Euarthropoda

Reconstruction of Mollisonia plenovenatrix by J. Liang, from Aria & Caron (2019).

Belongs within: Arthropoda.
Contains: Pycnogonida, Euchelicerata, Artiopoda, Marrellomorpha, Myriapoda, Fuxianhuiida, Hymenocarina, Oligostraca, Allotriocarida, Tantulocarida, Thecostraca, Copepoda, Malacostraca.

The name Euarthropoda refers to the most exclusive clade formed by crown-group arthropods. Recent analyses agree on a division of living arthropods between three major clades, the Chelicerata, Myriapoda and Pancrustacea, with the latter two usually being united to form a clade of arthropods possessing mandibles. Fossil euarthropod taxa not belonging to these clades are best known from the Cambrian period with the exception of the Artiopoda (the clade including the trilobites). The Chelicerata have a single pair of pre-oral appendages, the chelicerae, that are usually adapted for manipulating food. The Cambrian Mollisoniidae are a group of likely stem-chelicerates characterised by a broad, helmet-like head shield bearing wide eye notches, seven free segments in the anterior portion of the trunk extending into short pleurae, and posterior trunk segments fused into a pygidium (Aria & Caron 2019).

The clade Pancrustacea includes the modern crustaceans and insects with the latter a derived subgroup of the former (Regier et al. 2010). This last grouping is primarily supported by molecular data; potential synapomorphies of the Pancrustacea have all become modified in various subclades. Relationships within the pancrustaceans remain subject to analysis. A relationship between the Tantulocarida and Thecostraca has been suggested on the basis of gonopore position in the two groups; tantulocarids and basal thecostracans are also both sessile parasites.

