Orius niger, from Raupach et al. (2014).

Belongs within: Cimicomorpha.

The Anthocoridae, flower bugs, are a group of small bugs that mostly feed on insect eggs or haemolymph from other insects. They are characterised by the presence of a cuneus in the hemelytron, with the hemelytral membrane usually being veinless, the presence of ocelli, and an apparently three-segmented labium. Most species engage in haemocoelic insemination, with the exception of the subfamily Lasiochilinae (Carver et al. 1991).

    |  i. s.: Amphiareus YS01
    |         Bilia MM96
    |         Macrotracheliella laevis MM96
    |         Plochiocorella elongata CGW91
    |--Lasiochilus [Lasiochilinae] CGW91
    |    `--L. femoralis CGW91
    |--Lyctocorinae [Dufouriellinae, Lyctocoraria, Lyctocoridae, Xylocoraria] C61
    |    |  i. s.: Eoanthocoris cretaceus Popov 1990 SP02, RJ93
    |    |         Physopleurella CGW91
    |    |--Lyctocoris C61 [Lycotocorini W77]
    |    |    |--L. campestris (Fabricius 1794) [=Acanthia campestris] C61
    |    |    `--L. subelegans Breddin 1928 C61
    |    |--Xylocorini W77
    |    |    |--Xylocoris C61
    |    |    |    |  i. s.: X. hirsutus Carayon 1961 C61
    |    |    |    |--X. (Xylocoris) C61
    |    |    |    |    |--X. (X.) afer (Reuter 1884) [=Piezostethus afer] C61
    |    |    |    |    |--X. (X.) cursitans (Fallén 1807) [=Anthocoris cursitans] C61
    |    |    |    |    `--X. (X.) galactinus C61
    |    |    |    `--X. (Arrostus) flavipes (Reuter 1875) (see below for synonymy) C61
    |    |    `--Piezostethus galactinus W77, G89
    |    |--Almeidini W77
    |    |    |  i. s.: ‘Piezostethus’ machadoi Carvalho 1952 W77
    |    |    |--Australmeida Woodward 1977 W77
    |    |    |    `--*A. derricki (Gross 1954) [=Lasiochilus derricki] W77
    |    |    |--Almeida Distant 1910 W77
    |    |    `--Lippomanus Distant 1904 W77
    |    |--Cardiastethus C61 [Cardiastethini W77]
    |    |    |--C. affinis C61
    |    |    |--C. capensis Carayon 1961 C61
    |    |    |--C. fasciiventris C61
    |    |    |--C. megalophthalmus C61
    |    |    |--C. poweri WFS04
    |    |    |--C. pygmaeus C61
    |    |    `--C. rufescens C61
    |    `--Scolopini W77
    |         |--Lasiochiloides Champion 1901 C61
    |         `--Blaptostethus Fieber 1860 C61
    |              |--B. africanus (Carayon 1956) [=Lasiochiloides africanus] C61
    |              `--B. pleneti (Carayon 1958) [=Lasiochiloides pleneti] C61
    `--Anthocorinae C61
         |  i. s.: Wollastoniella bifoveata Carayon 1958 C61
         |--Anthocoris [Anthocorini] C61
         |    |--A. nemoralis C61
         |    |--A. nemorum H01
         |    |--A. pacificus Kirkaldy 1908 K08
         |    |--A. subcruciatus Walker 1872 C61
         |    `--A. sylvestris K08
         `--Oriini C61
              |--Lampronannella Poppius 1909 C61
              |--Dokkiocoris Miller 1951 C61
              |--Pseudotriphleps Poppius 1910 [incl. Horniella Poppius 1910 non Raffray 1904] C61
              |--Caffrocoris Carayon 1961 C61
              |    `--*C. brincki Carayon 1961 C61
              |--Bilianella C61
              |    |--B. microscopica C61
              |    `--B. minuta Carvalho 1951 C61
              |--Montandoniola Poppius 1909 [incl. Teisocoris Hiura 1959] C61
              |    |--*M. moraguesi (Puton 1896) [=Montandoniella moraguesi] C61
              |    |--‘Ectemnus’ pictipennis Esaki 1931 [=*Teisocoris pictipennis] C61
              |    `--M. thripodes Bergroth 1916 C61
              `--Orius Wagner 1952 [Oriini] C61
                   |  i. s.: O. armatus CGW91
                   |         O. cocciphagus (Hesse 1947) [=Triphleps cocciphagus] C61
                   |         O. fogoensis Wagner 1957 C61
                   |         O. insidiosus MM96
                   |         O. laevigatus WS07
                   |         O. minutum KU07
                   |         O. tantillus CGW91
                   |         O. tristicolor WT91
                   |--O. (Orius) C61
                   |    |--O. (O.) brunnescens (Poppius 1913) [=Triphleps brunnescens] C61
                   |    |--O. (O.) niger C61
                   |    `--O. (O.) thripoborus (Hesse 1940) [=Triphleps thripoborus] C61
                   `--O. (Dimorphella) lanatus Carayon 1961 C61

