, P., Y. I. Kantor, A. Sysoev & N. Puillandre. 2011. A new operational classification of the Conoidea (Gastropoda). Journal of Molluscan Studies 77: 273–308.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BW80] Bouchet, P., & A. Waren. 1980. Revision of the north-east Atlantic bathyal and abyssal Turridae (Mollusca, Gastropoda). Journal of Molluscan Studies Supplement 8: 1–119.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[C75] Cernohorsky, W. O. 1975. The taxonomy of some Indo-Pacific Mollusca. Part 3. With descriptions of new taxa and remarks on an Ecuadorian fossil species of Turridae. Records of the Auckland Institute and Museum 12: 213–234.

[C60] Cox, L. R. 1960. Gastropoda: general characteristics of Gastropoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I84–I169. Geological Society of America, and University of Kansas Press.

[C96] Crame, J. A. 1996. Evolution of high-latitude molluscan faunas. In: Taylor, J. D. (ed.) Origin and Evolutionary Radiation of the Mollusca pp. 119–131. Oxford University Press: Oxford.

[DK10] Darragh, T. A., & G. W. Kendrick. 2010. Eocene molluscs from the Merlinleigh Sandstone, Carnarvon Basin, Western Australia. Records of the Western Australian Museum 26 (1): 23–41.

[FP15] Fedosov, A., N. Puillandre, Y. Kantor & P. Bouchet. 2015. Phylogeny and systematics of mitriform gastropods (Mollusca: Gastropoda: Neogastropoda). Zoological Journal of the Linnean Society 175: 336–359.

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[F26] Finlay, H. J. 1926. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[F27] Finlay, H. J. 1927. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488–533.

[H79] Haast, J. von. 1879. Geology of the Provinces of Canterbury and Westland, New Zealand. A report comprising the results of official explorations. "Times" Office: Christchurch.

[H62] Häntzschel, W. 1962. Trace fossils and problematica. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt W. Miscellanea: Conodonts, Conoidal Shells of Uncertain Affinities, Worms, Trace Fossils and Problematica pp. W177–W245. Geological Society of America, and University of Kansas Press.

[H08] Hedley, C. 1908. Studies on Australian Mollusca. Part X. Proceedings of the Linnean Society of New South Wales 33: 456–489, pls 7–10.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[H22] Hedley, C. 1922. A revision of the Australian Turridae. Records of the Australian Museum 13 (6): 213–359, pls 42–56.

[H78] Houston, R. S. 1978. Notes on the spawning and egg capsules of two prosobranch gastropods: Nassarius tiarula (Kiener, 1841) and Solenosteira macrospira (Berry, 1957). Veliger 20 (4): 367–368.

[H86] Hutton, F. W. 1886. The Mollusca of the Pareora and Oamaru systems of New Zealand. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 205–237.

[J64] Jeffreys, J. G. 1864. Report of the committee appointed for exploring the coasts of Shetland by means of the dredge. Report of the British Association for the Advancement of Science 33: 70–81.

[JB12] Johnson, M. E., & B. G. Baarli. 2012. Development of intertidal biotas through Phanerozoic time. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 63–128. Springer.

[M54] Macpherson, J. H. 1954. Molluscs (sea shells and snails). In: Willis, J. M. (ed.) The Archipelago of the Recherche pp. 55–63. Australian Geographical Society: Melbourne.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[OT04] Olabarria, C., & M. H. Thurston. 2004. Patterns of morphological variation of the deep-sea gastropod Troschelia berniciensis (King, 1846) (Buccinidae) from the northeastern Atlantic Ocean. Journal of Molluscan Studies 70: 59–66.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. 2 pt 1. Stanford University Press: Stanford University (California).

[PP64] Peres, J. M., & J. Picard. 1964. Nouveau manuel de bionomie benthique de la mer Mediterranee. Recueil des Travaux de la Station Marine d'Endoume, Bulletin 31 (27): 5–137.

[P12] Petuch, E. J. 2012. Miocene asteroid impacts: proposed effects on the biogeography and extinction patterns of eastern North American gastropods. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 967–981. Springer.

[PP78] Poorman, F. L., & L. H. Poorman. 1978. Additional molluscan records from Bahía de Los Angeles, Baja California Norte. Veliger 20 (4): 369–374.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4^th ed. Whitcombe and Tombs Limited: Christchurch.

[P66] Powell, A. W. B. 1966. The molluscan families Speightiidae and Turridae: An evaluation of the valid taxa, both Recent and fossil, with lists of characteristic species. Bulletin of the Auckland Institute and Museum 5: 1–184, pls 1–23.

[RD75] Radwin, G. E., & A. D’Attilio. 1975. A catalogue of muricacean generic taxa. Transactions of the San Diego Society of Natural History 17 (20): 279–292.

[S11] Simone, L. R. L. 2011. Phylogeny of the Caenogastropoda (Mollusca), based on comparative morphology. Arquivos de Zoologia 42 (4): 161–323.

[S00] Sowerby, G. B. 1900. New species of Mollusca of the genera Voluta, Conus, Siphonalia, and Euthria. Annals and Magazine of Natural History, series 7, 5: 439–441, pl. 11.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

[WG71] Wilson, B. R., & K. Gillett. 1971. Australian Shells: illustrating and describing 600 species of marine gastropods found in Australian waters. A. H. & A. W. Reed: Sydney.

[ZC10] Zhang, S., & Z. Chen. 2010. Species composition and faunal characteristics of Buccinacea (Mollusca, Gastropdoa) of the Yellow Sea and Bohai Sea. In: China-Russia Bilateral Symposium: Proceedings of the China-Russia Bilateral Symposium of "Comparison on Marine Biodiversity in the Northwest Pacific Ocean", 10–11 October 2010, Qingdao (China) pp. 117–118. Institute of Oceanology, Chinese Academy of Sciences; A. V. Zhirmunsky Institute of Marine Biology, Far East Branch of the Russian Academy of Sciences.

[ZLK11] Zou, S., Q. Li & L. Kong. 2011. Additional gene data and increased sampling give new insights into the phylogenetic relationships of Neogastropoda, within the caenogastropod phylogenetic framework. Molecular Phylogenetics and Evolution 61: 425–435.

Last updated: 24 February 2022.