<==Euarthropoda (see below for synonymy)
    |  i. s.: Dala peilertae Müller 1983 ZS07
    |         Walossekia quinquespinosa Müller 1983 ZS07
    |--Panchelicerata AC19
    |    |--+--Sanctacaris Briggs & Collins 1988 AC19, CB04 [Sanctacarida, Sanctacaridea]
    |    |  |    |--‘Utahcaris’ orion Conway Morris & Robison 1988 B95
    |    |  |    `--Sanctacaris uncata Briggs & Collins 1988 WD02
    |    |  `--Habeliida B95
    |    |       |--Habelia Walcott 1912 AC19, CB04
    |    |       |    `--H. optata AC19
    |    |       `--Ecnomocaris LSE13
    |    `--Chelicerata AC19
    |         |  i. s.: Wisangocaris barbarahardyae Jago, García-Bellido & Gehling 2016 BS20
    |         |--Mollisoniidae [Mollisoniida] AC19
    |         |    |--Mollisonia Walcott 1912 AC19, CB04 [incl. Houghtonites Raymond 1931 AC19]
    |         |    |    |--*M. symmetrica Walcott 1912 [incl. M. rara Walcott 1912] AC19
    |         |    |    |--M. gracilis Walcott 1912 [=Houghtonites gracilis] AC19
    |         |    |    |--M. plenovenatrix Aria & Caron 2019 AC19
    |         |    |    `--M. sinica Zhang et al. 2002 AC19
    |         |    |--Thelxiope Simonetta & Delle Cave 1975 AC19
    |         |    |--Corcorania Jell 1980 AC19
    |         |    |    `--C. trispinosa AC19
    |         |    `--Urokodia Hou, Chen & Lu 1989 AC19
    |         |--Pycnogonida RS10
    |         `--Euchelicerata RS10
    `--+--Retifaciida LSE13
       |    |--Retifacies Hou, Chen & Lu 1989 LSE13, CB04
       |    |    `--R. abnormalis Hou et al. 1989 CB04
       |    `--+--Pygmaclypeatus LSE13
       |       `--+--+--Kiisortoqia AC19
       |          |  `--Siriocars LSE13
       |          `--Squamacula LSE13
       |               |--S. buckorum LSE13
       |               `--S. clypeatus LSE13
       `--+--Artiopoda AC19
          `--+--Marrellomorpha AC19
             `--+--Aquilonifer Briggs, Siveter et al. 2016 AC19, BS16
                |    `--*A. spinosus Briggs, Siveter et al. 2016 BS16
                `--Mandibulata (see below for synonymy) AC17
                     |  i. s.: Ercaia [Promandibulata] C12
                     |           `--E. minuscule C12
                     |         Musacaris LSE13
                     |         Cambropachycopidae LSE13
                     |           |--Cambropachycope clarksoni Walossek & Müller 1990 BWW93
                     |           `--Goticaris longispinosa Walossek & Müller 1990 BWW93
                     |         Devonopilio Tihelka, Tian & Cai 2020 P-GS21, TTC20
                     |           `--*D. hutchinsoni Tihelka, Tian & Cai 2020 TTC20
                     |--Myriapoda AC19
                     `--+--+--Fuxianhuiida AC19
                        |  `--Hymenocarina AC19
                        `--Pancrustacea (see below for synonymy) AC17
                             |  i. s.: Nippotantulus heteroxenus Huys, Ohtsuka & Boxshall 1994 MA01
                             |         Neomysis M02
                             |           |--N. americana M02
                             |           `--N. vulgaris CS77
                             |         Perosquilla armata K-MC02
                             |           |--P. a. armata K-MC02
                             |           `--P. a. capensis K-MC02
                             |         Mesosoma Otto 1821 C92
                             |         Mixodiaptomus laciniatus D51
                             |         Pleuroncodes planipes CH97
                             |         Phalangites priscus Münster 1836 D07
                             |         Cryptophthalmus Rafinesque 1814 BR05
                             |         Cymodocea Rafinesque 1814 BR05
                             |         ‘Cymodocea’ Leach 1818 nec Rafinesque 1814 nec Lamouroux 1816 BR05
                             |         ‘Dactylopus’ Claus 1862 non Gill 1859 BR05
                             |         Donovania Leach 1814 BR05
                             |         Orientalina Kolosnitsyna 1973 BR05
                             |         Pagodina van Beneden 1853 BR05
                             |         ‘Trichia’ de Haan 1839 non Haller 1768 BR05
                             |         ‘Typhis’ Risso 1816 non Montfort 1810 [Typhidae] BR05
                             |         Fenneropenaeus chinensis DH08
                             |         Limnadella kitei Girard 1854 DS89
                             |         Anatanais robusto BBB-S95
                             |         Anthonema Walther 1904 H75
                             |           `--*A. problematica Walther 1904 H75
                             |         ‘Acherusia’ Lucas 1846 non Laporte & Gory 1837 E12
                             |           `--A. dumerilii Lucas 1846 E12
                             |         Naesea edwardsii Lucas 1846 E12
                             |         Ovalia Latreille in Griffith & Pidgeon 1833 KA-Z11
                             |         Cycloidea FGS04
                             |           |--Cyclus de Koninck 1841 [incl. Paraprosopon Gemmellaro 1890] FGS04
                             |           `--Oonocarcinus Gemmellaro 1890 FGS04
                             |         Heymonsicambria EL11
                             |         Hymenodora glacialis CS77
                             |         Ozolus Latreille 1802 L02a
                             |           `--*O. gasterostei Latreille 1802 L02b (see below for synonymy)
                             |         Amymone L02b
                             |           |--A. baccha [=Monoculus bacchus] L02b
                             |           |--A. faunus [=A. fauna, Monoculus faunus] L02b
                             |           |--A. moenas L02b
                             |           |--A. satyrus [=A. satyra, Monoculus satyrus] L02b
                             |           |--A. silene [=A. silenus, Monoculus silenus] L02b
                             |           `--A. thyas L02b
                             |         Nauplius L02b
                             |           |--N. bracteatus [=Monoculus bracteatus] L02b
                             |           `--N. saltatorius [=Monoculus saltatorius] L02b
                             |         Microdentopus gryllotalpa PP64
                             |         Aristeomorpha foliacea PP64
                             |         Ligur ensiferus PP64
                             |         Paromola cuvieri PP64
                             |         Anamathia rissoana PP64
                             |         Ergasticus clouei PP64
                             |         Dorynchus thompsoni PP64
                             |         Cymopolia caroni PP64
                             |         Halopsyche de Saussure 1857 BR17
                             |         Palpipes Roth 1851 K18
                             |         Berndtia purpurea CG97
                             |         Trypetasa lampas CG97
                             |         Amblyops crozetii O01
                             |         Harpactocarcinus tumidus H79
                             |         Tuzoiidae O68
                             |           |--Tuzoia O68
                             |           `--*Dioxycaris argenta (Walcott 1888) [=Leperditia argenta, Isoxys argenta] O68
                             |         Corynuropsis Scott 1894 E00
                             |--Oligostraca AC19
                             `--Altocrustacea [Ostraca, Phyllopodomorpha, Thoracopoda] RS10
                                  |--Allotriocarida SC17
                                  `--Multicrustacea [Communostraca, Vericrustacea] SC17
                                       |  i. s.: Arenosicaris inflata SC17
                                       |--+--Tantulocarida LS02
                                       |  `--Thecostraca SC17
                                       `--+--Copepoda SC17
                                          `--Malacostraca SC17

Euarthropoda [Arachnata, Arachnomorpha, Aspidonia, Aspidota, Cormogonida, Eotrilobitacea, Myriochelata, Paradoxopoda, Polymeria, Skaracarida]

Mandibulata [Antennata, Atelocerata, Biantennata, Labiophora, Labrophora, Opisthogoneata, Tracheata] AC17

*Ozolus gasterostei Latreille 1802 L02b [incl. Argulus delphini L02b, Monoculus delphinus L02a, M. gyrini L02b]

Pancrustacea [Archoptera, Copepodoida, Copepodomorpha, Crustacea, Crustaceanomorpha, Entomostraca, Eucrustacea, Gymnota, Maxillopoda, Pneumonura, Progonomorpha, Pseudopoda, Tetraconata, Thecostracomorpha] AC17

*Type species of generic name indicated

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Last updated: 29 December 2021.