Xylocoris (Arrostus) flavipes (Reuter 1875) [=Piezostethus flavipes; incl. Triphleps frumenti Zacher 1922, T. ramae Narayanan & Chatterji 1953, T. sinui Narayanan & Chatterji 1952] C61

*Type species of generic name indicated


[C61] Carayon, J. 1961. Hemiptera (Heteroptera): Anthocoridae. In: Hanström, B., P. Brinck & G. Rudebeck (eds) South African Animal Life: Results of the Lund University Expedition in 1950–1951 vol. 8 pp. 533–557. Almqvist & Wiksell: Uppsala.

[CGW91] Carver, M., G. F. Gross & T. E. Woodward. 1991. Hemiptera (bugs, leafhoppers, cicadas, aphids, scale insects etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 429–509. Melbourne University Press: Carlton (Victoria).

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a , 7: 5–72.

[H01] Horváth, G. 1901. Hemipterák [Hemiptera]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 245–274. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[KU07] Khanjani, M., & E. A. Ueckermann. 2007. Natural enemies of the walnut pest Aceria tristriatus (Nal.) (Acari: Eriophyidae) in western Iran. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 451–453. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[K08] Kirkaldy, G. W. 1908. A catalogue of the Hemiptera of Fiji. Proceedings of the Linnean Society of New South Wales 33: 345–391, pl. 4.

[MM96] Mound, L. A., & R. Marullo. 1996. The thrips of Central and South America: an introduction (Insecta: Thysanoptera). Memoirs on Entomology, International 6: 1–487.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.

[SP02] Shcherbakov, D. E., & Yu. A. Popov. 2002. Superorder Cimicidea Laicharting, 1781: Order Hemiptera Linné, 1758. The bugs, cicadas, plantlice, scale insects, etc. [=Cimicida Laicharting, 1781, =Homoptera Leach, 1815+Heteroptera Latreille, 1810). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 143–157. Kluwer Academic Publishers: Dordrecht.

[WS07] Walzer, A., & P. Schausberger. 2007. Key factors of intraguild predation among Phytoseiulus persimilis, Neoseiulus californicus (Acari: Phytoseiidae) and Frankliniella occidentalis (Thysanoptera: Thripidae). In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 645–648. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[WT91] Wilson, L. T., P. J. Trichilo, D. L. Flaherty, R. Hanna & A. Corbett. 1991. Natural enemy-spider mite interactions: comments on implications for population assessment. In Dusbábek, F. & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 167–173. SPB Academic Publishing: The Hague.

[WFS04] Winks, C. J., S. V. Fowler & L. A. Smith. 2004. Invertebrate fauna of boneseed, Chrysanthemoides monilifera ssp. monilifera (L.) T. Norl. (Asteraceae: Calenduleae), an invasive weed in New Zealand. New Zealand Entomologist 27: 61–72.

[W77] Woodward, T. E. 1977. The occurrence of Almeidini (Hemiptera: Anthocoridae) in Australia, with the description of a new genus. Systematic Entomology 2: 255–260.

[YS01] Yoshizawa, K., & T. Saigusa. 2001. Phylogenetic analysis of paraneopteran orders (Insecta: Neoptera) based on forewing base structure with comments on monophyly of Auchenorrhycha (Hemiptera). Systematic Entomology 26: 1–13.

Last updated: 30 January 2019.


  1. I worked for the late Tom Woodward as a research assistant on Anthocoridae. Why don't you list any of his (and Tony Postle's) papers or species like Orius heterorioides Woodward & Postle, 1986 or Australmeida derricki (Gross, 1954)?

  2. I simply wouldn't have gotten to them yet; thanks for bringing them to my attention. I do update these pages over time; keep an eye on the "Recently Updated Posts" list to the right.


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